Cryptops stabilis

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Taxonavigation

Ordo: Scolopendromorpha
Familia: Cryptopidae
Genus: Cryptops

Name

Cryptops stabilis Chamberlin, 1944 – Wikispecies link – Pensoft Profile

• Cryptops stabilis Chamberlin, 1944 Notul. Nat. no. 147: 1.

Material examined

ANSP Holotype labelled Type 9921/lot 1398. Cryptops stabilis Ch. Malabar XII-1920 in a small unlabelled micro vial in a larger vial in which there was a second specimen placed in a second microvial and labelled Cryptops stabilis second specimen by J. G. E. L. The two are certainly conspecific.
NB. Chamberlin stated that the type from Goenoeng (=Gunung?) Malabar, Java was the only specimen seen.

Diagnosis

Length 11 mm. Without dark subcuticular pigment. Cephalic plate and all tergites without paramedian sutures or sulci. Each side of coxosternite with 2 large and 2–3 smaller submarginal setae. Poison calyx oval. Coxopleural pores large, pore field occupying anterior 60% of coxopleuron. Ultimate leg prefemur with medial longitudinal glabrous strip and with 6 tibial and 3 tarsal saw teeth. Legs 1–20 with minute pretarsal accessory spurs.

Description of holotype

(Chamberlin’s (1944) data in parentheses where relevant).
Length 11 mm. Cephalic plate and tergite 1 orange, trunk light brown with darker subcuticular pigment (head and first and last tergites yellow or orange, the remaining portion of body dark).
Two basal antennal articles with long setae, third with long and many short, fourth and subsequent with dense short setae and irregular basal whorl of long setae. Cephalic plate without sutures. Tergite 1 without sutures and overlying cephalic plate. Anterior margin of forcipular coxosternite curved (widely convex) with 2 large and 2 small submarginal setae on each side (Fig. 86). Poison gland calyx oval.
Tergites apparently without paramedian sutures or sulci in either specimen (median sulci absent). Lateral crescentic sulci on tergites 3–18. Sternites with cruciform sulci. Transverse skeletal thickening not observable. Sternite 21 in spm. 2 with posterior margin weakly convex.
Coxopleuron with 14+16 pores of varying size (Fig. 87), the pore field, which is without setae, occupying the anterior 64% of coxopleuron and 54% in spm 2. Two setae between the pore field and the posterior margin on which there are 4 large and 2 small setae. Prefemur of ultimate leg (Fig. 88) with strong setae (Fig. 89) of moderate length on prefemur, some long ventral setae with an elongated glabrous strip on medial face. Femur with some strong setae, tibia and tarsus 1 and 2 with fine setae. Tibia with 6, tarsus with 3 saw teeth (Fig. 90). Legs 1–19 with undivided tarsi and fine setae. Leg 20 with dense setae ventrally on distal half of prefemur and ventrally on femur and tibia. Pretarsi of legs 1–20 long, with minute accessory spurs (Fig. 91).

Table 1 Status of the species of the Cryptops hortensis group of the Old World.

