Gochnatia recticulifolia
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Ordo: Asterales
Familia: Asteraceae
Genus: Gochnatia
Name
Gochnatia recticulifolia H.Rob. & V.A.Funk sp. nov. – Wikispecies link – Pensoft Profile
Type
Peru. Department Ancash: Callejon de Huaylas, trail to cave across Río Santo [Río Santa] from Mancos, shrub to 3 m tall. Flowers yellow, phyllaries green. 9 April 1970, C. Earle Smith Jr. & Jacinto Blas 4901 (holotype US!). Collected in co-operation with the Seasonal Transhumance and Preceramic Occupation of the Callejon de Huaylas Project – Thomas F. Lynch, Director and according to the label the Vernacular name is Juanca blanca.
Description
Shrub to 3 m tall; stems gnarled, thickened and blackish near base, to ca. 8 mm wide, with scarcely noticeable narrow pith; younger stems as slender shoots, with internodes 3–10 mm long, covered with grayish tomentum. Leaves alternate, petioles ca. 4 mm long; laminae ovate oblong, mostly 1.6–2.5 cm long, 1.2–1.8 wide, flat or sometimes folded along midvein, base short-obtuse, apex usually rounded with slight mucro, margins flat, not recurved, entire, adaxial surface greenish with minute thin evanescent floccose puberulence, abaxial surface with grayish granular-looking pubescence consisting of slender highly contorted trichomes with few thin-walled cells at base, sometimes with slightly off-set apical cells separated by an oblique cross-wall, with a weakly prominulous midvein, 3 or 4 pairs of ascending, secondary veins, and a minute reticulum of veinlets evident on both surfaces that are not obscured by pubescence. Inflorescence of a solitary capitulum or 2–3 grouped together at tips of leafy stems; involucres broadly campanulate at anthesis, ca. 9 mm high and 11 mm diam., bracts ca. 40 in ca. 6 gradate series, basal bracts broadly ovate, 1.5–2.5 mm long, to 2.5 mm wide, rounded to obtuse at tips, inner bracts lanceolate, to ca. 7 mm long, acute, outer surface thinly pilosulous with weak indumentum near bases; receptacle slightly crested between areoles. Florets homogamous 35–40 per capitulum, corollas yellow, ca. 8–9 mm long, without long hairs outside, basal tube ca. 3.5 mm long, throat ca. 1.5 mm long, narrowly funnelform, lobes linear, ca. 3.5 mm long, ca. 0.2 mm wide; anther thecae ca. 1.8 mm long, tails ca. l.2 mm long, with a dense fringe of narrow hairs, apical appendage ca. 0.9 mm long, narrowly ovate with acuminate tip; pollen 30–40 µm in diam. and 50–55 µm long; styles slightly broadened and blunt at tips. Achene cylindrical, ca. 4 mm long, sericeous with slender setulae; pappus pale yellow with ca. 35 inner capillary bristles, up to 7 mm long, most with distinctly broader and more densely scabrous tips, and with numerous outer shorter weakly barbellate bristles of various lengths with slender tips.
Distribution
Known only from the type collection which places it in northern Peru.
Conservation status
DD (according IUCN 2019[1]).
Etymology
The epithet for Gochnatia recticulifolia is based on the minute reticulum of veinlets that is evident on both surfaces of the leaf.
Notes
The type specimen was original identified in the herbarium as G. curvifolia S.F.Blake, a potentially related species of mostly Bolivian distribution. However, G. curvifolia has pointed leaf tips and a minute reticulum of the leaf veins mostly obscured by pubescence. The corollas of G. recticulifolia have no hint of the pilosity seen in many but not all specimens of G. curvifolia and the closely related G. boliviana S.F. Blake. G. recticulifolia may actually be closer to the unseen G. vargasii Cabrera of the Department of Apurimac in Peru, but the latter is distinct in having more acute leaves and serrate leaf margins.
As a result of our investigations we emphasize the trichome character that provides an additional distinction between Gochnatia s.s. and Moquiniastrum: the trichomes on the abaxial surface of the leaf. In Gochnatia they are slender, highly contorted trichomes with a few thin-walled cells at base, sometimes with slightly off-set apical cells separated by an oblique cross-wall (Fig. 5C–E). In Moquiniastrum the trichomes are T-shaped, often long-stalked with an elongate cap-cell attached at the mid-point (Fig. 5F). We examined the generitypes of both genera as well as our two new species. Gochnatia vernonioides (Fig. 5E), the generitype of Gochnatia has some trichomes with two such oblique cross-walls and off-set cells. Certainly, the distribution of this character should be examined for all 10 species in an upcoming monograph of Gochnatia s.s. (Sancho et al., in prep.). The 10 species now recognized in Gochnatia can be distinguished by the following key. Many details are from Cabrera (1971)[2], especially regarding G. vargasii of which proper material has not been seen.
The gnarled appearance of the stems was found in the newly described G. recticulifolia and in one specimen of G. arequipensis Sandwith. The lack of this character in all other collections of Gochnatia may be an artifact of collecting and it may occur in other species but was omitted during the pressing of the plants.
Original Description
- Robinson, H; Funk, V; 2020: Two new species for Gochnatia Kunth (Asteraceae, Gochnatieae) and an Extension of the tribal Range into Ecuador PhytoKeys, 139: 51-62. doi
Images
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Other References
- ↑ IUCN (2019) The IUCN Red List of Threatened Species. Version 2019-2. http://www.iucnredlist.org [Downloaded on 18.07.2019]
- ↑ Cabrera A (1971) Revision del genero Gochnatia. Revista del Museo de La Plata, Nueva Serie.Sección Botánica12(66): 1–160.
- ↑ Panero J, Freire S (2013) Paquirea, a new Andean genus for Chucoa lanceolata (Asteraceae, Mutisioidea, Onoserideae).Phytoneuron2013: 1–11.
