Pupinidius pupinella altispirus (Wu, Min & Zheng, Wei 2009)
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Ordo: Stylommatophora
Familia: Enidae
Genus: Pupinidius
Species: Pupinidius pupinella
Name
Pupinidius pupinella altispirus Moellendorff, 1901 – Wikispecies link – Pensoft Profile
- Pupinidius pupinella Wu, Min, 2009, Zootaxa 2053: 19-20.
Description
Figs 1, 6 D–H, 8 G, 15
Materials Examined
Material examined. HBUMM-05723 (20 shell measured, 3 with broken apex; specimen- 1 for dissection; 1 young): 1661 m a.s.l., 34 °0΄ 127.6 ΄ N, 104 ° 25 ΄ 16.4 ΄ E, Dangchang, Gansu, loessial area, 2006 -X- 4, coll. J.Liu & W.Zheng; ZI RAS-N 602: type material, det. Moellendorff, 1885 -VI- 19, coll.? Potanin. SMF- 41990: Lectotype. Tan-tshang, Kansu, China. Slg. O.v.Moellendorff. SMF- 41991: Paratypes. 3 fmss. Tan-tshang (Dangchang of new spelling), Kansu, China. Slg. O.v.Moellendorff (measurement of types: ewh 1.500–1.656 – 1.750, wh 6.375–6.500 – 6.625, h 15.4–16.8 – 17.9, diam. maj. 11.4–11.8 –12.0, aph 8.7–9.3 – 9.7, apw 7.2–7.8 – 8.6, h/diam. maj. 1.35–1.40 – 1.49, h/aph 1.68–1.77 – 1.87).
Distribution
Distribution. Gansu (type locality, between Li-dshia-pu and His-gu-tsheng, between Li-dshia-pu and Guan-ting).
Materials Examined
Type locality. Dangchang.
Diagnosis
Diagnosis. Shell bandless; not or faintly striate. Aperture more or less distinctly with angular tubercle. Most swollen part at penultimate and body whorls. With 6.375–7.000 whorls; height 15.4–18.7, diam. maj. 11.4 –14.0. Penis without penial caecum; internally with more than 2 fine longitudinal pilasters. Papilla developed by fused proximal pilasters forming V-shaped structure absent. Epiphallus forming a few loops. Epiphallic caecum absent. Re-description of shell. Ovate; apex protruded. Most swollen part at penultimate and body whorls. Shell dextral; thin; solid; opaque; glossy; not speckled; with spiral grooves (only at umbilical region); with 6.375–6.588 – 7.000 whorls. Whorls not shouldered; flattened. Growth lines usually not very clear. Embryonic shell smooth; polished; with 1.375–1.529 – 1.750 whorls. Postnuclear whorls smooth. Suture normal, without narrow defined zone on beneath whorl. Last whorl gradually ascending towards aperture; rounded at periphery; without smoothed spiral peripheral depression nor rugate region. Shell 16.4–17.5 – 18.7 high; 12.0– 12.7 –14.0 in diam. maj. Height/width ratio 1.30–1.38 – 1.48. Aperture in a plane; completely attached to body whorl; subcircular or truncate-ovate; not adnate; oblique; not armed; more or less distictly with angular tubercle; 9.0–10.0– 10.9 high; 7.2–8.4 – 9.3 wide. Ratio of shell height to aperture height 1.60–1.75 – 1.92. Peristome reflexed, without distinct cuff; expanded; sharp. Parietal callus indistinct. Columellar margin reflexed. Columella vertical or slightly arched. Outer edge of columellar lip vertical. Umbilicus very narrowly open. Apical whorls normally coloured to differently tinted. Shell multicoloured; and umbilical region. in light greyish brown. From about the third or the fourth whorl on, with a broad whitish spiral band which occupies nearly the whole whorl. Aperture white. Genitalia. Vas deferens long; of even thickness; entering epiphallus apically with distinct demarcation. Epiphallus long; cylindrical; of uniform thickness; forming a few loops; externally smooth; without caecum. Flagellum short; tubular; proximally normal; with tip blunt. Penis with terminal entrance of epiphallus; clavate; evenly thick; without penial caecum; thin-walled; internally with more than 2 fine longitudinal pilasters; with 1 V-shaped pilaster. V-shaped pilaster with proximal free end approaching at penial retractor insertion; distally not developing into a papilla. Penial process nipple-shaped; situated at boundary between epiphallus and penis. Penial verge absent. Penial appendix long; branched off from penis at some distance from atrium; divided into sections sections including A- 1 +A- 2, A- 3 and A- 4 +A- 5. A- 1 and A- 2 fused; interiorly with more than 2 fine longitudinal pilasters. A- 2 near A- 3 internally without a ring of papilla. A- 2 and A- 3 not fused. A- 3 opening into A- 2 by no papilla. Boundary between A- 4 and A- 5 indistinct. A- 5 long; convoluted. Penial retractor biramous; attaching to penis distally and to A- 1 + A- 2; with penial retractor arms arising from diaphragm closed to each other. Muscular band connecting vagina and epiphallus absent.. Additional retractor other than penial or appendicular absent. Atrium short; with weak atrial retractor. Free oviduct short; much shorter than vagina. Vagina long; not swollen; straight; not lined with loose, spongy tissue; unpigmented. Bursa copulatrix duct rather long; proximally twisted. Bursa copulatrix with stalk; without apical ligament; normal in size; with long neck; well defined. Diverticle normally present; longer than reservoir; expanded. Bursa copulatrix and diverticle distinguishable; forked more distally from their base. Measurement of genitalia. P: 4.4; Ep: 19.0; Fl: 0.7; VD: 13.0; Va: 3.9; FO: 2.2; BCD: 26.1; BC: 1.6; D: 4.5; A- 1 +A- 2: 6.0; A 3: 1.6. A- 4 +A- 5: 20.2.
Taxon Treatment
- Wu, Min; Zheng, Wei; 2009: A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species, Zootaxa 2053: 19-20. doi
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