Pseudomyrmex ferrugineus (Ward, Philip S. 1993)
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BibTeX: @article{Ward1993JournalofHymenopteraResearch2, RIS/ Endnote: TY - JOUR Wikipedia/ Citizendium: <ref name="Ward1993Journal of Hymenoptera Research2">{{Citation |
Ordo: Hymenoptera
Familia: Formicidae
Genus: Pseudomyrmex
Name
Pseudomyrmex ferrugineus Ward, Philip S., 1993 – Wikispecies link – Pensoft Profile
- Pseudomyrmex ferrugineus Ward, Philip S., 1993, Journal of Hymenoptera Research 2: 135-139.
Description
(Figs. 18, 27,65,66, 70)
Materials Examined
Mexico Examined Guatemala Beccari Mexico Brakhoven Mexico Brakhoven Honduras Puerto Castilla J. Bequaert Mexico Chicoasen G. N. Collins Mexico Cordoba F. Knab Honduras Puerto Castilla J. Bequaert Honduras Bates
Description
Species Author (s) ferrugineus flavicornis nigrocinctus peperi spinicola veneficus Norton (1868a) flavidula (part) _ _ flavidula (part) _ _ Belt (1874) - bicolor - - - - Emery (1890,1891) belti fulvescens belti nigrocincta - spinicola - Wheeler (1913) belti fulvescens (part) spinicola (part) belti (part) nigrocincta - spinicola (part) belti fulvescens (part) belti (part) Wasmann (1915,1916) canescens belti nigrocincta - spinicola - Safford (1922) belti fulvescens (part) belti nigrocincta belti "varieties" (part) spinicola belti fulvescens (part) - Menozzi (1927a,b) - belti obnubila peltata - - - Skwarra (1934a,b) belti uiasmanni (part) - - belti uiasmanni (part) - - Wheeler (1942) belti fulvescens belti saffordi (part) belti vesana belti bequaerti belti uiasmanni belti belti fellosa belti obnubila nigrocincta peltata spinicola convarians belti saffordi (part) spinicola spinicola atrox spinicola gaigei spinicola infernalis spinicola scelerosa belti venefica (part) Janzen (1966,1967a) ferruginea spinicola (part) belti (part) nigrocincta (part) nigrocincta (part) spinicola (part) belti (part) Janzen (1967b, 1969, 1973, 1974, 1975, 1983) ferruginea (part) belti (part) nigrocincta (part) nigrocincta (part) and "undescribed sp." ferruginea (part) venefica Hoelldobler & Engel (1979), Hoelldobler & Engel-Siegel (1985) - - - - ferruginea - Mintzer (1982), Mintzer & Vinson (1985), Mintzer et al. (1987) ferruginea - - - - - Errard (1984) ferruginea - - - - - Espelie & Hermann (1988) ferrugineus - - - - - Ward (1989) ferrugineus flavicornis nigrocinctus peperi spinicola veneficus Young et al. (1990) - flavicornis nigrocinctus - spinicola - *Of the four other species in the P. ferrugineus group, P. satanicus has always been correctly identified (Wheeler, 1942; Janzen, 1969, 1973, 1974); P. mixtecus was mistaken for P. veneficus by Wheeler (1942) and for P. flavicornis (= belti) by Janzen (1966,1973); P. janzeni was referred to by Janzen (1973:729) as an undescribed species derived from P. ferrugineus-, and P. particeps has received no mention in the literature. Worker measurements (n = 69). - HL 0.99-1.33, HW 0.92-1.15, MFC 0.054-0.108, Cl 0.81-0.94, REL 0.42-0.48, REL2 0.48-0.54, 001 1.39-3.16, VI 0.60-0.78, FCI 0.054-0.101, SI 0.41-0.46, SI2 0.79-0.91, NI 0.61-0.72, PLI 0.54-0.69, PWI 0.56- 0.73, PPWI 1.34-1.70.
