Liogluta

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Klimaszewski J, Webster R, Langor D, Sikes D, Bourdon C, Godin B, Ernst C (2016) A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys (273) : 217–256, doi. Versioned wiki page: 2016-03-24, version 78694, https://species-id.net/w/index.php?title=Liogluta&oldid=78694 , contributors (alphabetical order): Pensoft Publishers.

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BibTeX:

@article{Klimaszewski2016ZooKeys,
author = {Klimaszewski, Jan AND Webster, Reginald P. AND Langor, David W. AND Sikes, Derek AND Bourdon, Caroline AND Godin, Benoit AND Ernst, Crystal},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae)},
year = {2016},
volume = {},
issue = {273},
pages = {217--256},
doi = {10.3897/zookeys.573.7878},
url = {http://zookeys.pensoft.net/articles.php?id=7878},
note = {Versioned wiki page: 2016-03-24, version 78694, https://species-id.net/w/index.php?title=Liogluta&oldid=78694 , contributors (alphabetical order): Pensoft Publishers.}

}

RIS/ Endnote:

TY - JOUR
T1 - A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae)
A1 - Klimaszewski J
A1 - Webster R
A1 - Langor D
A1 - Sikes D
A1 - Bourdon C
A1 - Godin B
A1 - Ernst C
Y1 - 2016
JF - ZooKeys
JA -
VL -
IS - 273
UR - http://dx.doi.org/10.3897/zookeys.573.7878
SP - 217
EP - 256
PB - Pensoft Publishers
M1 - Versioned wiki page: 2016-03-24, version 78694, https://species-id.net/w/index.php?title=Liogluta&oldid=78694 , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/zookeys.573.7878

Wikipedia/ Citizendium:

<ref name="Klimaszewski2016ZooKeys">{{Citation
| author = Klimaszewski J, Webster R, Langor D, Sikes D, Bourdon C, Godin B, Ernst C
| title = A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae)
| journal = ZooKeys
| year = 2016
| volume =
| issue = 273
| pages = 217--256
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.573.7878
| url = http://zookeys.pensoft.net/articles.php?id=7878
| pmc =
| accessdate = 2024-12-13

}} Versioned wiki page: 2016-03-24, version 78694, https://species-id.net/w/index.php?title=Liogluta&oldid=78694 , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Taxonavigation

Ordo: Coleoptera
Familia: Staphylinidae

Name

Liogluta Thomson, 1858 – Wikispecies link – Pensoft Profile

Diagnosis

Body length ranging from 2.8–5.4 mm, body narrowly subparallel (Figs 1, 8, 16, 23, 30, 35, 44, 50, 58, 65, 72, 79, 86, 90), moderately flattened; elytra and abdomen wider than head and pronotum; uniformly dark brown or reddish-brown with head and posterior abdomen dark brown to almost black; integument of forebody with strong meshed microsculpture; surface of elytra often granulose (Figs 50, 51); head with incomplete postocular carinae, postocular region of head long and in most species longer than diameter of eye; glossae split into two lobes; antennae long with articles VI-X subquadrate, slightly transverse or rarely slightly elongate; pronotum with hypomera fully visible in lateral view; pubescence of pronotum directed posteriad on midline of disc and posterolaterad to laterad on sides; mesocoxae narrowly separated, metasternal process short and acute; legs long, three basal articles of metatarsi highly elongate in most species; tarsal formula 5-5-5. Male. Apical margin of male tergite VIII often with broad, variably-shaped rectangular projection, with edge entire or bearing crenulation or variably shaped structures, with two lateral teeth and sometimes with additional median tooth; integument of disc often with broadly distributed asperate punctation near apex (Figs 3, 11, 18, 25, 38, 46, 53, 60, 67, 74, 81, 88, 92); median lobe of aedeagus simple with apical part variably shaped in lateral view (Figs 2, 9, 17, 24, 36, 45, 52, 59, 66, 73, 80, 87, 91). Female. Spermatheca with capsule narrowly club-shaped or tubular, with apical invagination moderate to deep, stem long, sinuate, looped or twisted posteriorly (Figs 7, 15, 22, 29, 34, 42, 43, 57, 64, 71, 78, 85, 96); sternite VIII with apical margin rounded or medially emarginate, antecostal suture arcuate, or slightly to distinctly sinuate (Figs 6, 14, 21, 28, 33, 41, 49, 56, 63, 70, 77, 84, 89, 95).

