Acalypha paxii
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Ordo: Malpighiales
Familia: Euphorbiaceae
Genus: Acalypha
Name
Acalypha paxii Aug.D.C., Bull. Herb. Boissier, sér. 2, 1: 567. 1901. – Wikispecies link – Pensoft Profile
- Acalypha urophylla Pax, Bot. Jahrb. Syst. 19: 96. 1894. nom. illeg. non A.urophylla Boivin ex Baill. (1861). Acalyphareticulatavar.urophylloides Pax & K.Hoffm., Pflanzenr. 147, 16 (Heft 85): 105. 1924. Acalyphafiliformisvar.urophylloides (Pax & K.Hoffm.) Govaerts, World Checkl. Bibliogr. Euphorbiaceae 63. 2000.
Type
Madagascar. Prov. Antsiranana: Nossibé, nordwest, Feb 1880, J. M. Hildebrandt 3356 (holotype B or WRSL, presumably destroyed; lectotype, designated here: BREM [BREM0001784]!; isolectotypes: JE [JE00004294!, JE00004293!], K [K000186531]!, M [M0110600]!, P [P00536741!, P00536742!, P00536743!]).
Acalyphapervilleana Baill., Adansonia 1: 273. 1861. Acalyphareticulatavar.pervilleana (Baill.) Müll.Arg., Linnaea 34: 32. 1865. Acalyphafiliformisvar.pervilleana (Baill.) Govaerts, World Checkl. Bibliogr. Euphorbiaceae 62. 2000. Type. Madagascar. Prov. Antsiranana: Nossibé, 1840, M. Richard 384 (lectotype, designated here: P [P00536745]!; isolectotype: P [P05604474]!, syn. nov.).
Syntypes. Madagascar, Prov. Antsiranana: Nossibé, M. Pervillé 368 P [P00536746!, P00536747!].
Acalyphahumblotiana Baill., Hist. Pl. Madag., Atlas (1891) t. 1891. Acalyphareticulatavar.urophyllaf.humblotiana (Baill.) Leandri, Notul. Syst. (Paris) 10: 262. 1942. Type: Comoros: Grande comore, 14 Nov 1885, L. Humblot 1461 (lectotype, designated here: P [P00196295]!; isolectotypes: P [P02712292!, P00196296!], syn. nov.).
Acalyphameiodonta Baill., Bull. Mens. Soc. Linn. Paris 2: 1197. 1895. Acalyphareticulatavar.meiodonta (Baill.) Leandri. Notul. Syst. (Paris) 10: 267. 1942. Type: Madagascar: Centr. Madag., Dec. 1883, R. Baron 2826 (lectotype, designated here: P [P05604378]!; isolectotype: K [K000186508]!, syn. nov.).
Syntype: Madagascar: “Centr. Madag.”, s.d., R. Baron 6581 K [n.v.].
Acalyphareticulatavar.meiodontaf.andronea Leandri, Notul. Syst. (Paris) 10: 268. 1942. Type: Madagascar, Prov. Mahajanga, Bemarivo (Boïna), Dec 1906, H. Perrier de la Bâthie 9561 (lectotype, designated here: P [P00513143]!; isolectotype: P [P00513144]!, syn. nov.).
Former syntypes: Madagascar, Prov. Mahajanga, bord du massif du Manongarivo, versant du Sambirano, 1909, H. Perrier de la Bâthie 9934 P [P00513149]!; Massif du Manongarivo, versant du Sambirano, Sep 1909, H. Perrier de la Bâthie 9939 P [P00513150]!; haut Bemarivo (Andranofosy), Boïna, Jan 1907, H. Perrier de la Bâthie 9635 P [P00513145]!; Massif du Manongarivo, Sambirano, Apr 1909, H. Perrier de la Bâthie 9928 P [P00513142]!; Manongarivo (Ambongo), Oct 1904, H. Perrier de la Bâthie 1677, P [P00513146!, P00513147!].
Distribution
Madagascar (Antsiranana, Mahajanga, Toamasina). Comoros Archipelago (Grande Comore, Anjouan, Mohéli and Mayotte).