Diagnosis
Worker diagnosis. - Medium-sized species (HW>0.91; LHT 0.75-1.06) with broad head (Cl> 0.80); anterior margin of median clypeal lobe straight or weakly concave, rounded laterally; palp formula 5,3; frontal carinae well separated (FCI> 0.05) and median lobe of antennal sclerite not strongly exposed (FCI 2 0.45-0.69); mesosomal profile typically as in Fig. 27, with mesonotum notably inclined and with basal face of propodeum rounding gradually into declivitous face, but deviations from this pattern occur; petiole relatively short, high and wide (see relevant metrics: PLI, PWI), with a distinct anterior peduncle (PWI 3 0.36-0.50); posterolateral angles of petiole moderately developed but not as pronounced as in P. peperi (compare Figs. 24 and 27); postpetiole broad. Head densely punctulate, predominantly opaque or subopaque but at least weakly shining on upper third of head between ocelli and compound eye; mesosoma punctulate to punctulate-coriarious, subopaque to sublucid; posterior portions of propodeum opaque to subopaque and usually overlain by larger but weak, irregular punctures or rugulae. Petiole, postpetiole and gaster with fine piligerous punctures, sublucid. Standing pilosity common; pubescence dense but fine and appressed on most surfaces. Color variable, from light reddish- or yellowish-brown to very dark brown, gaster (and usually head) somewhat darker than the mesosoma; mandibles, scapes, fronto-clypeal complex, and apices of legs usually lighter.
Discussion
Comments. - Workers and queens of P. ferrugineus can be recognized by features of head morphology (laterally rounded median clypeal lobe, well separated frontal carinae and correspondingly limited exposure of the median lobes of the antennal sclerites, and moderately broad worker head; see Fig. 18), head sculpture (densely punctulate and (sub)opaque, but weakly shining on upper third of head between the ocelli and compound eye), and coloration (variably brown, not black or orange-brown). This species is most likely to be confused with the allopatric P. janzeni and the partly sympatric P.flavicornis. See under those species for more specific discussion.
Distribution
Distribution and biology. - P. ferrugineus is distributed from eastern and southern Mexico to El Salvador and Honduras (Fig. 70). It is a common species and colonies have been recorded from all swollen-thorn acacia species growing within its range, i.e. Acacia chiapensis, A. collinsii, A. cookii, A. cornigera, A. gentlei, A. globulifera, A. hindsii, A. janzenii, A. mayana and A. sphaerocephala. P. ferrugineus is usually monogynous (Janzen 1967b, 1973) but a few alcohol nest series from Guatemala, belonging to apparently mature colonies (as judged by the presence of alates), contained more than one dealate queen. Janzen (1966, 1967b) conducted a massive experimental study of the interaction between P. ferrugineus and Acacia cornigera in Mexico, and the results of these experiments, together with copious additional observations by Janzen, showed conclusively that the ants protect the acacia from herbivores and competing plants (especially vines). P. ferrugineus was also the subject of a study on kinship and nestmate recognition among workers (Mintzer 1982; Mintzer et al. 1985) which demonstrated a worker-based and probably inherited component to colony odor. Statements about the biology of " P. ferruginea " from regions south of El Salvador and Honduras (e.g. Janzen 1983) refer to a different species, P. spinicola.
