Lethades

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Taxonavigation

Ordo: Hymenoptera
Familia: Ichneumonidae

Name

Lethades Davis, 1897 – Wikispecies link – Pensoft Profile

• Lethades Davis, 1897: 204. Type species: Adelognathus texanus Ashmead, 1890. Monobasic.

Diagnosis

Lethades can be distinguished from all other genera in the Pionini by the combination of the following characters: first flagellomere longer than second; second trochanter of hind leg rounded without a transverse ridge; glymma present; profile of the propodeum nearly rounded with short posterior field; dorsomedian and dorsolateral carinae of the T1 converging at base; ovipositor without subapical notch; cerci parallel-sided and protruding (Townes 1970, Cameron and Wharton 2011). The notaulus varies from absent to deep, but very short, in nearly all described species of Lethades. Only Lethades schaffneri Hinz has an elongate notaulus. The latter species was transferred to Lethades based on other characters, especially ovipositor morphology (Cameron and Wharton 2011).

Key to world species of the genus Lethades

Key to world species of the genus Lethades — A. Reshchikov , Z. Xu , H. Pang

1		Notauli distinctly impressed extending at least over the anterior 0.5 of the mesoscutum. Claws not pectinate     Lethades schaffneri Hinz
–		Notauli absent or weakly impressed, not extending the anterior 0.5 of the mesoscutum. Claws pectinate    2
2		Fore wing areolet absent    3
–		Fore wing areolet present    5
3		Body finely striated and weakly punctate. Fore femur and tibia uniformly red     Lethades amauroneinati (Hinz)
–		Body distinctly and densely punctate, mesopleuron weakly striated. Fore femur and tibia not uniformly red    4
4		Epicnemial carina reaching anterior edge of mesopleuron. Metasomal tergites without yellow bands on posterior margins. Fore femur and tibia red, dark apically     Lethades schmiedeknechti Hinz
–		Epicnemial carina not reaching anterior edge of mesopleuron. Metasomal tergites with yellow bands on posterior margins. Fore femur black, yellowish apically; fore tibia yellow     Lethades texanus (Ashmead)
5		Metasomal tergites black, or with narrow posterior margins light colored (Fig. 1    5
–		Middle metasomal tergites red, sometimes with dark maculae    11
6		Claw distinctly pectinate, teeth of hind claw more than 0.5 times as high as claw (Fig. 7). Longitudinal propodeal carina absent or complete    7
–		Hind claw with teeth less than 0.5 times as high as claw. Longitudinal propodeal carina present    8
7		Longitudinal propodeal carina mostly absent, only the area apicalis defined     Lethades punctatissimus (Strobl)
–		Longitudinal propodeal carina complete (Fig. 4)     Lethades orientalis Reshchikov & Xu, sp. n.
8		Head with parallel sides or expanded behind eyes dorsally. T1 with distinct dorsal longitudinal carinae reaching almost to posterior margin     Lethades erichsonii Hinz, 1996
–		Head narrowed behind eyes dorsally. T1 with weak dorsal longitudinal carinae reaching only 0.7 of length    9
9		Mesopleuron polished ventrally, finely and densely punctate. Propodeum with costula defined. T2 and T3 finely sculptured, polished. Metasomal tergites with narrow yellow posterior margins. Clypeus in female entirely or apically pale. Scape yellow ventrally     Lethades cingulator Hinz
–		Mesopleuron matt ventrally, shagreened or granulated    10
10		T2 and T3 with broad reddish-yellow bands on posterior margins. Pronotum, mesonotum, and mesopleuron in male with large yellow maculae. Female with clypeus and scape entirely black     Lethades laricis Hinz
–		T2 and T3 black (T3 slightly reddish-brown basally). Male with pronotum, mesonotum, and mesopleuron black. Clypeus in female with yellow maculae on sides     Lethades buriator Aubert
11		Ovipositor sheath 2 times as long as first tarsomere of hind leg     Lethades lapponicus (Holmgren)
–		Ovipositor sheath equal to or shorter than first tarsomere of hind leg    12
12		Temples and lower part of mesopleuron coriaceous and granulated with fine, dense punctures. T1 very densely punctate and striated, with elongate dorsal carinae reaching to its middle     Lethades facialis (Brischke)
–		Temples and lower part of mesopleuron striated, punctures not defined. T1 finely shagreened, with short dorsal carinae not reaching middle    13
13		Ovipositor sheath curved upwards, as long as first tarsomere of hind leg, curved upwards. Antenna with 24–26 flagellomeres; the basal flagellomeres stout, and apical flagellomeres transverse. T1 black; T2–T4 red     Lethades lapponator Hinz
–		Ovipositor sheath straight, shorter, 0.6–0.8 times as long as first tarsomere of hind leg. Antenna with 23–31 flagellomeres; the basal flagellomeres elongate, the apical flagellomeres cubic. T1 black with posterior margin red    14
14		Antenna with 28–31 flagellomeres. Female with third flagellomere 2.3–2.8 times as long as broad; male with third flagellomere 2.2–2.4 times as long as broad. Mesopleuron finely striated, finely and sparsely punctate. Body black. Palpi, mandibles, posterior edge of pronotum, and tegulae yellow. Legs (except coxae and hind tarsi), posterior margins of T1, and T2–T3 red. Male with clypeus, face, scape and pedicel ventrally, subtegular carina, fore and middle coxae and trochanters yellow     Lethades imperfecti Hinz
–		Antenna with 23–28 flagellomeres. Female with third flagellomere 1.9–2.4 times as long as broad; male with third falgellomere 1.9–2.2 times as long as broad. Mesopleuron distinctly coriaceous    15
15		Mesonotum and T1–T3 finely striated     Lethades scabriculus (Thomson)
–		Mesonotum and T1–T3 not striated    16
16		Mesonotum distinctly matt, finely and densely punctate     Lethades kukakensis (Ashmead)
–		Mesonotum distinctly polished, sparsely and indistinctly punctate     Lethades curvispina (Thomson)

Taxon Treatment

• Reshchikov, A; Xu, Z; Pang, H; 2017: First record of the genus Lethades Davis, 1897 from the Oriental region, with description of a new species (Hymenoptera, Ichneumonidae, Ctenopelmatinae) ZooKeys, (644): 43-50. doi

Images

Figures 1–7. Lethades orientalis Reshchikov & Xu, sp. n., female, holotype. 1 Habitus, lateral view 2 head, frontal view 3 head, dorsal view 4 propodeum, dorsal view 5 mesopleuron 6 apex of metasoma, lateral view 7 claw.