Distribution

The genus is holarctic in distribution (Smetana 2004[8]). Seevers (1978)[9] mentioned a few species from Africa and Jamaica but these records need verification.

Key to Canadian and Alaskan species of Liogluta

Key to Canadian and Alaskan species of Liogluta — J. Klimaszewski , R. Webster , D. Langor , D. Sikes , C. Bourdon , B. Godin , C. Ernst
1
Eyes small, each shorter than postocular region of head in dorsal view (Figs 1, 8, 16, 23, 30) 
  2
–
Eyes large, each subequal in length to postocular region of head in dorsal view (Figs 35, 44, 50, 58, 65, 72, 79, 86, 90) 
  6
2
Antennae enlarged (Fig. 16) 
   Liogluta quadricollis (Casey)
–
Antennae normally developed and not enlarged (Figs 1, 8, 23, 30) 
  3
3
Body more-or-less uniformly reddish- to yellowish-brown (Fig. 1) 
   Liogluta terminalis (Casey)
–
Body brown to dark brown with paler pronotum and elytra, or reddish-yellow with brown head, antennae and abdomen (Figs 8, 23, 30) 
  4
4
Body narrow (Fig. 8); abdomen subparallel, distinctly narrower at base than elytra (Fig. 8); male tergite VIII truncate apically, with two large lateral teeth and apical margin denticulate (Fig. 11); bulbus moderately large, tubus long with apical part moderately broad (Figs 9, 10); spermatheca with broad capsule, with apical invagination small and shallow, stem sinuate with moderately long posterior loop (Fig. 15) 
   Liogluta trapezicollis Lohse
–
Body broad (Figs 23, 30); abdomen arcuate laterally and broadest at middle of its length; genital structures shaped differently (Figs 24, 29, 34) 
  5
5
Pronotum about evenly arcuate laterally (Fig. 23); male tergite VIII truncate apically, with two small lateral teeth and apical margin slightly crenulate (Fig. 25); spermatheca with capsule narrow and apical invagination deep, stem sinuate with long posterior loop (Fig. 29) 
   Liogluta wickhami (Casey)
–
Pronotum trapezoidal in shape, narrowest at base and broadest in apical third (Fig. 30); male unknown; spermatheca with broad capsule, with apical invagination small and shallow, stem sinuate with moderately long posterior loop (Fig. 34) 
   Liogluta vasta (Mäklin)
6
Elytra broad, about 25% wider than maximum width of pronotum (Figs 35, 44, 90), integument moderately to highly glossy; male tergite VIII truncate apically and without distinct lateral teeth, not or moderately produced apically (Figs 38, 46, 92) 
  7
–
Elytra narrow, at most 20% wider than maximum width of pronotum (Figs 50, 58, 65, 72, 79, 86); integument moderately glossy in most species; male tergite VIII usually with well-defined lateral teeth, and with or without crenulation on apical margin (Figs 53, 60, 67, 74, 81, 88) 
  9
7
Body moderately glossy (Fig. 90); antennal articles VI-X subquadrate to slightly elongate (Fig. 90); terminalia and genitalia as illustrated (Figs 91–96) 
   Liogluta gigantea Klimaszewski & Langor
–
Body highly glossy (Figs 35, 44); antennal articles VI-X slightly to distinctly transverse (Figs 35, 44); terminalia and genitalia shaped differently (Figs 36–43, 45–49) 
  8
8
Antennal articles VI-X subquadrate to slightly transverse (Fig. 35); body usually uniformly dark brown to black (Fig. 35); male tergite VIII truncate and not produced apically (Fig. 38); remaining terminalia, median lobe of aedeagus, and spermatheca as illustrated (Figs 36, 37, 39–43) 
   Liogluta nigropolita (Bernhauer)
–
Antennal articles VI-X distinctly transverse (Fig. 44); body dark brown to black with brownish elytra (Fig. 44); male tergite VIII truncate and produced apically (Fig. 46); median love of aedeagus and remaining terminalia as illustrated (Figs 45–49) 
   Liogluta nitens (Mӓklin)
9
Punctures on pronotum and elytra sparse, distance between punctures greater than diameter of a puncture (Fig. 50); surface of disc strongly granulose (Figs 50, 51); male tergite VIII broadly triangularly produced apically and obtusely angular medially (Fig. 53); remaining terminalia and genitalia as illustrated (Figs 52, 54–57) 
