Trichomycterus trefauti
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Ordo: Siluriformes
Familia: Trichomycteridae
Genus: Trichomycterus
Name
Trichomycterus trefauti Wosiacki, Wolmar Benjamin, 2004 – Wikispecies link – Pensoft Profile
- Trichomycterus trefauti Wosiacki, Wolmar Benjamin, 2004, Zootaxa 592: 3-9.
Materials Examined
Holotype: MZUSP79911, 49.5 mm SL, riacho Andrequicé, tributary of rio Paraúna, itself a tributary of rio das Velhas (rio São Francisco basin), approximately 18 ° 30 ’S, 43 ° 30 ’W, Município de Trinta Réis, Minas Gerais, Brazil; collected by M.T.U. Rodrigues; 8 September 1986. Paratypes: Collected with the holotype: MZUSP 36966, 5 ex. (36.6–54.2 mm SL; 3 C&S); MPEG 7896, 2 ex. (37.2–47.3mm SL).
Diagnosis
Diagnosis: Trichomycterus trefauti differs from all other known members of the subfamily Trichomycterinae by the autapomorphic presence of an elliptical, vertically elongated, brown spot, at caudalfin base (vs. without elliptical spot at caudalfin base), and the combination of homogeneously gray color pattern (vs. yellowish, presence of stripes or bands, or lack of color pattern), first pectoralfin ray prolonged as a filament (vs. not prolonged), subterminal mouth (vs. inferior or terminal mouth), two supraorbital pores at interorbital space (vs. one supraorbital pore at mesial line), caudal fin truncate with attenuated edges (vs. caudal fin rounded), pelvic fins covering anus and urogenital openings (vs. not covering), interorbital space very wide — 39.8–45.9 % HL (vs. more or less than 39.8– 45.9 % HL), maxillary barbels very long — 84.2 –93.0 % HL (vs. more or less than 84.2 – 93.0 % HL), rictal barbels very long — 67.6–74.3 % HL (vs. more or less than 67.6–74.3 % HL).
Description
Description: Morphometric data for holotype and paratypes are given in Table 1. Body elongate, roughly cylindrical close to head and gradually more compressed towards caudal fin. Dorsal and ventral profiles of trunk slightly convex. Dorsal and ventral profiles of caudal peduncle straight (Fig. 2). Integument thick, especially over base of pectoral and caudal fins. Small papillae on lips, and scattered on dorsal surface of head. Head wide and depressed, trapezoidal, slightly longer than wide, transverse section at posterior tip of opercle wider than anteriorly at nostril, anterior margin slightly rounded. Head lateral to eye slightly swollen by jaw muscles in large and small specimens. Dorsal and ventral profiles of head convex. Eyes rounded, dorsolaterally oriented. Eye covered by thin skin, transparent at its center, gradually opaque towards rim, distinctly separated from surface of eyeball. Ocular structures readily visible on surface of skin, not deeply sunken. Orbital rim not free. Anterior nare slightly larger than posterior, surrounded by fleshy flap of integument. Posterior nare surrounded anteriorly by thin flap of integument. Anterior and posterior nares about one third of eye diameter. Gill membranes thick, united to isthmus only at anteriormost point, forming small free fold across the isthmus. Gill openings not constricted. Branchiostegal rays 6–8 (6 in Holotype) visible from below (7–8 in C&S). Mouth subterminal, its corners laterally oriented. Lower lip with conspicuous lateral fleshy lobes, internal to origin of rictal barbels. Anterior margin of upper lip slightly rounded. Small papillae on external surface of upper lip and large papillae inside mouth at region of teeth attachment. Barbels long (nasal 74.3 – 69.8; maxillary 93.0– 84.2; and rictal barbel length 74.3 – 67.6 % of head length). Barbels with broad bases, gradually narrowing towards tip. Nasal barbels reaching median odontodes of interopercle; maxillary barbels reaching base of pectoral fin; rictal barbels reaching last interopercular odontode. Origin of nasal barbels on posterolateral portion of integument flap around anterior