Buthus tunetatus
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Ordo: Scorpiones
Familia: Buthidae
Genus: Buthus
Name
Buthus tunetatus (Herbst, 1800) – Wikispecies link – Pensoft Profile
- Scorpio tunetanus Herbst 1800[1]: 68–69, pl. III, fig. 3 (not pl. II, fig. 2, as listed in the text); Latreille 1804[2]: 122–124.
- Androctonus (Leiurus) tunetanus: Ehrenberg in Hemprich and Ehrenberg 1829[3]: 354.
- Androctonus (Leiurus) tunetanus genuinus: Ehrenberg in Hemprich and Ehrenberg 1829[3]: 354.
- Androctonus (Liurus) tunetanus: Ehrenberg in Hemprich and Ehrenberg 1831[4] (pages unnumbered)
- Androctonus (Liurus) tunetanus genuinus: Ehrenberg in Hemprich and Ehrenberg 1831[4] (pages unnumbered).
- Androctonus tunetanus: C. L. Koch 1845[5]: 15–19, pl. CCCCI (sic), fig. 968; C. L. Koch 1850[6]: 90.
- Buthus occitanus (MIS): Kovařík and Whitman 2005 (part): 106.
- Buthus occitanus tunetanus: Birula 1903[7]: 107; Birula 1910[8]: 118; Borelli 1914a[9]: 154–155; Borelli 1914b[10]: 461; Borelli 1924[11]: 4–5; Borelli 1928[12]: 351; Giltay 1929[13]: 196–197; Werner 1929[14]: 30–31; Caporiacco 1932[15]: 395–396; Schenkel 1932[16]: 379–380; Werner 1932[17]: 300–305; Pallary 1934[18]: 99; Borelli 1934[19]: 169; Werner 1934b[20]: 84–85, fig. 4; Werner 1936[21]: 173; Caporiacco 1937[22]: 345; Schenkel 1949[23]: 186; Vachon 1949c[24]: 344–353, fig. 381–393; Vachon 1951a[25]: fig. 670; Vachon 1952a[26]: 272–281, fig. 381–393, 670; Vachon 1966[27]: 211; Peréz 1974: 22; Levy and Amitai 1980[28]: 16; El-Hennawy 1992[29]: 98, 121; Kovařík 1995[30]: 20; Kovařík 1997[31]: 179; Gantenbein et al. 1998[32]a: 51; Gantenbein et al. 1998[32]b: 33–39; Kovařík 1998[33]: 106; Fet and Lowe 2000[34]: 97; Lourenço 2002[35], p. 113, 115, fig. 8–9, 11, 13; Kovařík 2002[36]: 6; Gantenbein and Largiadèr 2003[37]: 120, 122; Ben Othmen et al. 2004[38]: 257; Touloun 2012[39]: 37, 41.
- Buthus (Buthus) occitanus tunetanus: Birula 1908[40]: 123–124; Birula 1909[41]: 507–508, fig. B; Birula 1910[8]: 156–157; Birula 1917a[42]: 223; Roewer 1943[43]: 206.
- Buthus tunetanus: Simon 1872[44]: 251–252; Lourenço 2003[45]: 897–899, fig. 57–61; Kovařík 2006[46]: 2, 8, 10, 15, fig. 16–19.; Sadine et al. 2011[47]: 6; Lourenço and Cloudsley-Thompson 2012[48]: 13–16, fig. 8; Lourenço and Simon 2012[49]: 12; Lourenço 2013[50]: 65–66; Rossi, Tropea and Yağmur 2013: 4–5, 7
- [[ | ]] Scorpion Tunetanus (ISS): Latreille 1817[51]: 106.
Type material
Types lost according to Fet and Lowe (2000)[34], Tunisia.
Distribution
The species is currently distributed across Algeria, Libya, Morocco and Tunisia, and doubtfully in the island of Malta.