Materials Examined
Belize 2.2 mi W Belize, rd. to Chetumal D. H. Janzen Belize 3.8 mi NW Belize, rd. to Chetumal D. H. Janzen Belize 4.9 mi SW Belize, rd. to Cayo D. H. Janzen Belize Belize Baker Belize Belize c. u. Belize Manatee [River] J. D. Johnson Belize Manatee [River] c. u. Belize nr. Belize N. L. H. Krauss Belize 36.1 mi SW Belize D. H. Janzen Belize 36.1 mi [S] W Belize U. Kansas Mex. Exped Belize El Cayo [= San Ignacio] N. L. H. Krauss Belize Pine Mtn. Ridge, Mecal R. G. D. Alpert Belize San Ignacio S. E. Schoenig Belize 13.5 mi S Sta. Elena (Louisville) D. H. Janzen Belize 15 mi S Sta. Elena (Louisville) D. H. Janzen Belize Orange Walk'. 5 mi SE Orange Walk, rd. to Belize D. H. Janzen Belize Punta Gorda H. Broomfield El Salvador 4.7 mi NW La Palma D. H. Janzen El Salvador 5.5 mi SE La Palma D. H. Janzen Guatemala San Joaquin, nr. San Cristobal Verapaz D. H. Janzen Guatemala Trece Aguas Schwarz, Barber Guatemala Yepocapa H. T. Dalmat Guatemala 2 mi W Coatepeque D. H. Janzen Guatemala 4.9 mi SW Sanarate D. H. Janzen Guatemala 1.7 mi S Escuintla [on CA-2] D. H. Janzen Guatemala 12.6 mi NE Escuintla, rd. to Cd. Guatemala D. H. Janzen Guatemala 15 km E Escuintla [on CA- 9] D. H. Janzen Guatemala 3 mi N Escuintla [on CA- 9] D. H. Janzen Guatemala 3 mi S Escuintla [on CA- 9] D. H. Janzen Guatemala 43 km S Cd. Guatemala [= 15 km E Escuintla] D. H. Janzen Guatemala 6.8 mi S Escuintla [on CA-2] D. H. Janzen Guatemala 8.3 mi N Escuintla [on CA- 9] D. H. Janzen Guatemala 9.2 mi N Escuintla [on CA-9] D. H. Janzen Guatemala Escuintla W. M. Wheeler Guatemala Pantaleon Champion Guatemala 16.2 mi NE Cd. Guatemala D. H. Janzen Guatemala 19 kmS Cd. Guatemala [on CA- 9] D. H. Janzen Guatemala 20 mi SE Cd. Guatemala [on CA-1] D. H. Janzen Guatemala 6.2 mi NE Cd. Guatemala D. H. Janzen Guatemala 7.9 mi S Cd. Guatemala [on CA-9] D. H. Janzen Guatemala 8.1 mi NE Cd. Guatemala D. H. Janzen Guatemala Cd. Guatemala Champion Guatemala Cd. Guatemala D. H. Janzen Guatemala Escuintla-Cd. Guatemala [= 19 km S Cd. Guatemala] D. H. Janzen Guatemala 12.4 mi SE Mex. border at Cd. Cuauhtemoc D. H. Janzen Guatemala 12.5 mi SE Mex. border at Cd. Cuauhtemoc D. H. Janzen Guatemala 37.6 mi NW Huehuetenango D. H. Janzen Guatemala 7.1 mi SE Mex. border at Cd. Cuauhtemoc D. H. Janzen Guatemala 9.3 mi SE Mex. border at Cd. Cuauhtemoc D. H. Janzen Guatemala 1.1 mi NE Quirigua D. H. Janzen Guatemala 2. lmi SW Morales D. H. Janzen Guatemala Lago Izabal, 1 km NE El Estor [= 1.5 km NE El Estor] D. H. Janzen Guatemala Los Amates Kellerman Guatemala Murciellago D. H. Janzen Guatemala Quirigua W. M. Wheeler Guatemala 6.7 mi N San