nostril. Interopercular patch of odontodes rounded, with 34–36 conical odontodes. Opercular patch of odontodes rounded, with 14–16 conical odontodes. Supraorbital canal complete and infraorbital incomplete. Infraorbital anterior section pores i 1 and i 3, and posterior section pores i 10 and i 11. Supraorbital pores s 1, s 2 and s 4. Two pores s 4 at interorbital space. TABLE1. Morphometric data for Trichomycterus trefauti based on the holotype and four paratypes. Holotype Range Mean Standard length (mm) Percents of standard length54.3 54.3–37.2Head length18.6 20.0–18.2 19.3Predorsal length66.5 66.6–65.3 66.1Prepelvic length61.3 63.0–59.7 61.8Preanal length70.9 93.3–68.9 75.1Pectoral girdle width14.7 17.4–14.2 15.3Trunk length46 46.0–41.9 44Pectoralfin length15.1 16.1–13.7 14.9Pelvicfin length7.4 9.1–7.4 8.3Pelvicfin base anus distance5.3 5.9–4.2 5.1Caudal peduncle length21.5 21.5–19.0 20.1Caudal peduncle depth10.5 11.9–9.5 10.8Body depth14.7 15.5–12.6 14.5Dorsalfin length9.8 12.3–9.8 10.8Analfin length9.6 11.3–8.9 9.8Percents of head length Head width92.1 96.5–82.4 92.1Nasal barbel length74.3 74.3–69.8 72.4Maxillary barbel length84.2 93.0–84.2 87.2Rictal barbel length74.3 74.3–67.6 71.1Snout length40.6 45.3–39.8 42.7Interorbital width40.6 45.9–39.8 43Mouth width30.7 36.1–29.7 32.1Eye diameter15.8 18.9–14.5 15.9Counts Dorsalfin rays II7II7Pectoralfin rays I6I6Pelvicfin rays5 5Analfin rays I6I6Caudalfin rays I11I I11I Pectoralfin margin truncate, I/ 6 rays, first ray longest with filamentous extension. Dorsal fin with margin semicircular when expanded, II/ 7 rays, third and fourth rays longest. Anal fin slightly elongate in overall shape, smaller than dorsal fin, I/ 6 rays, third ray longest, origin at vertical through sixth dorsalfin ray. Pelvic fins with origin anterior to dorsalfin origin, rounded margin beyond urogenital and anal openings when extended, 5 rays, second and third rays longest. Caudal fin truncate with attenuated margin edges, distinctly wider than remaining caudal region, I/ 11 /I principal rays, principal central rays splitting once and dorsal and ventral principal rays splitting twice. Only first dorsal and ventral caudalfin accessory rays visible. Anal and urogenital openings midway between pelvic fin base and analfin origin. Free vertebrae 37. Ribs 13–14 pairs, first thickest, second to 12–13 th pairs slightly longest, the last pair rudimentary and free. Dorsal pterygiophores 8, first in front of neural spine of 21 th free vertebrae. Anal pterygiophores 6, first in front of haemal spine of 24 th free vertebrae. Procurrent rays of dorsal lobe 14–16. Procurrent rays of ventral lobe 12–16. Caudal skeleton pleurostyle, hypurals 4 + 5, hypural 3, and fused parahypural and hypurals 1 + 2. Color in alcohol: Refer to figures 2 for general view of color pattern in alcohol. Uniform gray on dorsal surface and sides of head, trunk and caudal peduncle, darker at nape. Dark stripe from eyes to anterior nostril. Elliptical, vertically elongated dark spot with illdefined borders at base of caudal fin. Few irregular spots on caudal peduncle in large specimens (48.9–54.3 mm SL) and on trunk and caudal peduncle in small (37.2–42.4 mm SL) specimens. Ventral surface of head and abdominal region without pigmentation. Dorsal surface of pectoralfin base with few chromatophores, gradually scattered towards margin. Ventral surface of pectoral fin and pelvic fins unpigmented. Bases of dorsal and anal fin with chromatophores, more scattered towards margin over rays. Dorsal fin darker than anal fin. Chromatophores on base of caudalfin rays, gradually more scattered towards margin. Nasal barbels with chromatophores all over their surface; maxillary barbels with chromatophores on their dorsal surface. Rictal barbels unpigmented. Internal surface of all barbels lighter than outer margin.
Distribution
Distribution: Known only from the typelocality.
