Tytthus pygmaeus (Henry, Thomas J. 2012)
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Ordo: Hemiptera
Familia: Miridae
Genus: Tytthus
Name
Tytthus pygmaeus Zetterstedt – Wikispecies link – Pensoft Profile
- Tytthus pygmaeus Henry, Thomas J., 2012, ZooKeys 220: 65-67.
Diagnosis
Diagnosis. This highly variable species is distinguished by the black head, with relatively vague interocular spots; entirely pale (European material) to entirely fuscous to black (all North American material) pronotum, with intermediate color forms; very clear or translucent white hemelytra; the fuscous to black antennal segment I, with the apical one fourth pale yellow; antennal segments II-IV uniformly fuscous to black; and the uniformly pale yellow legs. All males and most females of this species are fully macropterous; only a few weakly brachypterous females, with the membrane extending to the apex of the abdomen, have been examined. Tytthus pygmaeus is superfically similar to the Holarctic Tytthus pubescens in overall size, coloration, and distribution. It can be distinguished from Tytthus pubescens by the fuscous antennal segment I having only the apex pale, the short, recumbent setae on antennal segments I and II, and the variably fuscous to pale pronotum (but never fuscous with pale humeral angles as in Tytthus pubescens).
Description
Description. Male (n = 10) (Figs 41, 43): Length to apex of hemelytron 2.50-2.94 mm, length to base of cuneus 1.98-2.14 mm, width across hemelytra 0.94-0.98 mm. Head: Length 0.29-0.32 mm, width across eyes 0.69-0.75 mm, interocular width 0.30-0.32 mm. Labium: Length 1.07-1.12 mm. Antenna: Segment I length 0.30-0.32 mm, II 0.98-1.04 mm, III 0.61-0.62 mm, IV 0.54-0.58 mm. Pronotum: Length 0.35-0.40 mm, basal width 0.77-0.82 mm. Coloration: Head: Shiny fuscous to black, with a relatively small, yellow, interocular spot near inner margin of each eye; eyes fuscous to dark reddish brown. Labium: Uniformly pale yellow, with apex of segment IV usually brown. Antenna: Segment I, fuscous to black, with only apical one fourth to one third pale yellow; segment II-IV black. Pronotum: Highly variable, ranging from uniformly pale yellow to entirely fuscous to black, with many intermediate forms, including all of disc dark and anterior half yellow and almost entirely dark with yellow across anterior margin and through middle of calli; some yellow specimens with only fuscous posterior angles. Mesoscutum and Scutellum: Uniformly fuscous to black. Hemelytron: Uniformly clear to very pale translucent white. Ostiolar evaporative area: Fuscous to black, even on palest specimens. Ventral surface: Thorax uniformly fuscous; abdomen fuscous along lateral margins and genital capsule, ventral area and sides pale yellow to whitish. Legs: Uniformly pale yellow. Structure, texture, and vestiture: Head: Shiny, impunctate, much broader than long; set with numerous, long, erect and semierect setae on vertex and frons. Labium: Extending to apices of meso- or bases of metacoxae; segment I extending just beyond head to anterior margin of xyphus just before procoxae. Antenna: Segment I sparsely set with recumbent setae and two, long, erect, subapical, bristlelike setae; segment II, thickly set with short, recumbent and semierect setae. Pronotum: Trapeziform, anterior angles nearly weakly rounded; lateral margins straight, gradually widening to posterior angles; posterior margin nearly straight or only weakly sintuate. Mesoscutum: Broadly exposed. Scutellum: Equilateral; sparsely set with scattered relatively short, semierect setae. Hemelytra: Macropterous, cuneus and membrane fully developed, extending well beyond apex of abdomen; evenly set with recumbent setae. Male genitalia: Left paramere (Fig. 175): Right arm long, broad, apically blunt; left arm short, apically pointed. Right paramere (Fig. 176): Elongate oval. Endosoma (Fig. 177): Strongly C-shaped, apically blunt. Phallotheca (Fig. 178): Slender, apically acute. Female (n = 10) (Figs 42, 44): Length to apex of hemelytron 2.92-3.17 mm, length to base of cuneus 2.21-2.34 mm, width across hemelytra 1.20-1.31 mm. Head: Length 0.34-0.35 mm, width across eyes 0.74-0.75 mm, interocular width 0.34-0.35 mm. Labium: Length 1.12-1.20 mm. Antenna: Segment I length 0.27-0.29 mm, II 0.86-0.88 mm, III 0.61-0.62 mm, IV 0.54-0.58 mm. Pronotum: Length 0.40-0.42 mm, basal width 0.91-0.96 mm.
Distribution
Distribution. This Holarctic species is known in Canada from Alberta, British Columbia, Labrador, Newfoundland, Ontario, Saskatchewan, and Yukon Territory and in the United States from New Mexico and Wyoming (Kelton 1980, Henry and Wheeler 1988, Wheeler and Henry 1992, Maw et al. 2000, Wheeler 2011). Colorado is a new state record. In the Old World, it is known in Asia from Russia (East Siberia, Far East, and West Siberia) and in Europe from Austria, Byelorussia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Hungary, Latvia, Luxembourg, Moldavia, The Netherlands, Norway, Poland, Russia (Central European and North European territories), Slovakia, Sweden, and the United Kingdom (Carvalho and Southwood 1955, Wheeler and Henry 1992, Kerzhner and Josifov 1999).
Taxon Treatment
- Henry, Thomas J.; 2012: Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae) ZooKeys, 220: 65-67. doi
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