Cristobal D. H. Janzen Guatemala 6.9 mi N San Cristobal D. H. Janzen Guatemala 9 mi SW Jutiapa D. H. Janzen Guatemala 70 km NW Tikal W. R. Tschinkel Guatemala Tikal T. H. Hubbell Guatemala Tikal N. L. H. Krauss Guatemala Tikal W. R. Tschinkel Guatemala 7.2 mi NE San Felipe, on Retalhuleu-QuezaltenangoRd. D. H. Janzen Guatemala 7.5 mi NE San Felipe, on Retalhuleu-Quezaltenango Rd. D. H. Janzen Guatemala 1.3 mi E Champerico D. H. Janzen Guatemala 2 mi NE Champerico D. H. Janzen Guatemala 5 mi W Retalhuleu D. H. Janzen Guatemala 5 mi W Retalhuleu, Hwy. CA- 2, at Rio Nil D. H. Janzen Guatemala Champerico Baker Guatemala Champerico F. Knab Guatemala Puenta Samala, 3.8 mi NE San Felipe D. H. Janzen Guatemala Retalhuleu Stoll Guatemala 25 mi SE Escuintla [on CA-2] D. H. Janzen Guatemala Pacific slope c. u. Guatemala Patulul W. M. Wheeler Guatemala 10 mi SW El Lobo [on CA-9] [= 9.2 mi NE Piedras de Afilar] D. H. Janzen Guatemala 2.6 mi SW El Lobo [on CA-9] [= 16.6 mi NE Piedras de Afilar] D. H. Janzen Guatemala 8.1 mi SW Los Amates [on CA-9] [=0.8 mi NE El Lobo] D. H. Janzen Guatemala 8.2 mi NE Piedras de Afilar [on CA-9] [= 12.2 mi NE Rio Hondo] D. H. Janzen Guatemala Beccari Guatemala Concepcion C. N. Ainslie Guatemala Mimosa, " Cocepcion " [Concepcion] C. N. Ainslie Honduras Tela (quarantine New Orleans, U.S.A.) T.J. Baker Honduras Puerto Castilla J. Bequaert Honduras Comayaena S. Passoa Honduras 11.7 mi S San Antonio D. H. Janzen Honduras 15.7 mi N Siguatepeque D. H. Janzen Honduras Minas de Oro J. B. Edwards Honduras 10.4 mi S Sta. Rosa de Copan D. H. Janzen Honduras 11.9 mi S Sta. Rosa de Copan D. H. Janzen Honduras 17.3 mi N Sta. Rosa de Copan D. H. Janzen Honduras 24.6 mi SW San Pedro Sula D. H. Janzen Honduras 6.8 mi S San Pedro Sula D. H. Janzen Honduras San Pedro Sula S. C. Bruner Honduras San Pedro Sula W. M. Mann Honduras 4.4 mi E [Nueva] Ocotepeque D. H. Janzen Honduras 7.3 mi E Quimistan D. H. Janzen Honduras 13.7 mi SW Quimistan D. H. Janzen Honduras Coyoles, W of Olanchito Echtemacht Honduras Honduras Bates Mexico 0.1 mi S Tenabo D. H. Janzen Mexico 0.8 mi E Campeche D. H. Janzen Mexico 10.6 mi E Campeche D. H. Janzen Mexico 20 mi E Campeche, Hwy. 180 D. H. Janzen Mexico 29 mi E & 12 mi S Campeche (Ruinas Edzna) D. H. Janzen Mexico 48 mi NE Puerto Real (Isla Aguada), Hwy. 180 D. H. Janzen Mexico Ruinas Edzna U. Kansas Mex. Exped Mexico 10 mi NE [NW?] Tapachula [on Hwy. 200?] D. H. Janzen Mexico 10.8 mi S Pichucalco D. H. Janzen Mexico 11 km SE Pichucalco P. S. Ward Mexico 11 mi E Arriaga D. H. Janzen Mexico 1 km WSW Palenque P. S. Ward Mexico 26 km E Cintalpa W. MacKay Mexico 2 