Etymology
Etymology: Named after Dr. Miguel Trefaut Rodrigues, who discovered and collected the first specimens of the species. Systematic and taxonomic comments: The genus Trichomycterus has been recognized as nonmonophyletic (Arratia, 1990, 1998, de Pinna, 1989; 1998), as is also true for the subfamily Trichomycterinae (de Pinna, 1989; 1998). Arratia (1990) proposed four “unique derived characters” for Trichomycterinae (1 “basioccipital”…” with well developed anterior membranous processes which lie ventrolateral to parasphenoid and prootics”, 2 “an enarthrodial articulation between preopercle and opercle”…” is present in adults”, 3 “vomer with only one long posterior process”, and 4 “pronounced notch”…” on the posteroventral margin of ceratobranchial”). These features are present in all analyzed Trichomycterinae species prepared for C&S and Xray, including Trichomycterus trefauti. However, a more recent analysis (Costa & Bockmann, 1993) proposed Scleronema and the genus Ituglanis composed of species previously located in Trichomycterus, as two sequential monophyletic groups more related to the Glanapteryginae, Sarcoglanidinae, Tridentinae, Stegophilinae, and Vandelliinae, than to the other trichomycterine species. The lack of knowledge about the relationships among trichomycterine species, absence of definition of the genera Trichomycterus and Eremophilus, and the superficial definition of Hatcheria, Bullockia, Silvinichthys, and Rhizosomichthys make the allocation of new species difficult. Unfortunately, Trichomycterus has been considered as a group of species that do not possess the features present in other genera of trichomycterine. For this reason Trichomycterus has grown to a total of 96 nominal species (Barbosa & Costa, 2003; Fernández & Schaefer, 2003; de Pinna & Wosiacki, 2003; and Wosiacki & Garavello, 2004) in contrast with the other genera Eremophilus, Bullockia, Hatcheria, Rhizosomichthys, and Silvinichthys with only one species each. The lack of pelvic fins and girdle has been used to diagnose Eremophilus, the only recognized distinction from Trichomycterus. These features were observed and discussed by Myers (1944), Miranda Ribeiro (1949), de Pinna (1989, 1998), Costa & Bockmann (1993), Trajano & de Pinna (1996), Fernández & Vari (2000), and de Pinna & Wosiacki (2003). These papers have demonstrated that the lack of pelvic fins and girdle are highly homoplastic and for this reason, weak features to define a genus. Recently de Pinna & Wosiacki (2003) reallocated Eremophilus candidus MirandaRibeiro to the genus Trichomycterus (T. candidus) based on detailed phylogenetic analysis which indicates that it is more related to Trichomycterus species than to E. mutisii Humboldt. Arratia et al. (1978) proposed the genus Bullockia based on several features slightly similar to those of Hatcheria diagnosed in the same paper and in Arratia & MenuMarques (1981).These features are clearly distinct from those observed in species of Trichomycterus. Characters like a narrow and strongly compressed caudal peduncle and a long dorsal fin with more than 17 rays are present in Hatcheria and Bullockia but are not present in species of Trichomycterus. Silvinichthys differs by having the entire surface of body skin perforated by pores of ampullary organs and a unique combination of characters (Arratia, 1998). Rhizosomichthys totae (Miles) is an atypical freshwater fish that has the unique character of thick fat tissue forming folds covering the entire body. Its osteology is unknown, and its relationship among trichomycterine is totally