Raveniola\according to Li et al 2015

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This page should be cited as follows (rationale):
Li S, Zonstein S (2015) Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae). ZooKeys (519) : 1–32, doi. Versioned wiki page: 2015-08-26, version 72908, https://species-id.net/w/index.php?title=Raveniola%5Caccording_to_Li_et_al_2015&oldid=72908 , contributors (alphabetical order): Pensoft Publishers.

Citation formats to copy and paste

BibTeX:

@article{Li2015ZooKeys,
author = {Li, Shuqiang AND Zonstein, Sergei},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae)},
year = {2015},
volume = {},
issue = {519},
pages = {1--32},
doi = {10.3897/zookeys.519.8784},
url = {http://zookeys.pensoft.net/articles.php?id=6246},
note = {Versioned wiki page: 2015-08-26, version 72908, https://species-id.net/w/index.php?title=Raveniola%5Caccording_to_Li_et_al_2015&oldid=72908 , contributors (alphabetical order): Pensoft Publishers.}

}

RIS/ Endnote:

TY - JOUR
T1 - Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae)
A1 - Li S
A1 - Zonstein S
Y1 - 2015
JF - ZooKeys
JA -
VL -
IS - 519
UR - http://dx.doi.org/10.3897/zookeys.519.8784
SP - 1
EP - 32
PB - Pensoft Publishers
M1 - Versioned wiki page: 2015-08-26, version 72908, https://species-id.net/w/index.php?title=Raveniola%5Caccording_to_Li_et_al_2015&oldid=72908 , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/zookeys.519.8784

Wikipedia/ Citizendium:

<ref name="Li2015ZooKeys">{{Citation
| author = Li S, Zonstein S
| title = Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae)
| journal = ZooKeys
| year = 2015
| volume =
| issue = 519
| pages = 1--32
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.519.8784
| url = http://zookeys.pensoft.net/articles.php?id=6246
| pmc =
| accessdate = 2024-12-12

}} Versioned wiki page: 2015-08-26, version 72908, https://species-id.net/w/index.php?title=Raveniola%5Caccording_to_Li_et_al_2015&oldid=72908 , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Taxonavigation

Ordo: Araneae
Familia: Nemesiidae

Name

Raveniola Zonstein, 1987 – Wikispecies link – Pensoft Profile

Type species

Brachythele virgata Simon, 1891, from Central Asia, by the original designation.

Diagnosis

Raveniola, similar to Sinopesa Raven & Schwendinger, 1995, has two enlarged retroventral distal spines on tibia I in males and divided receptacles in females, as well as the absence of a serrula and metatarsal preening combs. The leg scopula in Raveniola is more developed than it is in Sinopesa. By contrast, the male intercheliceral tumescence in Raveniola is lost or vestigial, whereas in Sinopesa, it is well-developed. Like Sinopesa, members of Raveniola have more or less reduced PMS, which are completely lost in some species. The apical segment of the PLS in Raveniola is usually shorter than that in Sinopesa.

Composition

Raveniola currently comprises 29 species, including the new species described here; 14 of them occur in China: Raveniola alpina sp. n., Raveniola bellula sp. n., Raveniola chayi sp. n., Raveniola gracilis sp. n., Raveniola guangxi (Raven & Schwendinger, 1995), Raveniola hebeinica Zhu, Zhang & Zhang, 1999, Raveniola montana Zonstein & Marusik, 2012, Raveniola rugosa sp. n., Raveniola shangrila Zonstein & Marusik, 2012, Raveniola songi Zonstein & Marusik, 2012, Raveniola spirula sp. n., Raveniola xizangensis (Hu & Li, 1987), Raveniola yajiangensis sp. n. and Raveniola yunnanensis Zonstein & Marusik, 2012.

Key to East Asian Raveniola species

Females of Raveniola alpina sp. n., Raveniola gracilis sp. n., Raveniola guangxi, Raveniola rugosa sp. n., Raveniola shangrila, Raveniola songi, Raveniola spirula sp. n. and Raveniola yunnanensis are unknown.