mi S Pichucalco D. H. Janzen Mexico 32 mi W [San] Cristobal de las Casas, Hwy. 190 D. H. Janzen Mexico 3 km ENE Chiapa de Corzo P. S. Ward Mexico 3 mi N Soyalo [on Hwy. 195] D. H. Janzen Mexico 4 mi NW Ocosingo R. C. Bechtel, E. I. Schlinger Mexico 4 mi NW Ocosingo R. F. Smith Mexico 4 mi S Simojovel R. C. Bechtel, E. I. Schlinger Mexico 4 mi S Simojovel E. I. Schlinger Mexico 56.9 mi NE [NW?] Tapachula [on Hwy. 200?] D. H. Janzen Mexico 5 mi SE Tapilula [on Hwy. 195] D. H. Janzen Mexico 6.9 mi N Tapilula D. H. Janzen Mexico 7 mi SW Teapa [on Hwy. 195] D. H. Janzen Mexico 8.5 mi S La Trinitaria U. Kansas Mex. Exped Mexico 8.5 mi S La Trinitaria, Hwy. 190 D. H. Janzen Mexico 8 mi W Las Cruces, Hwy 190 D. H. Janzen Mexico 96 km S Tuxtla Gutierrez D. E. & J. A. Breedlove Mexico Acapetahua Las Bugamvillas F. Diaz Mexico Arriaga D. H. Janzen Mexico Cd. Cuauhtemoc D. H. Janzen Mexico Chicoasen G. N. Collins Mexico El Real Goodnight, Stannard Mexico Ixtacomitan R. Andrews Mexico Llano Grande G. N. Collins Mexico Palenque c. u. Mexico Pichucalco G. N. Collins Mexico Rio Huixtla, Huixtla J. G. Pereira Mexico Ruinas Palenque E. M. Fisher Mexico San Sebastian [near Tuxtla Gutierrez; see Safford 1922:393] G. N. Collins Mexico Santo Domingo, 15 mi SE [SW?] Simojovel R. F. Smith Mexico Tapilula D. H. Janzen Mexico Tonala D. H. Janzen Mexico Tonola [prob. = Tonala] A. Petrunkewitch Mexico Tuxtla Gutierrez N. L. H. Krauss Mexico Tuxtla Gutierrez W. P. Stoutamize Mexico Yaxoquintela J. E. Rawlins Mexico Yerba Santa G. N. Collins Mexico 10 mi NE Acapulco D. H. Janzen Mexico 15.8 mi S Chilpancingo D. H. Janzen Mexico 18 mi S Chilpancingo F. D. Parker Mexico 18 mi S Chilpancingo F. D. Parker, L. A. Stange Mexico 25.4 mi S Chilpancingo D. H. Janzen Mexico 55 mi N Acapulco, Hwy. 95 Cornell Univ. Mex. Field Party Mexico 55 mi N Acapulco, Hwy. 95 c. u. Mexico 57.8 mi S Chilpancingo D. H. Janzen Mexico 59.9 mi N Acapulco D. H. Janzen Mexico 62.4 mi N Acapulco D. H. Janzen Mexico 74 km N Acapulco W. MacKay Mexico Acapulco L. J. Lipovsky Mexico Revolcadero N. L. H. Krauss Mexico San Geronimo de Juarez W. von Hagen Mexico 2 km W Orizatlan W. MacKay Mexico 15 mi S Cuernavaca E. S. Ross Mexico 15 mi S Cuernavaca W. S. Ross Mexico Cuernavaca W. M. Wheeler Mexico 11.4 - 17.0 mi W Tehuantepec D. H. Janzen Mexico 1 mi W Temascal D. H. Janzen Mexico 22.2 mi N Puerto Escondido D. H. Janzen Mexico 25 mi W Tehuantepec D. H. Janzen Mexico 3.2 mi NE Tehuantepec D. H. Janzen Mexico 42 km N San Pedro Pochutla W. MacKay Mexico 44 mi W Tehuantepec E. E. Gilbert, C. D. MacNeil Mexico 5 mi E Temascal D. H. Janzen Mexico 9.6 km E