obscure. Trichomycterus trefauti does not have the features above mentioned for Eremophilus, Bullockia, Hatcheria, Rhizosomichthys, and Silvinichthys and can not be allocated to any of these genera. In order to maintain nomeclatural stability it does not seem desirable to propose a new genus for one new species. It seems more parsimonious and coherent to allocate the new species in Trichomycterus. The majority of the species of Trichomycterus are morphologically very similar to each other and the diagnosis of most species is only possible by a combination of several characters. Sometimes these features concern to internal anatomy and are difficult to analyze. Few species, like Trichomycterus castroi de Pinna, T. papilliferus Wosiacki & Garavello, T. trefauti have external autapomorphies that make them easy to identify Phylogenetic assignment and comparisons: The presence of stripes or bands in Trichomycteridae has been frequently recorded (e.g., Scleronema operculatum (Eigenmann), T. barbouri (Eigenmann), T. castroi, T. itatiayae MirandaRibeiro, T. reinhardti (Eigenmann), T. taeniops Fowler, T. taenia Kner). A transverse band at the caudal fin has been recorded only in S. operculatum, T. castroi, and T. trefauti. The presence of a vertical caudalfin band in T. castroi and S. operculatum was discussed by de Pinna (1992 a) who interpreted these as autapomorphies for these species. The spot at the caudalfin base of Trichomycterus trefauti has a different topology from the band of T. castroi that is at the distal margin of the caudal fin. In addition, the vertical spot at the caudalfin base distinguishes T. trefauti from its sympatric congeners in the rio São Francisco basin (T. brasiliensis Lütken, T. concolor Costa, T. itacarambiensis Trajano & Pinna, T. reinhardti, T. variegatus Costa), and all other trichomycterins species. The spot at the caudalfin base of T. trefauti is unique within trichomycterids and it is hypothesized to be an autapomorphy for this species. The first pectoralfin ray prolonged as a filament, present in T. trefauti, is present in all species of Ituglanis, Trichomycterus hasemani (Eigenmann), T. johnsoni (Fowler) and several other species (T. candidus, T. alternatus (Eigenmann), T. punctulatus Valenciennes, T. brasiliensis, T. reinhardti, T. itacarambiensis, T. auroguttatus Costa, T. longibarbatus Costa, T. bahianus Costa, T. mimonha Costa, T. mirissumba Costa, T. itatiayae Miranda Ribeiro, and T. immaculatus (Eigenmann & Eigenmann)) among others. According to Costa & Bockmann (1993), Ituglanis is a monophyletic sister group of the Glanapteryginae, Sarcoglanidinae, Tridentinae, Stegophilinae, and Vandelliinae, and T. hasemani and T. johnsoni could be more closely related to other subfamilies (de Pinna, 1989). Therefore it is hypothesized that the first pectoralfin rays prolonged as a filament is a homoplastic feature that has evolved several times in Trichomycteridae. Ecology: The specimens were collected in a stream 2–4m wide, with an average depth of 10cm, and with a bottom of pebbles of variable size and rocks, at an elevation of approximately 1000m. The vegetation was of “campos rupestres”.
Materials Examined