References
Müller Argoviensis (1866[1]: 852) as A.reticulatavar.pervilleana; Müller Argoviensis (1882[2]: 26) as A.reticulatavar.urophylla; Müller Argoviensis (1866[1]: 852) as A.reticulatavar.pervilleana; Baillon (1892[3]: 1004) as A.pervilleana; Baillon (1895b[4]: 1197) as A.humblotiana; Palacký (1907[5]: 25, 26); Pax and Hoffmann (1924[6]: 105, 112) as Acalyphareticulatavar.urophylloides and A.reticulatavar.pervilleana; Leandri (1942[7]: 258, 260) as A.reticulatavar.pervilleana; Govaerts et al. (2000[8]: 62, 63, 75, 104 105, 107).
Notes
Acalyphapaxii was proposed by August De Candolle as a replacement name for the illegitimate A.urophylla Pax. Leandri (1942)[7] treated A.paxii as a synonym of A.reticulatavar.pervilleana. We consider that A.paxii is a well-differentiated species.
Acalyphapaxii, A.pervilleana and A.meiodonta have been considered synonyms of A.reticulata, which we consider to be a synonym of A.filiformis. A.paxii can be differentiated from A.filiformis by its sessile, dentate female bracts with a prominent central tooth vs. pedicellate, crenate to subentire female bracts in A.filiformis (see notes in A.filiformis).
Acalyphapaxii can be distinguished from A.urophylla, which has also been placed within A.reticulata, mainly by its leaves with usually caudate apices, its dentate female bracts with a prominent central tooth and eglandular margins vs. leaves with usually acuminate apices, dentate female bracts without a prominent central tooth and with small sessile glands at the margins. See notes under A.gracilipes Baill. for the differences between A.paxii and that species.
Acalyphahumblotiana was first illustrated, without description, in Baillon`s (1891)[9]Histoire Naturelle des Plantes, but this illustration is not consistent with Baillon`s (1895b)[4] later description nor with the specimen on which the description presumably is based (L. Humblot 1461).
Although Richard 384 [P00536745]! and 385 [P04779454]! have labels giving the locality as “Bourbon”, now Réunion, these are not Richard’s original labels and the species is otherwise unknown from the Mascarene Islands. We do not have evidence that it occurs there.
Taxon Treatment
- Muñoz, I; Cardiel, J; Levin, G; 2018: Nomenclatural review of Acalypha (Euphorbiaceae) of the Western Indian Ocean Region (Madagascar, the Comoros Archipelago, the Mascarene Islands and the Seychelles Archipelago) PhytoKeys, (108): 85-116. doi
Other References
- ↑ 1.0 1.1 Müller-Argoviensis J (1866) Acalypha. In: De Candolle AP (Ed.) Prodromus Systematis Naturalis Regni Vegetabilis. Vol 15(2). Paris, Treuttel et Würtz, 799−889.
- ↑ Müller-Argoviensis J (1882) Euphorbiaceen. In: Buchenau, Fr (Ed.) Reliquiae Rutenbergianae I (Botanik, Taf. I, II). Abhandlungen herausgegeben vom Naturwissenschaftlichen Verein zu Bremen 7: 24−30.
- ↑ Baillon H (1892) Liste des plantes de Madagascar (suite de la page 996).Bulletin Mensuel de la Société Linnéenne de Paris2(126): 1003–1005.
- ↑ 4.0 4.1 Baillon H (1895b) Liste des plantes de Madagascar (suite de la page 1181).Bulletin Mensuel de la Société Linnéenne de Paris2(151): 1193–1200.
- ↑ Palacký J (1907) Catalogus plantarum Madagascariensium II. Pragae, 1−89.
- ↑ Pax F, Hoffmann K (1924) Acalypha. In: Engler A (Ed.) Das Pflanzenreich. IV, 147-16 (heft 85), 1−231.
- ↑ 7.0 7.1 Leandri J (1942) Acalypha. Contribution a l'etude des Euphorbiacées de Madagascar (VI).Notululae Systematicae10: 252–290.
- ↑ Govaerts R, Frodin D, Radcliffe-Smith A (2000) World Checklist and Bibliography of Euphorbiaceae (and Pandaceae). Royal Botanic Gardens, Kew, 1−1621.
- ↑ Baillon H (1891) Histoire Naturelle des Plantes Vol. 34, Tome 4 (Atlas 2), 3ª partie, 27 fascicule. In: Grandidier A. (ed.) Histoire Physique, Naturelle et Politique de Madagascar. M. le Grade des Sceaux, à l’Imprimerie Nationale, Paris, 161–204.