Santiago Astata W. MacKay Mexico Bahias de Huatulco W. MacKay Mexico Pinotepa Nacional S. W. T. Batra Mexico Salina Cruz D. H. Janzen Mexico Tehuantepec F. Knab Mexico Tehuantepec W. P. Stoutamize Mexico Temascal H. V. Daly Mexico Temascal D. H. Janzen Mexico Temascal D. H. Janzen Mexico Tuxtepec J. Camela G. Mexico Tuxtepec W. M. Mann Mexico Valle Nacional G. V. Halffter Mexico 12.2 mi S Peto, Q. Roo-Yucatan border D. H. Janzen Mexico 16.9 mi W Cd. Chetumal, rd. to Peto D. H. Janzen Mexico 22.5 mi S Felipe Carillo Puerto D. H. Janzen Mexico 6.4 mi E Polyuc D. H. Janzen Mexico 7 mi W Felipe Carillo Puerto D. H. Janzen Mexico 8.1 mi S Felipe Carillo Puerto D. H. Janzen Mexico 8 mi S Felipe Carillo Puerto D. H. Janzen Mexico Sian Ka'an A. Dejean Mexico Sian Ka'an Reserve, nr. Felipe Carillo Puerto A. Dejean Mexico Vallarta A. Dejean Mexico 27 mi N Tamazunchale D. H. Janzen Mexico 40 - 50 mi NW Cd. del Maiz W. S. Ross Mexico 4 mi N Valles W. S. Creighton Mexico Cd. Valles D. H. Janzen Mexico El Bonito, 7 mi S Cd. Valles P. H. & M. Arnaud Mexico El Salto A. Mintzer Mexico El Salto c. u. Mexico Huichihuayan L. J. Lipovsky Mexico Rio Amahac, Tamazunchale W. S. Creighton Mexico Tamazunchale D. H. Janzen Mexico Tamazunchale W. S. Ross Mexico locality not specified, prob. Tanquian [see Safford 1923: 390] Safford Mexico 0.6 mi S Paraiso onrd. to Cardenas D. H. Janzen Mexico 0.9 mi S Chontalpa D. H. Janzen Mexico 12.8 mi S Chontalpa D. H. Janzen Mexico 2.1 mi E Frontera D. H. Janzen Mexico 3 mi W Cardenas D. H. Janzen Mexico 30.2 mi W Cd. El Carmen D. H. Janzen Mexico 37.8 mi E Coatzacoalcos, Hwy. 180 D. H. Janzen Mexico 6.6 mi S Chontalpa D. H. Janzen Mexico Chontalpa, 26 mi S Cardenas D. H. Janzen Mexico Cardenas D. H. Janzen Mexico Teapa J. C. & D. Pallister Mexico Teapa H. H. Smith Mexico Teapa W. P. Stoutamize Mexico 22.7 mi S Cd. Victoria D. H. Janzen Mexico 41 mi S Cd. Victoria C. W. Obrien Mexico 50 mi N Valles G. E. Bohart Mexico 5 mi N Cd. Mante A. Mintzer Mexico 7 km WSW El Encino P. S. Ward Mexico Antiguo Morelos c. u. Mexico Cd. Madero F. Infante M. Mexico Ciudad Mante N. E. & M. A. Evans Mexico Ciudad Mante N. L. H. Krauss Mexico El Limon H. E. Evans Mexico Forlon L. J. Lipovsky Mexico Gonzales c. u. Mexico Llera W. E. LaBerge Mexico Mesa de Llera A. Mintzer Mexico N of Antiguo Morelos A. Mintzer Mexico Rio Guayalejo at Hwy. 85 A. S. Menke, L. A. Stange Mexico S of Cd. Mante A. Mintzer Mexico Tampico Brakhoven Mexico Tampico D. L. Crawford Mexico Tampico H. Jourdan Mexico Tampico N. L. H. Krauss Mexico Tampico Locke Mexico Tampico E. Palmer Mexico Tampico E. A. Schwarz Mexico Tampico