Comparative material Copionodon pecten MZUSP48962 (7 ex. Alc. 3 ex. C&S); MZUSP42462 (5 ex.) Paratypes; C. orthiocarinatus MZUSP42464 (1 ex.) Paratype; C. lianae MZUSP42470 (2 ex.) Paratypes; MZUSP39995 (1 ex. C&S); Glaphyropoma rodriguesi MZUSP42466 (3 ex.) Paratypes; Trichogenes longipinnis MZUSP40238 (2 ex); MZUSP63478 (15 ex. Alc., 3 ex. C&S); Bullockia maldonadoi CAS63842, Holotype; CAS63843 (43 ex.) Paratypes; MZUSP36958 (1 ex. C&S); Eremophilus candidus MNRJ11762 (2 ex. C&S) Paratypes; E. mutisii MZUSP35409 (1 ex. C&S) AMNH56092 (1 ex. C&S); CAS6595 (13 ex.); Hatcheria macraei MZUSP35687 (3 C&S); H. patagoniensis CAS63844 (2 ex.) Paratypes; H. titcombi CAS28557, Holotype; Ituglanis amazonicus MUSP30449 (1 ex.); I. eichorniarum MZUSP37763 (1 ex.); MNRJ780 (2 ex.) Syntypes; I. gracilior FMNH53264 (Xray) Holotype; I. guianensis FMNH52676 (Xray) Holotype; I. laticeps CAS32458 (1 ex.); I. metae CAS118214 (1 ex.); CAS124227 (1 ex.); CAS58138, Holotype; I. parahybae FMNH58576 (Xray) Holotype; I. proops MZUSP84190 (12 ex. Alc, 2 ex. C&S); MNRJ781 (3 ex.) Syntypes; Ituglanis n. sp.MNRJ11489 (13 ex. 5 C&S); Pygidium angustirostris MNRJ3605 (2 ex.) Syntypes; P. f l o re n s e MNRJ3751, Holotype; P. paquequerensis MNRJ1159, Holotype; P. parkoi MNRJ3849, Holotype; P. travassosi MNRJ5424, Holotype; Rhizosomichthys totae SU 37074 (2 ex.) Paratypes; Scleronema minutum MCP11169 (3 ex. C&S); S. operculatum FMNH58080 (Xray) Holotype; MCP9315 (1 ex.); Scleronema n. sp.UFRGS3955 (2 ex.); Trichomycterus albinotatus MZUSP35817 (2 ex.); T. alternatus CAS64575 (4 ex.) Paratypes; FMNH58082 (Xray) Holotype; FMNH58083 (62 ex. 2 Xray) Paratypes; MZUSP67913 (5 ex.); MZUSP52541 (4 ex. Alc., 1 ex. C&S); T. areolatus CAS15649 (47 ex.); T. atochae CAS64576, Holotype; CAS64577 (4 ex.) Paratypes; USNM301837 (5 ex.); T. auroguttatus MZUSP84191. (1 ex.); T. bahianus MZUSP38636 (1 ex.) Paratype; T. banneaui CAS58127 (26 ex.) Paratypes; FMNH56025 (Xray) Holotype; T. barbouri CAS64578 (1 ex.) Paratype; FMNH53946 (5 ex. 2 Xray). Syntypes; ROM – OR# 403409 (1 ex.); ROM51722 (1 ex.); T. bogotense CAS58118 (27 ex.) Paratypes; FMNH56030 (Xray) Holotype; FMNH56031 (264 ex. 2 Xray) Paratypes; T. boylei MZUSP84192 (2 ex.); T. brasiliensis MZUSP37145 (2 ex.); T. caliense CAS132081 (3 ex.); FMNH56029 (Xray) Holotype; T. candidus MPEG6711 (1 ex.); MZUSP79910 (4 ex.); MPEG6712 (5 ex.); MHNCI8075 (2 ex.); T. castroi NUPELIA uncat. (1 ex.– Rio Iguaçu, Segredo, PR); NUPELIA uncat. (1 ex.– Rio Iguaçu, Segredo, PR); MHNCI7643 (2 ex. 1 C&S); T. chapmani FMNH56027 (Xray) Holotype; FMNH56028 (10 ex. 2 Xray) Paratype; FMNH69813 (16 ex. 2 Xray) Paratypes; CAS58128 (8 ex.) Paratypes; T. chiltoni ANSP91039 (1 ex.); CAS57596, Holotype; CAS57597 (68 ex.) Paratypes; CAS57598 (4 ex.) Paratypes; T. conradi CAS58257 (ex. 1) Paratype; FMNH53721 (Xray) Holotype; T. cubataonis MNRJ12490, Holotype; MNRJ12491 (9 ex.) Paratypes; T. davisi FMNH54242 (9 ex. 2 Xray) Syntypes; FMNH54242 (9 ex.) Syntypes; FMNH60309 (Xray) Holotype; MCP10646 (2 ex.); T. dorsostriatus CAS64579 (2 ex.) Paratypes; FMNH58096 (Xray) Holotype; FMNH58097 (Xray) Paratype; T. duellmani UMMZ204202 (3 ex.); T. eichorniarum MNRJ780 (2 ex.) Syntypes; T. emanueli (CAS147810); T. gabrieli CAS64583 (4 ex.) Syntypes; T. gabrieli SU 36556 (1 ex.) Syntype; T. hasemani CAS54584 (36 ex.) Paratypes; FMNH56424 (Xray) Holotype; FMNH56425 (2 Xray) Paratypes; MZUSP23600 (5 ex. C&S); T. heterodontus CAS58139 (1 ex.) Paratype; T. iheringi CAS64585, Holotype; CAS64586 (3 ex.) Paratypes; FMNH58074 (2 ex.) Paratypes; T. itacarambiensis MZUSP67914 (3 ex. C&S); MZUSP40329 (5 ex.); T. johnsoni MZUSP59367 (1 ex.); T. latidens CAS76335, Holotype; T. latistriatus FMNH58449 (Xray) Holotype; T. longibarbatus MZUSP67915 (3 ex. C&S); T. mboycy MPEG6695Holotype; MPEG6696 (2 ex.) Paratypes; MPEG6697Paratype; MPEG6698 (2 ex.) Paratypes; T. meridae CAS147809 (14 ex.); UMMZ145374 (5 ex.); T. mimonha MZUSP49486 (3 ex. C&S); T. mirissumba MZUSP44500 (3 ex. C&S); T. naipi