   Liogluta granulosa Lohse
–
Punctures on pronotum and elytra dense, distance between punctures about equal to diameter of a puncture (Figs 58, 65, 72, 79, 86); surface of disc slightly granulose; male tergite VIII truncate apically (Figs 60, 67, 74, 81, 88) 
  10
10
Body length 2.8 mm; pronotum with microsculpture weakly defined (Fig. 86); elytra slightly longer than pronotum (Fig. 86); male tergite VIII truncate apically and evenly serrated, not angulate medially (Fig. 88); median lobe of aedeagus as illustrated (Figs 87); known only from British Columbia 
   Liogluta atriventris (Casey)
–
Body length 3.9–5.4 mm; pronotum with distinct microsculpture; elytra as long as pronotum or rarely slightly longer (Figs 58, 65, 72, 79); male tergite VIII truncate apically, crenulate or entire, and sometimes angulate medially (Figs 60, 67, 74, 81) 
  11
11
Antennal articles IV-XI yellowish to light brown (Fig. 79); pronotum one-sixth broader than long (Fig. 79); forebody reddish to reddish-yellow (Fig. 79); basal metatarsal article distinctly longer than following article (Fig. 79); male tergite VIII distinctly crenulate apically (Fig. 81); median lobe of aedeagus with apical part of tubus narrow and straight in lateral view (Fig. 80); female sternite VIII emarginate apically (Fig. 84); spermatheca as illustrated (Fig. 85) 
   Liogluta intermedia Klimaszewski & Langor
–
Antennal articles IV-XI dark brown to black (Figs 58, 65, 72); pronotum usually one-fifth broader than long (Figs 58, 65, 72); forebody dark reddish-brown to almost black (Figs 58, 65, 72); basal metatarsus as long as or slightly longer than following article (Figs 58, 65, 72); male tergite VIII without or with slight crenulation (Figs 60, 67, 74); median lobe of aedeagus with apical part of tubus wide or narrow and arched ventrad in lateral view (Figs 59, 66, 73); female sternite VIII truncate or emarginate medially at apex (Figs 63, 70, 77); spermatheca as illustrated (Figs 64, 71, 78) 
  12
12
Pronotum and elytra dark reddish-brown, without apparent dark brown irregularly shaped spots (Fig. 65); male tergite VIII truncate apically with two lateral teeth (Fig. 67); median lobe of aedeagus with apical part of tubus broad in lateral view (Fig. 66); female sternite VIII with antecostal suture distinctly sinuate and subangulate medially (Fig. 70); spermatheca with very long stem (Fig. 71) 
   Liogluta castoris Klimaszewski & Webster, sp. n.
–
Pronotum and elytra dark reddish-brown, mottled with dark brown or black irregularly shaped spots (Figs 58, 72); male tergite VIII truncate apically without or with three small apical teeth (Figs 60, 74); median lobe of aedeagus with apical part of tubus narrow in lateral view (Figs 59, 73); female sternite VIII with antecostal suture slightly sinuate (Figs 63, 77); spermatheca with moderately long stem (Figs 64, 78) 
  13
13
Pronotum distinctly transverse (Fig. 72); two basal antennal articles brown (Fig. 72); apical margin of male tergite VIII evenly rounded (Fig. 74); median lobe of aedeagus with apical part of tubus straight in lateral view (Fig. 73); female sternite VIII broadly rounded apically (Fig. 77); spermatheca as illustrated (Fig. 78) 
   Liogluta pseudocastoris Klimaszewski & Webster, sp. n.
–
Pronotum slightly transverse, appearing subquadrate (Fig. 58); two basal antennal articles reddish-brown (Fig. 58); apical margin of male tergite VIII with subrectangular broad projection with two lateral and one median teeth (Fig. 60); median lobe of aedeagus with apical part of tubus arched ventrad in lateral view (Fig. 59); female sternite VIII emarginate apically (Fig. 63); spermatheca as illustrated (Fig. 64) 
   Liogluta microgranulosa Klimaszewski & Webster, sp. n.