Remarks
Because of the description of new species from Tunisia, and the lack of both type specimen and locality (beyond the country), a neotype for B. tunetanus is necessary to stabilize the taxonomy of Tunisian Buthus. As explained for B. paris, this is further complicated by recent diagnoses of B. tunetanus that differ from those offered by Vachon (1952a)[26], and as such if Vachon’s B. tuntetanus material is found in the MNHN it should be given priority in the future designation of a neotype. Vachon (1952a)[26] did not formally described any variety of B. tunetanus, but he again split the specimens that compose the species into four regions: 1) the typical region, corresponding to north and central Tunisia; 2) the southern montane region of Algeria, specimens from which have subsequently been described as B. tassili; 3) the Algerian Saharan Atlas and the southern region of the High Plateau; and 4) the disjunct desert regions of southern Tunisia, western central Algeria and eastern central Morocco. It is unclear whether region 3 or 4 might either correspond to B. dunlopi or B. saharicus. As explained in Fet and Lowe (2000)[34], the name A. (Leiurus) t. genuinus refers to the nominotypical form of the species and as such the adjective “genuinus” is not an available subspecific name.
Taxon Treatment
- Sousa, P; Arnedo, M; Harris, D; 2017: Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae) ZooKeys, (686): 15-84. doi
Other References
- ↑ Herbst J (1800) Natursystem der ungeflügelten Insekten. Naturgeschichte der Skorpionen. Gottlieb August Lange, Berlin, 86 pp.
- ↑ Latreille P (1804) Histoire des Scorpions. In: Histoire naturelle, générale et particulière des Crustacés et des Insectes. Ouvrage faisant suite aux oeuvres de Leclerc de Buffon, et partie de cours complet d'histoire naturelle rédigée par C. S. Sonnin, member de plusieurs Sociétés savantes. Imprimerie de F. Dufart, Paris, 110–129.
- ↑ 3.0 3.1 Hemprich F, Ehrenberg C (1829) Vorläufige Uebersicht der in Nord-Afrika und West-Asien einheimischen Scorpione und deren geographischen Verbreitung, nach den eigenen Beobachtungen von Dr. Hemprich und Dr. Ehrenberg. Verhandungen der Gessellschaft naturforschende Freunde in Berlin 1: 348–362. https://books.google.pt/books?id=8OoTAAAAQAAJ
- ↑ 4.0 4.1 Hemprich F, Ehrenberg C (1831) Animalia articulata. Arachnoidea. Scorpiones africani et asiatici. In: Hemprich et Ehrenberg Symbolae Physicae. Animalia evertebrata exclusis insectis. Percensuit Dr. C. G. Ehrenberg. Series prima cum tabularum decade prima. Continent Animalia Africana et Asiatica 162. Berolini ex officina Academica, 12 (unnumbered). https://doi.org/10.5962/bhl.title.107403
- ↑ Koch C (1845) Die Arachniden. Getreu nach der Natur abgebildet und beschrieben. 12. C.H. Zeh'schen Buchhandlung, Nürnberg, 166 pp. http://www.biodiversitylibrary.org/item/55844.
- ↑ Koch C (1850) Skorpionen. Scorpiones. In: Übersicht des Arachnidensystems. Fünftes Heft. Verlag von J. L. Lotzbeck, Nürnberg, 86–92. http://www.biodiversitylibrary.org/bibliography/43744.
- ↑ Birula A (1903) Bemerkungen über einige neue oder wenig bekaunte Skorpionnenformen Nord-Afrikas. Bulletin de l'Académie Impériale des Sciences de St-Pétersbourg [Izviestiia Imperatorskoi akademii nauk] 19: 105–113.
- ↑ 8.0 8.1 Birula A (1910) Ueber Scorpio maurus Linné und seine Unterarten. Horae Societatis Entomologicae Rossicae 39: 115–192. https://www.ntnu.no/ub/scorpion-files/Birula_1910.pdf
- ↑ Borelli A (1914a) Contributo allo studio della fauna Libica. Materiali raccolti nelle zone di Misurata e Homs (1912-13). Scorpioni. Annali del Museo Civico di Storia Naturale di Genova 6: 148–159.
- ↑ Borelli A (1914b) Gli scorpioni del Museo Civico di Storia Naturale di Milano. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano 53: 456–464.
- ↑ Borelli A (1924) Missione zoologica del Dr. E. Festa in Cirenaica. XVI. Scorpioni e Solifughi. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 39: 1–16.
- ↑ Borelli A (1928) Risultati zoologici della Missione inviata dalla R. Societa Geografica Italiana per l’explorazione dell’oasi di Giarabud (1926-1927). Scorpioni e Solifughi. Annali del Museo civico di Storia Naturale Giacomo Doria 52: 346–355.
- ↑ Giltay L (1929) Mission saharienne Augieras-Draper, 1927–1928. Scorpions. Bulletin du Muséum National d'Histoire Naturelle, Paris, 2eme série 1: 193–197.
