Moullava
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Ordo: Fabales
Familia: Leguminosae
Name
Moullava Adans., Fam. Pl. 2: 318. 1763, descr. emended E. Gagnon & G. P. Lewis – Wikispecies link – Pensoft Profile
- Wagatea Dalzell (1851).
- Cinclidocarpus Zoll. & Moritzi (1846).
- Caesalpinia sect. Cinclidocarpus (Zoll. & Moritzi) Benth. & Hook. (1865).
Diagnosis
Moullava is related to Mezoneuron, but differs by its fleshy, oblong-elliptic, indehiscent, sub-torulose, wingless pods, with thickened sutures (vs. laterally compressed, chartaceous, coriaceous or ligneous, indehiscent pods, with a longitudinal wing along the upper suture), and by its subglobular (vs. compressed) seeds.
Type
“H.M. 6 t. 6” (= Rheede`s Hortus Malabaricus 6, plate 6, 1686) = Moullava spicata.
Emended description
Lianas and scrambling shrubs, armed with deflexed prickles on shoots. Stipules not seen. Leaves alternate, bipinnate, ending with a pair of pinnae, 12–40 cm long, glabrous to pubescent-tomentose, with a pair of prickles at the insertion of each pinna; pinnae opposite, in 7–20 pairs; leaflets in 5–40 opposite pairs per pinna, sessile, narrowly oblong to ovate-oblong, apex rounded to emarginate, sometimes mucronate, base asymmetrical to rounded, blades eglandular, glabrous to pubescent, 4–20 × 2–6 mm. Inflorescence an elongated terminal or axillary raceme, the flowers subsessile, pedicels, when present, 10–25 mm long, the racemes sometimes aggregated into panicles, 8–60 cm long, unarmed or with a few prickles at the base. Flowers bisexual, sub-actinormophic or zygomorphic; calyx comprising a hypanthium with 5 sepals, 6–12 × 2–4 mm, the lower sepal strongly cucullate, covering the other 4 sepals in bud, all sepals eglandular and glabrous; petals 5, free, yellow, the median and lateral petals sometimes streaked red, eglandular; stamens 10, free, barely exserted beyond the corolla, densely pubescent on lower half of filaments, 8–15 mm long; ovary glabrous or pubescent. Fruit fleshy, oblong-elliptic, unarmed, indehiscent, sub-torulose, with thickened sutures, the apex apiculate, 35–50 (–80) × 15–30 mm, drying black (immature fruits of Moullava spicata red-tomentose), exocarp and endocarp strongly adnate, glabrous, 1–4-seeded. Seeds sub-globular, 12–20 mm in diameter, olive-brown to black.
Geographic distribution
A genus of four species, three in south Asia: India, Nepal, Myanmar (Burma), Thailand, Laos, Cambodia, Sri Lanka, southern China (Yunnan and Hainan), and the Malay Peninsula and Archipelago, and one in Africa: Cameroun, Gabon, the Democratic Republic of Congo, Angola, Zambia (Kabompo Dist.), Uganda and Tanzania (Kigoma Dist.).
Habitat
The Asian species are found in seasonally dry tropical semi-evergreen forest margins, secondary thickets, and on mountain slopes, up to 1200 m elevation. The African species occurs mostly in riverine habitats in lowland rainforests.
Etymology
Derived from the vernacular name of Moullava spicata, “mulu” (Malayalam: spiny), a spiny climber.
References
Brenan (1963[1], 1967[2]); Hattink (1974)[3]; Vidal and Hul Thol (1976)[4]; Nicolson (1980)[5]; Ansari (1990)[6]; Sanjappa (1992[7]: 33); Brummitt et al. (2007[8], see both Moullava and Mezoneuron welwitschianum); Chen et al. (2010a)[9].
Taxon Treatment
- Gagnon, E; Bruneau, A; Hughes, C; de Queiroz, L; Lewis, G; 2016: A new generic system for the pantropical Caesalpinia group (Leguminosae) PhytoKeys, (71): 1-160. doi
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Other References
- ↑ Brenan J (1963) Notes on African Caesalpinioideae. Kew Bulletin 17: 197–218. doi: 10.2307/4118939
- ↑ Brenan J (1967) Leguminosae, part 2: subfamily Caesalpinioideae. In: Milne-Redhead E Polhill R (Eds) Flora of Tropical East Africa. Crown Agents for Oversea Goverments and Administration, London.
- ↑ Hattink T (1974) A revision of Malesian Caesalpinia, including Mezoneuron (Leguminosae–Caesalpiniaceae). Reinwardtia 9(1): 1–69.
- ↑ Vidal J, Hul Thol S (1976) Révision des Caesalpinia asiatiques. Bulletin du Muséum National d’histoire naturelle, 3e série 395: 27–81.
- ↑ Nicolson D (1980) Moullava Adanson, recently Wagatea Dalzell (Fabaceae-Caesalpinioideae). In: Manilal K (Ed.) Botany and History of Hortus Malabaricus. Oxford & IBH Publi. Co., New Delhi, 181–185.
- ↑ Ansari A (1990) Extended distribution of an endemic monotypic genus Wagatea Dalz. Journal of Economic and Taxonomic Botany 14(3): 746–747.
- ↑ Sanjappa M (1992) Legumes of India. Bishen Singh Mahendra Pal Singh, Dehra Dun, India, 1–338.
- ↑ Brummitt R, Chikuni A, Lock J, Polhill R (2007) Leguminosae. In: Timberlake J Pope G Polhill R Martins E (Eds) Flora Zambesiaca, vol. 3(2). Kew Royal Botanic Gardens, 1–218.
- ↑ Chen D, Zhang D, Hou D (2010a) Caesalpinia. In: Wu C Raven P (Eds) Flora of China, vol. 10. Missouri Botanical Garden Press, St-Louis, USA, 41–47.