Taxon Treatment

  • Klimaszewski, J; Webster, R; Langor, D; Sikes, D; Bourdon, C; Godin, B; Ernst, C; 2016: A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae) ZooKeys, (273): 217-256. doi

Images

Figures 1–7. Liogluta terminalis (Casey): 1 habitus in dorsal view 2 median lobe of aedeagus in lateral view 3 male tergite VIII 4 male sternite VIII 5 female tergite VIII 6 female sternite VIII 7 spermatheca. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 1–7. Liogluta terminalis (Casey): 1 habitus in dorsal view 2 median lobe of aedeagus in lateral view 3 male tergite VIII 4 male sternite VIII 5 female tergite VIII 6 female sternite VIII 7 spermatheca. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 8–15. Liogluta trapezicollis Lohse: 8 habitus in dorsal view 9, 10 median lobe of aedeagus in lateral view 11 male tergite VIII 12 male sternite VIII 13 female tergite VIII 14 female sternite VIII 15 spermatheca. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 8–15. Liogluta trapezicollis Lohse: 8 habitus in dorsal view 9, 10 median lobe of aedeagus in lateral view 11 male tergite VIII 12 male sternite VIII 13 female tergite VIII 14 female sternite VIII 15 spermatheca. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 16–22. Liogluta quadricollis (Casey): 16 habitus in dorsal view 17 median lobe of aedeagus in lateral view 18 male tergite VIII 19 male sternite VIII [16–19 based on male from BC] 20 female tergite VIII 21 female sternite VIII 22 spermatheca (19–22 based on female holotype). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 16–22. Liogluta quadricollis (Casey): 16 habitus in dorsal view 17 median lobe of aedeagus in lateral view 18 male tergite VIII 19 male sternite VIII [16–19 based on male from BC] 20 female tergite VIII 21 female sternite VIII 22 spermatheca (19–22 based on female holotype). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 23–29. Liogluta wickhami (Casey): 23 habitus in dorsal view 24 median lobe of aedeagus in lateral view 25 male tergite VIII 26 male sternite VIII 27 female tergite VIII 28 female sternite VIII 29 spermatheca [23, 27–29 based on female holotype]. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 23–29. Liogluta wickhami (Casey): 23 habitus in dorsal view 24 median lobe of aedeagus in lateral view 25 male tergite VIII 26 male sternite VIII 27 female tergite VIII 28 female sternite VIII 29 spermatheca [23, 27–29 based on female holotype]. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 30–34. Liogluta vasta (Mäklin): 30 habitus in dorsal view 31 female pygidium (terminal segments) 32 female tergite VIII 33 female sternite VIII (30–33 based on female holotype) 34 spermatheca (based on YT specimen after Lohse and Smetana 1895). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 30–34. Liogluta vasta (Mäklin): 30 habitus in dorsal view 31 female pygidium (terminal segments) 32 female tergite VIII 33 female sternite VIII (30–33 based on female holotype) 34 spermatheca (based on YT specimen after Lohse and Smetana 1895). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 35–43. Liogluta nigropolita (Bernhauer): 35 habitus in dorsal view 36 median lobe of aedeagus in lateral view 37 median lobe of aedeagus in dorsal view (36, 37 after Lohse et al. 1990[2]) 38 male tergite VIII 39 male sternite VIII 40 female tergite VIII 41 female sternite VIII 42, 43 spermatheca (43 after Lohse et al. 1990[2]). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 35–43. Liogluta nigropolita (Bernhauer): 35 habitus in dorsal view 36 median lobe of aedeagus in lateral view 37 median lobe of aedeagus in dorsal view (36, 37 after Lohse et al. 1990[2]) 38 male tergite VIII 39 male sternite VIII 40 female tergite VIII 41 female sternite VIII 42, 43 spermatheca (43 after Lohse et al. 1990[2]). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 44–49. Liogluta nitens (Mäklin): 44 habitus in dorsal view (morphotype with broad and long elytra) 45 median lobe of aedeagus in lateral view 46 male tergite VIII 47 male sternite VIII 48 female tergite VIII 49 female sternite VIII. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 44–49. Liogluta nitens (Mäklin): 44 habitus in dorsal view (morphotype with broad and long elytra) 45 median lobe of aedeagus in lateral view 46 male tergite VIII 47 male sternite VIII 48 female tergite VIII 49 female sternite VIII. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 50–57. Liogluta granulosa Lohse: 50 habitus in dorsal view 51 elytron 52 median lobe of aedeagus in lateral view 53 male tergite VIII 54 male sternite VIII [50–54 based on male holotype] 55 female tergite VIII 56 female sternite VIII 57 spermatheca [55–57 based on female from YT]. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 50–57. Liogluta granulosa Lohse: 50 habitus in dorsal view 51 elytron 52 median lobe of aedeagus in lateral view 53 male tergite VIII 54 male sternite VIII [50–54 based on male holotype] 55 female tergite VIII 56 female sternite VIII 57 spermatheca [55–57 based on female from YT]. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 58–64. Liogluta microgranulosa Klimaszewski & Webster, sp. n.: 58 habitus in dorsal view 59 median lobe of aedeagus in lateral view 60 male tergite VIII 61 male sternite VIII 62 female tergite VIII 63 female sternite VIII 64 spermatheca (58–64 based on type specimens). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 58–64. Liogluta microgranulosa Klimaszewski & Webster, sp. n.: 58 habitus in dorsal view 59 median lobe of aedeagus in lateral view 60 male tergite VIII 61 male sternite VIII 62 female tergite VIII 63 female sternite VIII 64 spermatheca (58–64 based on type specimens). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 65–71. Liogluta castoris Klimaszewski & Webster, sp. n.: 65 habitus in dorsal view 66 median lobe of aedeagus in lateral view 67 male tergite VIII 68 male sternite VIII 69 female tergite VIII 70 female sternite VIII 71 spermatheca (65–71 based on type specimens). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 65–71. Liogluta castoris Klimaszewski & Webster, sp. n.: 65 habitus in dorsal view 66 median lobe of aedeagus in lateral view 67 male tergite VIII 68 male sternite VIII 69 female tergite VIII 70 female sternite VIII 71 spermatheca (65–71 based on type specimens). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 72–78. Liogluta pseudocastoris Klimaszewski & Webster, sp. n.: 72 habitus in dorsal view 73 median lobe of aedeagus in lateral view 74 male tergite VIII 75 male sternite VIII 76 female tergite VIII 77 female sternite VIII 78 spermatheca (72–78 based on type specimens). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 72–78. Liogluta pseudocastoris Klimaszewski & Webster, sp. n.: 72 habitus in dorsal view 73 median lobe of aedeagus in lateral view 74 male tergite VIII 75 male sternite VIII 76 female tergite VIII 77 female sternite VIII 78 spermatheca (72–78 based on type specimens). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 79–85. Liogluta intermedia Klimaszewski & Langor: 79 habitus in dorsal view 80 median lobe of aedeagus in lateral view 81 male tergite VIII, 82 male sternite VIII 83 female tergite VIII 84 female sternite VIII 85 spermatheca. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 79–85. Liogluta intermedia Klimaszewski & Langor: 79 habitus in dorsal view 80 median lobe of aedeagus in lateral view 81 male tergite VIII, 82 male sternite VIII 83 female tergite VIII 84 female sternite VIII 85 spermatheca. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 86–89. Liogluta atriventris (Casey): 86 habitus in dorsal view 87 median lobe of aedeagus in lateral view 88 male tergite VIII 89 male sternite VIII (86–89 based on male lectotype). Female unknown. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 86–89. Liogluta atriventris (Casey): 86 habitus in dorsal view 87 median lobe of aedeagus in lateral view 88 male tergite VIII 89 male sternite VIII (86–89 based on male lectotype). Female unknown. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 
Figures 90–96. Liogluta gigantea Klimaszewski & Langor: 90 habitus in dorsal view 91 median lobe of aedeagus in lateral view 92 male tergite VIII 93 male sternite VIII 94 female tergite VIII 95 female sternite VIII 96 spermatheca (90, 94–96 based on holotype, 91–93 based on male from Ontario). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
Figures 90–96. Liogluta gigantea Klimaszewski & Langor: 90 habitus in dorsal view 91 median lobe of aedeagus in lateral view 92 male tergite VIII 93 male sternite VIII 94 female tergite VIII 95 female sternite VIII 96 spermatheca (90, 94–96 based on holotype, 91–93 based on male from Ontario). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm. 