- ↑ Werner F (1929) Wissenschaftliche Ergebnisse einer zoologischen Forschungsreise nach Westalgerien und Marokko. III. Scorpiones. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Wien 138: 1–34. http://www.zobodat.at/publikation_articles.php?id=169321.
- ↑ Caporiacco L (1932) Spedizione scientifica all’Oasi di Cufra (Marzo-Luglio 1931). Scorpioni e Solifugi. Annali del Museo civico di Storia Naturale Giacomo Doria 55: 395–408.
- ↑ Schenkel E (1932) Notizen üeber einige Scorpione und Solifugen. Revue Suisse de Zoologie 39: 375–396.
- ↑ Werner F (1932) Ergebnisse einer zoologischen Forschungsreise nach Marokko. VI. Skorpione. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Wien 141: 285–306. http://www.zobodat.at/publikation_articles.php?id=180394.
- ↑ Pallary P (1934) Scorpions du Sahara central. Mission du Hoggar III (Février à Mai 1928). Mémoires de la Société d'Histoire Naturelle d'Afrique du Nord 4: 90–100.
- ↑ Borelli A (1934) Scorpiones. In: Zavattari E (Ed.) Prodromo della fauna della Libia. Tipografia già Cooperativa, Pavia, 169–173.
- ↑ Werner F (1934b) Über skorpione aus Marokko. Verhandlungen der Naturforschenden Gesellschaft in Basel 45: 82–88.
- ↑ Werner F (1936) Neu-Eingänge von Skorpionen im Zoologischen Museum in Hamburg. Festschrift zum 60. Geburtstage von Professor Dr. Embrik. Strand, Riga 2: 171–193.
- ↑ Caporiacco L (1937) Risultati scientifici della Missione del Prof. G. Scortecci nel Fezzan e sui Tassili (1936). Scorpioni e Solifughi. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano 76: 340–354.
- ↑ Schenkel E (1949) Mitteilungen über Spinnentiere. Skorpione aus Marokko. Verhandlungen der Naturforschenden Gesellschaft in Basel 60: 186–201.
- ↑ Vachon M (1949c) Études sur les Scorpions. III (suite). Déscription des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algérie 27: 334–396.
- ↑ Vachon M (1951a) Études sur les Scorpions. (suite et fin). Détermination des Scorpions du Nord-Ouest de l'Afrique. Archives de l'Institut Pasteur d'Algérie 29: 46–104.
- ↑ 26.0 26.1 26.2 Vachon M (1952a) Études sur les scorpions. (published 1948–1951 in Archives de l'Institut Pasteur d'Algérie, 1948, 26: 25–90, 162–208, 288–316, 441–481; 1949, 27: 66–100, 134–169, 281–288, 334–396; 1950, 28: 152–216, 382–413; 1951, 29: 46–104). Institut Pasteur d'Algérie, Alger, 482 pp. https://www.ntnu.no/ub/scorpion-files/vachon.php.
- ↑ Vachon M (1966) Liste des scorpions connus en Egypte, Arabie, Israeli, Liban, Syrie, Jordanie, Turquie, Irak, Iran. Toxicon 4: 209–218.
- ↑ Levy G, Amitai P (1980) Fauna Palaestina. Arachnida I: Scorpiones. The Israel Academy of Sciences an d Humanit es, Jerusalem, 172 pp.
- ↑ El-Hennawy H (1992) A catalogue of the scorpions described from the Arab countries (1758-1990) (Arachnida: Scorpionida). Serket 2: 95–153.
- ↑ Kovařík F (1995) First report of Orthochirus innesi (Scorpionida: Buthidae) from Morocco. Klapalekiana 31: 19–21.
- ↑ Kovařík F (1997) A check-list of scorpions (Arachnida) in the collection of the Hungarian Natural History Museum, Budapest. Annales Historico-Naturales Musei Nationalis Hungarici 89: 177–185.
- ↑ 32.0 32.1 Gantenbein B, Braunwalder M, Scholl A (1998) Allozyme studies on scorpions from the Aegean Region and from Morocco. Abstracts XIV International Congress Arachnology \ 22 Annual Meeting of the American Arachnological Society, Chicago, 51.
- ↑ Kovařík F (1998) Štiří. Madagaskar, Jihlava, 176 pp.