W. P. Stephen Mexico Tampico c. u. Mexico Tampico, dunes at Cd. Madero D. H. Janzen Mexico Xicotencatl H. C. Millender Mexico N. M., Vera Cruz Townsend Mexico Vera Cruz G. Seurat Mexico Vera Cruz H. H. Smith Mexico 10 mi W Veracruz G. E. Bohart Mexico 14 km ENE La Tinaja P. S. Ward Mexico 15 mi W Veracruz U. Kansas Mex. Exped Mexico 16 km E Cuilahuac [= Cuitlahuac] W. MacKay Mexico 24.9 mi NW Acayucan D. H. Janzen Mexico 28 km N Cardel, Morro de la Mancha V. Rico-Gray Mexico 30 mi S Acayucan F. D. Parker Mexico 3 mi N Sayula R. C. Bechtel Mexico 3 mi N Sayula R. C. Bechtel, E. I. Schlinger Mexico 4 mi NE Minatitlan R. C. Bechtel, E. I. Schlinger Mexico 4 miNWRinconada Antigua U. KansasMex. Exped. Mexico 4 mi W Puente Nacional U. Kansas Mex. Exped. Mexico 6.5 km N Tierra Blanca W. MacKay Mexico 7 mi S Cardel c. u. Mexico 9 km NNW Sontecomapan P. S. Ward Mexico Boca del Rio U. Kansas Mex. Exped. Mexico Buen Pais R. C. Bechtel, E. I. Schlinger Mexico Camaron E. Skwarra Mexico Cordoba F. Knab Mexico Cordoba R. R. Snelling Mexico Cordoba c. u. Mexico Cotaxtla Exp. Sta., Cotaxtla D. H. Janzen Mexico Est. Biol. Los Tuxtlas H. A. Hespenheide Mexico Est. Biol. " Los Tuxtlas ", nr. San Andres Tuxtla G. Ibarra M. Mexico Fortin c. u. Mexico Jalapa Rangel Mexico Jalapa W. M. Wheeler Mexico Jalapa c. u. Mexico Jalapilla, mun. Jilotepec G. Aleman Mexico Los Cocos A. Petrunkewitch Mexico Los Tuxtlas, 10 km NNW Sontecomapan P. S. Ward Mexico Mirador E. Skwarra Mexico Mocambo D. H. Janzen Mexico Orizaba c. u. Mexico Palma Sola Halffter, Reyes Mexico Panuco F. Parker, D. Miller Mexico Playa Azul, Catemaco W. P. Stoutamize Mexico Pueblo Nuevo, nr. Tezonapa Cornell Univ. Mex. Field Party Mexico Remutadero E. Skwarra Mexico Santa Lucrecia [= Jesus Carranza] F. Knab Mexico Santa Lucrecia [= Jesus Carranza] P. Knab Mexico Tamarindo E. Skwarra Mexico Tamos F. C. Bishop Mexico Tinajas [presumably La Tinaja, on Hwy. 150] F. D. Parker, L. A. Stange Mexico Tlacocintla E. Skwarra Mexico Veracruz G. E. Bohart Mexico Veracruz N. L. H. Krauss Mexico Veracruz E. Skwarra Mexico Veracruz L. A. Stange Mexico 14.8 mi S Ticul, " Hwy. 180 " [prob. Hwy. 261] D. H. Janzen Mexico 8 mi E Merida (rd. to Pto. Juarez) D. H. Janzen Mexico Merida D. H. Janzen Mexico Merida N. L. H. Krauss Mexico Merida (S margin of town) D. H. Janzen Mexico Tekax, 33 mi W Peto D. H. Janzen Mexico Temax Gaumer Mexico Mex Norton
Taxon Treatment
- Ward, Philip S.; 1993: Systematic studies on Pseudomyrmex acacia-ants (Hymenoptera: Formicidae: Pseudomyrmecinae), Journal of Hymenoptera Research 2: 135-139. doi
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