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- Lacconotus LeConte 1862: 255. – Gemminger and Harold 1870: 2179; Horn 1879: 338; Austin 1880: 41; LeConte and Horn 1883: 401; Fall 1901: 177; Dury 1902: 174; Blatchley 1910: 1302; Seidlitz 1917: 99; Leng 1920: 240; Leng and Mutchler 1933: 25, 36; Blair 1928: 33; Spilman 1951: 48; Spilman 1952: 10-11; Spilman 1954: 89; Arnett 1963: 717; Hatch 1965: 88; Campbell 1991: 267; Lawrence and Newton 1995: 896; Poole and Gentili 1996: 315; Arnett 2000: 473; Pollock 2002: 532; Bouchard et al. 2011: 443. Type species: Lacconotus punctatus LeConte, by monotypy.
[note: “Lacconotus” indicates the character states for subgenus Lacconotus, while “Alcconotus” refers to the corresponding states in subgenus Alcconotus (see below)].
Body elongate oval (TL/GEW 2.8-3.4), parallel-sided to widened posterior of middle, slightly (Alcconotus) to moderately (Lacconotus) flattened dorsally. TL 4.6–7.5 mm.
Head relatively short, narrowed slightly posterior of eyes; eyes moderately large, distinctly convex, inner margins convergent anteriorly; facets moderately coarse, with intrafacetal setae (especially noticeable in Alcconotus); frontoclypeal suture indistinct, indicated by slight furrow only; antennal insertions slightly concealed dorsally by raised lateral margins of frons; labrum transverse, anterior margin straight to shallowly sinuate.
Antennae relatively short (Fig. 1) (Lacconotus) to moderately elongate (Figs 2–3) (Alcconotus), not exhibiting distinct sexual dimorphism; scape and pedicel moniliform (Lacconotus) to slightly elongate (Alcconotus); antennomere 3 elongate, antennomeres 4–10 wider than long, submoniliform (Lacconotus) to elongate, triangular to subserrate (Alcconotus); antennomere 11 narrowed distally; sensilla present on distal surface of antennomeres 5–10. Mandibles relatively short, stout, slightly asymmetrical, apically bidentate; terebral teeth absent, or represented by several minute crenulae; molae approximately equal in size, subquadrate, with slightly developed surface texture; both mandibles with abrupt incision distal of mola; ventral row of microtrichia absent; prostheca distinct, about half length of mandible, inserted near distal edge of mola; maxilla with galea slightly longer than lacinia; galea bluntly rounded distally, relatively densely pubescent; maxillary palpi elongate, apical palpomere securiform (Lacconotus) to slightly cultriform (Alcconotus); inner margins of palpomeres 1 and 2 straight (Lacconotus) to slightly sinuate (Alcconotus); mentum short, about 2 × wider than long, posterior suture straight (Alcconotus) or distinctly arcuate (Lacconotus).
Thorax. Prothorax subquadrate (Figs 1–3), slightly wider than long (GPW/PL = 0.90–1.29); pronotal margins straight and convergent anteriorly, to slightly arcuate and widest near midlength; pronotal disc flat to slightly convex, with variously developed shallow, paired depressions; lateral pronotal carinae absent, margin smooth; posterior margin with pair of small, deep, punctiform pits; prosternum anterior of procoxae short (Lacconotus) to moderately elongate (Alcconotus), surface flat to slightly sunken medially; intercoxal process short, knife-like, extended to about half length of procoxae; procoxae rounded (Lacconotus) to elongate (Alcconotus); procoxal cavities open externally and internally; protrochantin concealed. Elytra elongate, subovate, disc flat (Lacconotus) to slightly convex (Alcconotus), upper surface uniformly and moderately coarsely punctate and setose (slightly more coarse in Lacconotus), setae closely appressed to elytral surface; apical elytral patch present, but not conspicuous dorsally, not contrasting in color with respect to remainder of elytron; epipleuron narrow, traceable only to abdominal ventrite 3 or 4; mesosternum with posterior intercoxal process parallel-sided, extended posteriorly to near posterior extent of mesocoxae; mesocoxae narrowly but completely separated, trochantins exposed; coxal cavities partly closed laterally by mesepimera; metasternum large, convex, anterior margin with indistinct (Lacconotus) to distinct (Alcconotus) process, in contact with posterior mesosternal process; median impressed line distinct to at least half distance to anterior margin of metasternum; metendosternite with long, relatively wide stalk; anterior tendons inserted on anterior margin of metendosternite body; laminae large, produced and somewhat angulate laterally.
Wing (Figs 7–8) fully developed, membrane beyond distinct radial cell moderately short (esp. in Lacconotus); venation similar in both species, but wing membrane and veins relatively darkly pigmented in Alcconotus (Fig. 8), very pale in Lacconotus (Fig. 7); wedge cell present; 3 MP veins reaching wing margin, proximal to CuA+AA; pigmented patches (flecks) present near junction of RP and MP, and near radial cell (Alcconotus), indistinct in Lacconotus.
Legs well developed, similar in relative shape and size on all thoracic segments; middle and hind femora slightly more expanded than front femora; femora relatively slender, but distinctly widened toward midlength; tibiae straight, about same length as femora, tibial spurs very short, equal in length; tarsomeres slender, 5–5-4; all tarsomeres simple ventrally, except for penultimate tarsomere with large ventral lobe; basal tarsomere on hind tarsus subequal in length to other tarsomeres combined; tarsal claws with large blunt tooth.
Abdomen with all ventrites freely articulated, uniformly punctate and setose, except for male sex patch; sex patch of two forms: small, longitudinally oval, setose patch on ventrite 2, not contrasting in color with ventrite (Fig. 6) (Alcconotus); or large, somewhat bulbous, glabrous area occupying and somewhat distorting the shape of ventrite 2, distinctly contrasting in color to dark ventrite surface (Fig. 5) (Lacconotus).
Male genitalia with median lobe dorsal to tegmen; sternite 9 forming ring-like sclerite, U-shaped in Alcconotus (Fig. 11), Y-shaped in Lacconotus (Fig. 9); tegmen relatively short, stout; basale broader than long, proximal margin deeply emarginate; length of apicale subequal to that of basale (along lateral margins); parameres of apicale short (Fig. 11) (Alcconotus) to slightly elongated (Fig. 9) (Lacconotus), with distal, inwardly-directed hook; median lobe (Figs 10, 12) stout, longer than tegmen; ventral side deeply emarginate, dorsal side proximally subquadrate, laterally produced, explanate; apex of median lobe triangular, relatively blunt.
Female genitalia with elongate, flexible, and only slightly sclerotized ovipositor; coxites 4-segmented, sparsely setose; distal segment short, distinctly more sclerotized than remainder of coxite; styli short, setose, with several very long distal setae; dorsal and ventral baculi well developed, extended to base of coxites; spiculum long, far exceeding length of segment 8; bursa copulatrix small (Fig. 13) (Lacconotus) to very large (Fig. 14) (Alcconotus), separated from vagina by narrow constriction, without conspicuous surface texture; spermatheca present, inserted near or at base of bursa, with elongate spermathecal gland.
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