- ↑ 34.0 34.1 34.2 Fet V, Lowe G (2000) Genus Buthus Leach, 1815. In: Fet V Sissom W Lowe G Braunwalder M (Eds) Catalog of the Scorpions of the World (1758-1998). The New York Entomological Society, New York, 91–97.
- ↑ Lourenço W (2002) Considérations sur les modèles de distribution et différentiation du genre Buthus Leach, 1815, avec la description d'une nouvelle espèce des montagnes du Tassili des Ajjer, Algérie (Scorpiones, Buthidae). Biogeographica 78: 109–127.
- ↑ Kovařík F (2002) A checklist of scorpions (Arachnida) in the collection of the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany. Serket 8: 1–23.
- ↑ Gantenbein B, Largiadèr C (2003) The phylogeographic importance of the Strait of Gibraltar as a gene flow barrier in terrestrial arthropods: a case study with the scorpion Buthus occitanus as model organism. Molecular Phylogenetics and Evolution 28: 119–130. https://doi.org/10.1016/S1055-7903(03)00031-9
- ↑ Ben Othmen A, Chatti N, Ben Ali-Haouas Z, Ouldbrahim I, Said K (2004) Allozymic differentiation of Tunisian populations of Androctonus species and Buthus occitanus (Scorpiones: Buthidae). Biological Journal of the Linnean Society 81: 255–265. https://doi.org/10.1111/j.1095-8312.2003.00285.x
- ↑ Touloun O (2012) Chapitre III. Ecologie et repartition de la scorpiofaune du Sud Ouest Marocain. In: Les peuplements de scorpions du sud ouest marocain: Ecologie, Biogéographie et Epidémiologie des envenimations. Éditions universitaires européennes, Saarbrücken, 30–82.
- ↑ Birula A (1908) Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Ägyptischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Wien 117: 121–152. http://www.zobodat.at/publikation_articles.php?id=148339
- ↑ Birula A (1909) Scorpione und Solifugen von Tripolis und Barka. Nach der Sammlung von Dr. Bruno Klaptocz im jahre 19-06. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 28: 505–522.
- ↑ Birula A (1917a) [authored as Byalynitskii-Birulya, A. A.] Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones. Zapiski Kavkazskogo Muzeya, Seriia A [Mémoires du Musée du Caucase, Série A] 5: 1–253. [English translation: Byalynitskii-Birulya AA, 1964. Arthrogastric Arachnids of Caucasia. Part I. Scorpions. Israel Program for Scientific Translations, Jerusalem. 170 pp.]
- ↑ Roewer C (1943) Über eine neuerworbene Sammlung von Skorpionen des Natur-Museums Senckenberg. Senckenbergiana 26: 205–244.
- ↑ Simon E (1872) Arachnides de Syrie rapportés par M. Charles Piochard de la Brûlerie (Scorpions et Galeodes). Annales de la Société Entomologiqe de France, 5eme série 2: 245–266.
- ↑ Lourenço W (2003) Compléments à la faune de scorpions (Arachnida) de l'Afrique du Nord, avec des considérations sur le genre Buthus Leach, 1815. Revue Suisse de Zoologie 110: 875–912.
- ↑ Kovařík F (2006) Review of Tunisian Species of the Genus Buthus with Descriptions of Two New Species and a Discussion of Ehrenberg’s Types (Scorpiones: Buthidae). Euscorpius 34: 1–16. http://www.science.marshall.edu/fet/euscorpius/p2006_34.pdf.
- ↑ Sadine S, Samia B, Ould Elhadj M (2011) Premieres donnees sur la diversite scorpionique dans la region du Souf (Algerie). Arachnides 61: 1–10.
- ↑ Lourenço W, Cloudsley-Thompson J (2012) A new species of Buthus Leach, 1815 from Egypt (Scorpiones, Buthidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 16: 11–18.
- ↑ Lourenço W, Simon E (2012) Confirmation of a new species of Buthus Leach, 1815 from Alexandria, Egypt (Scorpiones, Buthidae). Serket 13: 8–15.
- ↑ Lourenço W (2013) A new species of Buthus Leach, 1815 from Algeria (Scorpiones, Buthidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 16: 63–68.
- ↑ Latreille P (1817) Tome III. Contenant les Crustacés, les Arachnides et les Insectes. In: Cuvier G (Ed.) Le règne animal distribué d'après son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparée. Imprimerie de A. Belin, Paris, 653.