Species	Authority	Distribution	Status
Cryptops basilewskyi	Matic & Darabantu, 1977	St. Helena	= Cryptops nigropictus Syn. n.
Cryptops brignolii	Matic, 1977	Turkey	Valid pro tem
Cryptops campestris	Attems, 1953	Rwanda & DRC	Nomen dubium
Cryptops compositus	Chamberlin, 1952	Turkey	Nomen dubium
Cryptops covertus	Chamberlin, 1951	Angola	Valid
Cryptops daszaki	Lewis, 2002	Mauritius	Valid
Cryptops decoratus	Lawrence, 1960	Madagascar	Valid
Cryptops decoratus	sensu Lewis, 2002, 2007	Mauritius, Seychelles	= Cryptops melanotypus Syn. n.
Cryptops h. hortensis	(Donovan, 1810)	Europe, Central Asia, widely introduced	Valid
Cryptops hortensis atlantis	Pocock, 1891	Madeira (?Azores, Canaries)	Cryptops atlantis Stat. n.
Cryptops inermipes	Pocock, 1888	Christmas I.	Valid
Cryptops kalobensis	Goffinet, 1971	Democratic Republic Congo	= Cryptops nigropictus Syn. n.
Cryptops lobatus	Verhoeff, 1931	France, Italy	Valid
Cryptops mauritianus	Verhoeff, 1939	Mauritius	Valid
Cryptops medius	Verhoeff, 1901	Greece	Nomen dubium
Cryptops melanotypus	Chamberlin, 1941	Philippines, Mauritius, Seychelles	Valid
Cryptops nigropictus	Takakuwa, 1936	Afrotropical & Oriental regions	Valid
Cryptops niloticus	Lewis, 1967	Afrotropical region	= Cryptops nigropictus Syn. n.
Cryptops orthus	Chamberlin, 1951	Angola	Nomen dubium
Cryptops parisi parisi	Brolemann, 1920	Europe, Turkey, Canada (introduced)	Valid
Cryptops parisi cottarellii	Matic, 1977	Turkey	= Cryptops parisi parisi Syn. n.
Cryptops parisi rhenanus	Verhoeff, 1931	Germany	= Cryptops parisi parisi Syn. n.
Cryptops parisi sebini	Verhoeff, 1934	Italy	Nomen dubium.
Cryptops penicillatus	Lawrence, 1960	Madagascar	= Cryptops decoratus Syn. n.
Cryptops philammus gabonensis	Demange, 1968	Gabon	Nomen dubium
Cryptops songi	Song et al., 2010	China	Valid
Cryptops stabilis	Chamberlin, 1944	Indonesia (Java)	Valid

Distribution

Indonesia (Java).

Remarks

If absence of tergite paramedian sutures and sulci is disregarded (possible juvenile character) then the species is close to Cryptops melanotypus.

Provisional key to Cryptops s. str. species of the hortensis group

Provisional key to Cryptops s. str. species of the hortensis group — J. Lewis

NB.		Lead statement missing!    Lewis (2009) concluded that some species of Cryptops exhibit dark subcuticular pigmentation, some may or may not exhibit it and others are never pigmented. The character is used in the key but should be treated with circumspection.
1		Tibia and tarsus of ultimate leg without saw teeth. Christmas Island    Cryptops inermipes Pocock, 1888
–		Tibia and tarsus of ultimate leg with saw teeth    2
2		With the anterior edge of the forcipular coxosternite strongly bilobed. Each lobe with five short marginal setae (Fig. 66). France, Italy    Cryptops lobatus Verhoeff, 1931
–		Anterior edge of the forcipular coxosternite not strongly lobed (at most as in Cryptops covertus) (Fig. 17)    3
3		Prefemur of ultimate leg with glabrous longitudinal ventral groove    4
–		Prefemur of ultimate leg without glabrous longitudinal ventral groove    5
4		Coxosternal setae submarginal (Fig. 39), tibial saw teeth 5–8, tarsal 2–4. Europe and Atlantic Islands, western Central Asia. Widely introduced elsewhere    Cryptops hortensis (Donovan, 1810)
–		Coxosternal setae fine, marginal (Fig. 46), tibial saw teeth 15–17, tarsal 5–6. Madeira, ?Azores and Canary Is    Cryptops atlantis Pocock, 1891
