Trichosiopsis tuberculigera
Ordo: Diptera
Familia: Sciaridae
Genus: Trichosiopsis
Name
Trichosiopsis tuberculigera Tuomikoski, 1960 – Wikispecies link – ZooBank link
- Trichosiopsis tuberculigera Tuomikoski, 1960[1]: 29, fig. 5f
- Leptosciarella (Trichosiopsis) tuberculigera (Tuomikoski, 1960)[2]: 82-83, fig. 33
Type material
Holotype ♂, in MZH, leg. Tuomikoski, 16.06.1958
Type locality
Finland, Vihti, Vihtijärvi
Material studied
Finland: 1 ♂, Sa, Utula, Lake Simaa, Kyläniemi; spruce/birch/pine forest, sweep netting, Jaschhof, 20.-21.06.2004, PKHH 6223; 1 ♂, Ta, Seitseminen N.P., Kuru, N Pitkajärvi: birch/pine swamp forest, sweep netting, Jaschhof, 05.06.2004, PKHH 6354; 2 ♂, Ab, Salmi, Vihti, bushes along forest edge, sweep netting, Jaschhof, 27.05.2004, PASS 182, ZSMC 2438; 1 ♂, St, Huittinen, Mutilahti: birch forest with rowan, spruce, pine, sweep netting, Jaschhof, 01.06.2004, MZH 9049; Sweden: 2 ♂, Darlana, Orsa, Österaberget Nat. Res.; mixed forest (spruce, birch, willow), Malaise trap, Jaschhof & Jaschhof, 3.7.-17.08.2005, PKHH 5980, NHRS 1121; 6 ♂, Härjedalen, Nyvallen, Nyvallens fäbod: alpine birch and spruce wood, Malaise trap, Swedish Malaise Trap Project, 4.7.-04.08.2004, NHRS 1703, 3264, 3265, 3423, 4340, 4825.
Description (male)
Head. Eye bridge 4–5 rows of facets. LW-index of 4th antennal flagellar segment 1.7–2.1; neck 0.2–0.3 of segment width; Transition of basal part to neck pronounced. Colour of neck unicolour. Antennal hairs shorter than segment width; dense; adjacent. Palps bright; normal; palpomeres 3. First palpomere of normal shape; with 2–3 bristles; with only some sparse sensillae. Second palpomere shortly oval. Third palpomere as long as first segment. Thorax. Colour reddish. Notum unicolorous. Thoracic setae normal; darkened, or bright. Mesonotum with some weaker central bristles. Posterior pronotum setose. Postpronotal setae 2–6; fine. Laterotergite bare. Legs. Colour yellow. Hind coxae of same colour as femora. Hairs on fore coxae bright. Front tibia apically with a patch of setae. Front tibial organ bright. Front tibial organ not bordered. Tibial setae on hind legs weak, inconspicuous. Tibial spurs of equal length. Claws untoothed. Wings. Wings slightly darkened; of normal shape. Wing membrane without macrotrichia. Wing venation weak, with faint m-base. M-fork of normal shape. R1 inserting clearly before base of m-fork; posterior veins bare; bM bare; r-m with a few setae; bM:r-M 0.75–0.95; st-Cu:bM 0.3–0.45; r1:r 1.05–1.35; C:w 0.6–0.75. Halteres dark; of normal length. Abdomen. Abdominal setae weak; dorsally white; ventrally white. Hypopygium bicolour; 0.48–0.65 × longer than wide. Base of gonocoxites with normal, weak hairs; gonocoxites broadly separated; inner margin of gonocoxites normally U-shaped; inner part of hypopygium densely setose; elongated setae on valves of hypopygium absent. Gonostylus globular; 1.6–2 × longer than wide; Inner margin with complex structure; apex equally rounded. Apical tooth present; 2–2.5 × longer than broad; weak. Awl-like setae normal; below apex absent. Megasetae on inner part of gonostylus absent. Whiplash-hair absent. Tegmen 0.42–0.6 × longer than broad; rectangular with rounded edges, or rectangular with sharp edges; normal; Central process absent. Length of aedeagus/hypopygium 25–35 %; Aeadeagal apical structure absent. Measurements. Body size 2.4–2.9 mm. Hind tibia 1.3–1.5 mm. Wing length 2.2–2.7 mm.
Diagnosis
Trichosiopsis tuberculigera is a small species and differs from most other Trichosiopsis by the lacking macrotrichia on the posterior wing veins and the unusually shaped gonostylus.
DNA Barcoding
The COI sequence is assigned to BIN BOLD:ACU4750 (average distance 0.1%, max. 0.15%, n=3, K2P: 7.22%).
Discussion
Trichosiopsis tuberculigera was the only species of the montypic genus Trichosiopsis. It was later allocated in a separate subgenus by Mohrig & Menzel (1997)[2] for the lacking apical tooth on the gonostylus. However Vilkamaa & Salmela (2005)[3] observed, that there is indeed a very tiny tooth present on the inner middle lobe. This indicates, that the inner lobe is homologous to the apex of normal Trichosiopsis and the upper lobe is only a luxury structure formed by the dorsal part of the gonostylus, just as in some species of Leptospina or in some species of Sciara. Therefore the concept of Trichosiopsis is now broadened to include as well all other species of the former subgenus Leptosciarella s. str. sensu Mohrig & Menzel (1997).
Etymology
tuber lat. = hump; gerere lat. = bear. The species name refers to the tubercle-like structures on the inner side of the gonostylus.
Ecology
Until now Trichosiopsis tuberculigera has been found only in Scandinavia northern to 60°N, so it seems to be a boreal species, with a restricted distribution, but locally not rare.
Distribution
Finland[1][3], Norway[4], Sweden[5].
Images
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References
- ↑ 1.0 1.1 Tuomikoski, R. 1960: Zur Kenntnis der Sciariden (Dipt.) Finnlands. Annales Zoologici Societatis Zoologicae Botanicae Fennicae “Vanamo”, 21, 1–164.
- ↑ 2.0 2.1 Mohrig, W.; Menzel, F. 1997: Revision der paläarktischen Arten von Trichosia Winnertz sensu Tuomikoski, 1960 (Diptera, Sciaridae). – Teil II. Gattungen Leptosciarella Tuomikoski, 1960 und Trichodapus gen. nov. Studia dipterologica, 4(1), 41–98.
- ↑ 3.0 3.1 Salmela, J.; Vilkamaa, P. 2005: Sciaridae (Diptera) from central Finland: faunistics and taxonomy. Entomologica Fennica, 16, 287–300.
- ↑ GBIF Trichosiopsis tuberculigera
- ↑ Heller, K.; Vilkamaa, P.; Hippa, H. 2009: An annotated check list of Swedish black fungus gnats (Diptera, Sciaridae). Sahlbergia, 15(1), 23-51.