Key to East Asian Raveniola species — S. Li , S. Zonstein
1
Males 
  2
–
Females 
  15
2
PMS present 
  3
–
PMS absent 
  9
3
Carapace length > 10 mm. Embolus with distinct subapical keel 
  xizangensis
–
Carapace length 3.0–7.3 mm. Embolic keel absent or vestigial 
  4
4
Palpal tibia relatively short, with a length/width ratio of 3.0–3.2 (Fig. 11A–C). Embolus short and with deep subbasal ridges (Fig. 12A–C) 
  montana
–
Palpal tibia and embolus relatively long, with a length/width ratio of 3.6–4.5 (Figs 1A–C, 2A–C, 3A–C, 5A–C, 6A–C, 7A–C, 9A–C, 10A–C, 13A–C, 14A–C, 15A–C, 16A–C, 17A–C, 19A–C) 
  5
5
Embolus with distally hooked tip (Figs 9A–C, 10A–C) 
  gracilis sp. n.
–
Embolic tip not hooked 
  6
6
Embolus more or less twisted 
  7
–
Distal part of embolus curved gradually (Zonstein and Marusik 2012[1]: fig. 39) 
  hebeinica
7
Tibia I equal in length to or shorter than metatarsus (as in Fig. 17E). Few spines on cymbium (as in Fig. 19A–C) 
  8
–
Tibia I considerably longer than metatarsus (see Zonstein and Marusik 2012[1]: fig. 29). Cymbium with numerous dorsal spines (Op. cit.: fig. 35) 
  yunnanensis
8
Embolus only slightly twisted (Op. cit.: fig. 42) 
  songi
–
Embolus distinctly twisted (Figs 19A–C) 
  yajiangensis sp. n.
9
Embolus with hooked tip (see Zonstein and Marusik 2012[1]: figs 37, 38) 
  guangxi
–
Embolic tip not hooked 
  10
10
Embolus strongly spiralled as in Figs 15A–C, 16A–C 
  spirula sp. n.
–
Embolus more or less curved, slightly spiralled or bent as in Figs 1A–C, 2A–C, 3A–C, 5A–C, 6A–C, 7A–C, 13A–C, 14A–C 
  11
11
Embolus slightly spiralled as in Figs 1A–C, 2A–C 
  alpina sp. n.
–
Embolus curved or gradually twisted 
  12
12
Entire embolus arched as in Figs 13A–C, 14A–C 
  rugosa sp. n.
–
Embolus more or less distinctly twisted as in Figs 3A–C, 5A–C, 6A–C, 7A–C 
  13
13
Abdomen with dorsal and ventral spotted pattern (Figs 3D, G, 6D, G). Embolus only slightly twisted (Figs 3A–C, 5A–C, 6A–C, 7A–C) 
  14
–
Abdomen uniformly dark brown. Embolus noticeably twisted (see Zonstein and Marusik 2012[1]: fig. 41) 
  shangrila
14
Dorsal abdominal pattern consists of numerous darker spots on a lighter background (Fig. 3D). Metatarsus I very gently curved (Fig. 3E). Embolus moderately long without ridges (Figs 3A–C, 5A–C) 
  bellula sp. n.
–
Dorsal abdominal pattern consists of numerous lighter spots on a darker background (Fig. 6D). Metatarsus I noticeably curved (Fig. 6E). Embolus long and tapering with ridges (Figs 6A–C, 7A–C) 
  chayi sp. n.
15
PMS present 
  16
–
PMS absent 
  19
16
Carapace length > 10 mm. Median (ental) branch of receptacle bifurcate (Op. cit., fig. 50) 
  xizangensis
–
Carapace length < 8 mm. Median (ental) branch of receptacle entire 
  17
17
Median (ental) branch of receptacle curved inward as shown in Figs 18, 19D 
  yajiangensis sp. n.
–
Shape of receptacles different 
  18
18
PLS: apical segment triangular. Receptacular bases narrow (Zonstein and Marusik 2012[1], fig. 47) 
  hebeinica
–
PLS: apical segment digitiform. Receptacular bases widened (Op. cit., fig. 48) 
  montana
19
Receptacles as shown in Figs 4A, 5D 
  bellula sp. n.
–
Receptacles as shown in Figs 8B–C 
  chayi sp. n.