Other References

  1. ↑ Lohse G (1974) Hypocyphtinae und Aleocharinae. [in part; pp. 7–72, 221–292] In: Freude H Harde K Lohse G (Eds) Die Käfer Mitteleuropas. Band 5. Staphylinidae II. Goecke & Evers, Krefeld, 381 pp.
  2. ↑ 2.0 2.1 2.2 Lohse G, Klimaszewski J, Smetana A (1990) Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin 44: 121–202.
  3. ↑ Casey T (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St. Louis 16: 125–434.
  4. ↑ Casey T (1910) New species of the staphylinid tribe Myrmedoniini. In: Memoirs on the Coleoptera. I. New Era Printing Co., Lancaster, Pennsylvania, 1–183. doi: 10.5962/bhl.title.1159
  5. ↑ 5.0 5.1 5.2 Fenyes A (1920) 173B fascicule. Coleoptera. Fam. Staphylinidae. Subfam. Aleocharinae [pp. 111–414] In: Wytsman P (Ed.) Genera Insectorum. Vol. XXVII. L. Desmet-Verteneuil, Bruxelles, 453 pp. + 7 pls.
  6. ↑ 6.0 6.1 6.2 Bernhauer M, Scheerpeltz O (1926) Pars 82: Staphylinidae VI [pp. 499–988] In: Coleopterorum Catalogus. Volumen V. Staphylinidae. W. Junk, Berlin, 988 pp.
  7. ↑ 7.0 7.1 7.2 Moore I, Legner E (1975) A catalogue of the Staphylinidae of America north of Mexico (Coleoptera). University of California, Division of Agricultural Sciences, Special Publication No. 3015: 1–514.
  8. ↑ Smetana A (2004) Subfamily Aleocharinae Fleming, 1921. In: Löbl I Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Vol. 2. Hydrophiloidea - Histeroidea – Staphylinoidea. Apollo Books, Stenstrup, 353–494.
  9. ↑ Seevers C (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology 71: i-vi, 1–289.
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