MPEG6699Holotype, MPEG6700 (2 ex.) Paratypes, MPEG6701Paratype, MPEG6702 (2 ex.) Paratypes; T. oroyae CAS58104 (43 ex.); T. paolence FMNH58085 (Xray) Holotype; FMNH58575 (Xray) Paratype; T. papilliferus MPEG6692Holotype; MPEG6693 (4 ex.) Paratypes; MPEG6694 (3 ex.) Paratypes; NUPELIA 1615Paratypes; MPEG6703Paratype; T. paquequerense MNRJ1159, Holotype; T. pardus ROM51717 (3 ex.); T.plumbeus MPEG6686Holotype; MPEG6688 (20 ex.) Paratypes; MPEG6687 (2 ex.) Paratypes; NUPELIA 1614 (3 ex.) Paratypes; T. proops MNRJ781 (3 ex.) Syntypes; T. punctulatus ROM51725 (2 ex.); CAS58094 (4 ex.); T. punctulatus piurae CAS58119, Holotype; CAS58120 (14 ex.) Paratypes; FMNH77904 (5 ex. 2 Xray) Paratypes; FMNH58672 (3 ex. 2 Xray) Paratypes; T. quechuorum AMNH2035 /sw (1 ex.); T. regani CAS64591, Holotype; T. reinhardti FMNH58081 (Xray) Holotype; MZUSP uncat. (2 ex.Rio Bonito, Lauras, MG); T. rivulatus CAS58092 (2 ex.); CAS58093 (3 ex.); ROM40769 (1 ex.); T. santaeritae FMNH58577 (Xray) Holotype; MNRJ14380 (3 ex. 1 C&S); T. septentrionale CAS77008 (3 ex.); CAS77009 (3 ex.); FMNH59522 (Xray) Holotype; FMNH59195 (Xray) Paratype; T. stawiarski NUPELIA uncat. (1 ex. Rio Jordão, Segredo, PR); T. stellatus CAS58121 (3 ex.) Paratypes; FMNH58100 (3 ex. 2 Xray) Paratypes; FMNH58101 (Xray) Holotype; FMNH58102 (3 ex. 2 Xray) Paratypes; T. stramineus CAS58105 (4 ex. 1 Xray) Paratypes; CAS58148 (1 ex.) Paratype; FMNH58092 (3 ex. 1 Xray) Paratype; FMNH58105 (Xray) Holotype; T. striatum ANSP104215 (1 ex.); T. striatus CAS64593 (1 ex.) Paratype; FMNH26660 (1 ex.), 26661 (Xray), 26662 (Xray) Paratype; FMNH55233 (Xray) Paratype; FMNH7579 (Xray) Holotype; T. taczanowskii CAS76966 (12 ex.); T. taroba MPEG6689Holotype; MPEG6690 (15 ex.) Paratypes; MPEG6691 (10 ex.) Paratypes; NUPELIA 1616 (3 ex.) Paratypes; T. travassosi MNRJ5424, Holotype; T. triguttatus FMNH58670 (Xray) Holotype; FMNH58671 (3 ex. 2 Xray) Paratypes; CAS64596 (1 ex.) Paratype; T. unicolor ROM51718 (3 ex.); T. vermiculatus FMNH58077 (Xray) Holotype; MZUSP67912 (4 ex. C&S); T. zonatus FMNH58572 (1 ex. Rray) Paratype; FMNH58573 (Xray) Holotype; FMNH58574 (2 ex. 1 Xray) Paratypes; MZUSP67913 (6 ex. C&S); T. cf. nigricans MCP10649 (1 ex. C&S); Trichomycterus sp. n. (Bahia) MZUSP67910; Trichomycterus sp. n. (Mato Grosso) MZUSP67905 (1 ex.); MZUSP67906 (1 ex.); MZUSP67909 (1 ex.); Trichomycterus sp. n. (“pelvic less”) MPEG uncat. (1 ex. – Rio Roseira, Concórdia, RS); Vandellia beccarii AMNH55625 sw (1 ex.); V. cirrohosa AMNH20497 (1 ex.); V. i x MZUSP29155 (2 ex.) Paratypes; V. plazai USNM302683 (1 ex.); Paracanthopoma sp. MZUSP30401 (2 ex.); Homodiaetus haemomyzon ANSP137581 (1 ex.); H. anisitsi MCP9987 (1 ex.); Pseudostegophilus nemurus MZUSP30431 (2 ex.); Pareiodon sp. MZUSP32522 (3 ex.); Stegophilus passarelli UFRJ 0 653 (2 ex.); S. insidiosus MZUSP40048 (1 ex.); Henonemus sp. MZUSP30414 (2 ex.); Acanthopoma annectens MZUSP30420 (1 ex.); A. bondi MZUSP30418 (2 ex.); Megalocentor echthrus MZUSP30391 (1 ex.); M. echthrus MZUSP30380 (1 ex.); Parastegophilus maculatus MZUSP35736 (2 ex.); Ochmacanthus orinoco MZUSP30477 (2 ex.); O. alternus MZUSP30473 (3 ex.); Haemomaster venezuelae MZUSP30375 (1 ex.); Listrura nematopteryx MZUSP37138 (2 ex.). Tridensimilis venezuelae USNM121291 (2 ex.); Tridentopsis sp. MZUSP63092; Sarcoglanis simplex INPA8165 (2 ex.).
Taxon Treatment
- Wosiacki, Wolmar Benjamin; 2004: New species of the catfish genus Trichomycterus (Siluriformes, Trichomycteridae) from the headwaters of the rio São Francisco basin, Brazil, Zootaxa 592: 3-9. doi
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