5		Cephalic plate with paramedian sutures occupying posterior 30% (Fig. 78), labrum usually tripartite (Fig. 79), with 50 or more small coxal pores. Europe and SW Asia (Turkey)    Cryptops parisi Brolemann, 1920
–		Cephalic plate without (Fig. 1) or with very short posterior paramedian sutures, labrum unipartite, coxopleuron with maximum of 26 (rarely to 31) pores    6
NB.	1	Lead statement missing!    Up to 50 pores in C. cf. hortensis a high altitude (1,000–1,300 m) population described from Karnten, Austria, by Pichler (1987). A hortensis-like population but lacking the ventral groove on the femora of the ultimate legs.
NB.	2	Lead statement missing!    The inadequately described Cryptops brignolii near here.
6		Anterior edge of forcipular coxosternite bilobed with 6+6 stout marginal setae (Fig. 17) and about 25 coxopleural pores. Angola    Cryptops covertus Chamberlin, 1951
–		Anterior edge of forcipular coxosternite not markedly bilobed and with fine setae    7
7		20–32 mm, with 33–65 coxopleural pores, pretarsal accessory spurs minute (Fig. 70), 3+4–5+5 long submarginal coxosternal setae (Fig. 68). Mauritius    Cryptops mauritianus Verhoeff, 1939
–		Maximum length 20 mm, with 7–25 (rarely 31 in Sudanese Cryptops nigropictus) pretarsal accessory spurs short to long (except in Cryptops stabilis where they are also minute)    8
8		Tergites without paramedian sutures or sulci, pretarsal accessory spurs minute. Length 11 mm. Java (may be an immature specimen)    Cryptops stabilis Chamberlin, 1944
–		Tergites with paramedian sutures (may be incomplete), pretarsal accessory spurs may be short but not minute    9
9		Pretarsi of ambulatory legs each with a single long accessory pretarsal spur    10
–		Pretarsi of ambulatory legs with two accessory spurs, the second sometimes very small and easily overlooked    11
Couplet_ID_missing!		Lead statement missing!    Cryptops songi Song et al., 2010 a species with short cephalic posterior paramedian sutures, without subcutaneous pigment and with 26+26 coxopleural pores, China (near here).
Couplet_ID_missing!		Lead statement missing!    (The inadequately described Cryptops orthus Chamberlin, 1951, Angola, and Cryptops campestris Attems, 1953, Rwanda and Democratic Republic of Congo also near here).
10		Forcipular coxosternite with 2+3 fine marginal setae. Ultimate leg tibia with 3 saw teeth, tarsus with 2. Ambulatory legs with single long pretarsal accessory spurs (Fig. 27). 7.5 mm. Without dark subcuticular pigment. Mauritius (Ile aux Aigrettes)    Cryptops daszki Lewis, 2002
–		Forcipular coxosternite generally with 2+2 long submarginal setae and several shorter setae. Ultimate leg tibia with 4–8 saw teeth, tarsus with (2)3–4. With dark subcutaneus pigment. St. Helena, Sudan, Eritrea, Somalia, ?Yemen, Democratic Republic of Congo, Rodrigues, Taiwan    Cryptops nigropictus Takakuwa, 1936
11		With 6–9 coxopleural pores, 5–9 setae between pore field and coxopleural margin, ultimate prefemur without medial longitudinal glabrous area. Madagascar    Cryptops decoratus Lawrence, 1960
–		With 7–16 coxopleural pores, 1–3(4) setae between pore field and coxopleural margin, ultimate prefemur with broad medial longitudinal glabrous area. Philippines, Mauritius, Seychelles    Cryptops melanotypus Chamberlin, 1941

