Gochnatia lojaensis
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Ordo: Asterales
Familia: Asteraceae
Genus: Gochnatia
Name
Gochnatia lojaensis H.Rob. & V.A.Funk sp. nov. – Wikispecies link – Pensoft Profile
Type
Ecuador. Prov. Loja: La Toma – Catacocha road ca. km 26, shrub 3 m tall, heads yellow, 79°28'53"W, 03°58'40"S, 2300 m alt., 3 Sep 2000, Jens Elgaard Madsen with Orlandro A. Sanchez, 7209 (holotype AAU!; isotypes, LOJA, US frag.!).
Description
Shrub to 3 m tall, with numerous branches. Stems grayish, wrinkled when dry; internodes 2–5 mm long, surface with yellowish-gray, evanescent, granular appearing pubescence, composed of tightly glomerulous contorted trichomes, pith solid, ca. 2 mm wide. Leaves spirally alternate, petioles 7–8 mm long; blades narrowly elliptical, 3–6.5 cm long, 0.9–1.4 cm wide, base obtuse, margins entire, plane, apex subacute, adaxial surface green with thin evanescent pubescence in young leaves, with minute reticulum of prominulous veinlets (veins slightly prominent), secondary veins spreading from midvein at ca. 45° angles; abaxial surface covered with dense yellowish tomentum of slender highly contorted trichomes, trichomes with few thin-walled cells at base, sometimes with slightly off-set apical cells separated by an oblique cross-wall, costa prominent to near leaf-tip. Inflorescence corymbiform, with clusters of 10–15 heads apical on leafy branches, cluster usually becoming over-topped by younger branches and longer leaves, with small bracteoles 1.0–1.5 cm. long among heads; peduncles 0.7–l.0 cm long, longer peduncles with minute scale-like bracteoles; involucres campanulate, 1–1.2 cm high, ca. 0.8–0.9 cm wide at anthesis, with ca. 50 subimbricate appressed bracts in ca. 6 series, bracts progressing from basal scales ca. 1.5 mm long and wide to many progressively longer lanceolate median bracts to few somewhat deciduous linear inner bracts ca. 9 mm long and 1 mm wide, outer surfaces of bracts glabrous on most exposed surfaces, castaneous, yellow along margins, bases of bracts with yellowish tomentum of slender contorted trichomes. Upper surface of receptacles glabrous, alveolate. Florets homogamous, ca. 15 per capitulum; corollas yellowish with darkened tips, glabrous outside, ca. 9 mm long, basal tube ca. 4 mm. long, throat 1 mm long, narrowly funnelform, lobes 3.5 mm long, linear, 0.4 mm wide, coiled backward at anthesis Fig. 3A), with protuberant slightly rugulose elongate cells inside; anthers ca. 2.5 mm long (Fig. 3B), basal tails lanceolate, ca. 1 mm long, with fringed with retrorse teeth near and at tips (Fig. 3C), apical appendage indurate, ca. 1 mm long, oblong-ovate with marked apical apiculus; pollen prolate, 30–35 µm diam. and 35–40 µm long; styles slightly broadened and blunt at tips. Achenes ca. 2.5 mm long, 5-costate (Fig. 3E), densely villosulous with ascending twin-hairs, hairs not cleft at tips usually with one cell longer than other; carpopodium annuliform; pappus of ca. 50 capillary bristles, whitish, mostly ca. 7 mm long, broader and more strongly barbellate near tips (Fig. 3F), some shorter outer bristles with slender tips.
Distribution and ecology
Known only from the type collection which places it in “Matorral vegetation and ravine with disturbed remnants of montane forest.”
Conservation status
DD (according IUCN 2019[1]).
Etymology
Gochnatia lojaensis is named after the Ecuadorian province where it was collected.
Notes
Distinguishing characteristics include the corymbiform clusters of numerous heads and the narrow castaneous involucral bracts with narrowly blunted tips.
The position of the new species was at first in doubt. It was near the geographic range of Gochnatia typified by G. vernonioides Kunth from Peru, but it had the more elongate leaves often associated with the presently recognized separate genus Moquiniastrum. A detailed study of the plant now confirms a position in Gochnatia: the pubescence is particularly indicative, being a thick tomentum and not the loose stalked T-shaped hairs common in Moquiniastrum (Fig. 5F). The hairs do show one interesting tendency toward the T-shaped form, with some hairs having an apical cell that is obliquely mounted on the longer contorted cell. Such an apical cell usually has the lower end slightly projecting, a sub-T-shaped specialization. This remains totally different from the well-developed T-shape form seen in Moquiniastrum (Fig. 5F).
Original Description
- Robinson, H; Funk, V; 2020: Two new species for Gochnatia Kunth (Asteraceae, Gochnatieae) and an Extension of the tribal Range into Ecuador PhytoKeys, 139: 51-62. doi
Images
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Other References
- ↑ IUCN (2019) The IUCN Red List of Threatened Species. Version 2019-2. http://www.iucnredlist.org [Downloaded on 18.07.2019]