Taxon Treatment

  • Li, S; Zonstein, S; 2015: Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae) ZooKeys, (519): 1-32. doi

Images

Figure 1. Raveniola alpina sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 1.0 mm (A–C); 2.0 mm (D–G).
Figure 1. Raveniola alpina sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 1.0 mm (A–C); 2.0 mm (D–G). 
Figure 2. Raveniola alpina sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm.
Figure 2. Raveniola alpina sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm. 
Figure 3. Raveniola bellula sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D, G); 1.0 mm (E, F).
Figure 3. Raveniola bellula sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D, G); 1.0 mm (E, F). 
Figure 4. Raveniola bellula sp. n., female paratype. A vulva, dorsal view B genital area, ventral view. Scale bars: 0.25 mm.
Figure 4. Raveniola bellula sp. n., female paratype. A vulva, dorsal view B genital area, ventral view. Scale bars: 0.25 mm. 
Figure 5. Raveniola bellula sp. n., male holotype (A–C) and female paratype (D). A palp, prolateral view B palp, ventral view C palp, retrolateral view D vulva, dorsal view. Scale bars: 0.25 mm.
Figure 5. Raveniola bellula sp. n., male holotype (A–C) and female paratype (D). A palp, prolateral view B palp, ventral view C palp, retrolateral view D vulva, dorsal view. Scale bars: 0.25 mm. 
Figure 6. Raveniola chayi sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G).
Figure 6. Raveniola chayi sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G). 
Figure 7. Raveniola chayi sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm.
Figure 7. Raveniola chayi sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm. 
Figure 8. Raveniola chayi sp. n., female paratype. A habitus, dorsal view B, C vulva, dorsal view. Scale bars: 5.0 mm (A); 0.2 mm (B–C).
Figure 8. Raveniola chayi sp. n., female paratype. A habitus, dorsal view B, C vulva, dorsal view. Scale bars: 5.0 mm (A); 0.2 mm (B–C). 
Figure 9. Raveniola gracilis sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I (right side), ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G).
Figure 9. Raveniola gracilis sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I (right side), ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G). 
Figure 10. Raveniola gracilis sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm.
Figure 10. Raveniola gracilis sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm. 
Figure 11. Raveniola montana Zonstein & Marusik, 2012, male (Sichuan Prov.). A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G).
Figure 11. Raveniola montana Zonstein & Marusik, 2012, male (Sichuan Prov.). A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G). 
Figure 12. Raveniola montana Zonstein & Marusik, 2012, male (Sichuan Prov.). A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm.
Figure 12. Raveniola montana Zonstein & Marusik, 2012, male (Sichuan Prov.). A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm. 
Figure 13. Raveniola rugosa sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G).
Figure 13. Raveniola rugosa sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G). 
Figure 14. Raveniola rugosa sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm.
Figure 14. Raveniola rugosa sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm. 
Figure 15. Raveniola spirula sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 4.0 mm (D, G); 2.0 mm (E, F).
Figure 15. Raveniola spirula sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 4.0 mm (D, G); 2.0 mm (E, F). 
Figure 16. Raveniola spirula sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm.
Figure 16. Raveniola spirula sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm. 
Figure 17. Raveniola yajiangensis sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G).
Figure 17. Raveniola yajiangensis sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (A–C); 2.0 mm (D–G). 
Figure 18. Raveniola yajiangensis sp. n., female paratype. A vulva, dorsal view B genital area, ventral view. Scale bars: 0.25 mm.
Figure 18. Raveniola yajiangensis sp. n., female paratype. A vulva, dorsal view B genital area, ventral view. Scale bars: 0.25 mm. 
Figure 19. Raveniola yajiangensis sp. n., male holotype (A–C) and female paratype (D). A palp, prolateral view B palp, ventral view C palp, retrolateral view D vulva, dorsal view. Scale bars: 0. 5 mm (A–C); 0.25mm (D).
Figure 19. Raveniola yajiangensis sp. n., male holotype (A–C) and female paratype (D). A palp, prolateral view B palp, ventral view C palp, retrolateral view D vulva, dorsal view. Scale bars: 0. 5 mm (A–C); 0.25mm (D). 

Other References

  1. ↑ 1.0 1.1 1.2 1.3 1.4 Zonstein S, Marusik Y (2012) A review of the genus Raveniola (Araneae, Nemesiidae) in China, with notes on allied genera and description of four new species from Yunnan. ZooKeys 211: 71–99. doi: 10.3897/zookeys.211.3060
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