Taxon Treatment

• Lewis, J; 2011: A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha journal of myriapodology/ International Journal of Myriapodology, 4: 11-50. doi

Images

Figures 1–7. Cryptops basilewskyi. 1 Cephalic plate and tergites 1 and 2 (specimen 1615/V) 2 Clypeus (specimen 6269/V) 3 Anterior margin of forcipular coxosternite (specimen 6269/V) 4 Terminal segments, ventral (specimen 1615/V) 5 Detail of coxopleuron 6 Pretarsus leg 2 (specimen 425/V). Cryptops brignolii. 7 Labrum after Matic 1977. Scale line Figs 1, 4 = 0.5 mm, Figs 2, 5 = 0.5 mm, Figs 3, 6 = 0.1 mm.  Figures 8–23. Cryptops campestris syntypes. 8 Clypeus 9 Anterior margin of forcipular coxosternite (slide 1) 10 Anterior margin of forcipular coxosternite (slide 2) 11 Poison gland calyx (slide 2) 12 Ultimate leg (slide 2) 13 Ventromedial spine from ultimate prefemur 14 Tibial saw teeth ultimate leg 15 Tarsal saw teeth ultimate leg 16 Pretarsus leg 3 (slide 2). Cryptops covertus lectotype17 Anterior margin of forcipular coxosternite 18 Detail of left half of forcipular coxosternite 19 Ultimate leg coxopleuron 20 Ultimate leg median 21 Spine-like seta from ultimate prefemur 22 Tibial and tarsal saw teeth ultimate leg 23 Pretarsus of leg 16. Scale lines = 0.1 mm except Figs 12, 19, 20 = 0.5 mm, Figs 17, 22 = 0.25 mm and Figs 13, 21 = 0.05 mm.  Figures 24–38. Cryptops daszaki holotype after Lewis 2002. 24 Anterior margin of forcipular coxosternite 25 Poison gland calyx 26 Terminal segments ventral 27 Pretarsus leg 11. Cryptops decoratus holotype. 28 Left coxopleuron and prefemur of ultimate leg dorsolateral 29 Seta from ultimate leg prefemur 30 Pretarsus of an ambulatory leg. “Cryptops decoratus” 31 Anterior margin of forcipular coxosternite (Round Island Mauritius specimen 2). 32 Poison gland (Round Island after Lewis 2002) 33 Pretarsus leg 12 (Round Island specimen 3) 34 Clypeus (Aride, Seychelles, after Lewis 2007b) 35 Sternite 21 (Aride, after Lewis 2007b) 36 Ultimate leg coxopleuron (Aride specimen 2) 37 Tibial and tarsal saw teeth ultimate leg (Silhouette, after Lewis 2007b) 38 Pretarsus, leg mid body region (Silhouette specimen L2). Scale lines = 0.1 mm except Fig. 28 = 0.5 mm, Figs 35, 36 = 0.25 mm, Figs 29, 30, 32, 38 = 0.05 mm.  Figures 39–44. Cryptops hortensis hortensis. 39 Anterior margin of forcipular coxostrernum (St Helena specimen 1) 40 Left forcipule and coxosternite (La Guardia, Spain specimen C17) 41 Terminal segments, left coxopleuron and part of left prefemur of ultimate leg (Bournemouth, U.K.) 42 Tibial saw teeth of ultimate leg (after Brolemann 1930) 43 Tarsal saw teeth of ultimate leg (after Brolemann 1930) 44 Pretarsus leg 12 (Bournemouth, U.K.). Scale lines = 0.5 mm except Figs 39 = 0.1 mm, Fig. 44 = 0.05 mm.  Figures 45–54. Cryptops hortensis atlantis holotype. 45 Clypeus 46 Anterior margin of forcipular coxosternite 47 Right forcipule 48 Detail of poison gland calyx 49 Sternite 21 and right coxopleuron (sternite setae omitted) 50 Left ultimate leg 51 Ultimate leg tibial saw teeth 52 Detail of saw teeth and setae 53 Ultimate leg tarsal saw teeth 54 Pretarsus of ambulatory leg. Scale lines= 0.5 mm except Figs 48, 51, 53, 54 = 0.1 mm.  Figures 66–77. Cryptops lobatus. 6. Forcipular coxosternite and right forcipule (after Verhoeff 1931) 67 Ultimate leg coxopleuron (after Iorio and Geoffroy 2008). Cryptops mauritianus. 68 Anterior margin of forcipular coxosternite (after Lewis 2002) 69 Left coxopleuron (after Lewis 2002) 70 Pretarsus leg 4 (after Lewis 2002). Cryptops melanotypus holotype 71 Anterior margin of forcipular coxosternite 72 Ultimate leg tibial and tarsal saw teeth 73 Pretarsus of ambulatory leg (freehand sketch). Cryptops niloticus 74 Anterior margin of forcipular coxosternite (holotype Sudan) 75 Anterior margin of forcipular coxosternite (Yemen) 76 Anterior margin of forcipular coxosternite (Rodrigues, after Lewis 2002) 77 Pretarsus leg 5 (Rodrigues specimen 5). Scale lines: Figs 68, 69 = 0.5 mm. Figs 71, 24–26 = 0.25 mm. Figs 70, 72, 73, 77 = 0.1 mm.  Figures 78–91. Cryptops parisi parisi. 78 Cephalic plate (after Brolemann 1930) 79 Labrum (after Pichler 1987) 80 Anterior margin of forcipular coxosternite (after Lewis 2009) 81 Calyx of poison gland (after Lewis 2009) 82 Coxopleuron (after Eason 1964) 83 Ultimate leg tibial saw teeth (after Brolemann 1930) 84 Ultimate leg tarsal saw teeth (after Brolemann 1930) 85 Pretarsus leg 5 (after Lewis 2009). Cryptops stabilis.86 Anterior margin of forcipular coxosternite (specimen 2) 87 Right coxopleuron (holotype) 88 Prefemur of ultimate leg medial (holotype) 89 Ventral seta from ultimate leg prefemur (holotype) 90 Ultimate leg tibial and tarsal saw teeth (holotype) 91 Pretarsus leg 16 (holotype). Scale lines = 0.1 except Fig. 84 = 0.5 mm. Fig. 87 = 0.25 mm. Figs 85 & 89 = 0.05 mm.