Kunzea sinclairii (de Lange, Peter J. 2014)
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Ordo: Myrtales
Familia: Myrtaceae
Genus: Kunzea
Name
Kunzea sinclairii (Kirk) W. Harris – Wikispecies link – Pensoft Profile
- Kunzea sinclairii de Lange, Peter J., 2014, PhytoKeys 40: 1-1.
{{Section|class=etymology|heading = Etymology|content=Etymology. The specific epithet sinclairii honours Dr Andrew Sinclair (1794-1861) who, according to Kirk (1869, 1899), first discovered the species during a brief visit to the island. I have been unable to determine how Kirk came by that information, or when Sinclair's visit to Aotea (Great Barrier Island) happened, or why such an assiduous plant collector as Sinclair, so prominent in the early annals of early New Zealand botany (Hooker 1867; Cheeseman 1906, 1925), failed to procure specimens. = Leptospermum ericoides var. pubescens Kirk in T.P.N.Z.I. 1 (1869), 146-147.}}
Etymology
Etymology. The varietal epithet pubescens, though not elaborated on by Kirk (1869), probably refers to leaves, which he described in his paper as 'more or less pubescent'.
Description
Description (Figs 49, 50, 51).Growth habit mostly decumbent, trailing, silvery grey to grey, reddish-grey or grey-green, shrubs up to 3 x 1 m, very rarely forming a small tree up to 6 m tall; irrespective of stature, branches widely spreading and densely leafy, sometimes rooting on contact with soil or rock. Trunk 1(-4 or more), usually shortly erect between 0.2-1.0 m tall before branching but sometimes indistinguishable due to branches arising at ground level, 0.05-0.12(-0.16) m d.b.h.; basal portion of trunks covered with layers of somewhat firm to loose, stringy, pale grey to light brown chartaceous bark. Bark early bark dark brown to brown; firmly stringy, chartaceous to subcoriaceous, +/- elongate, lying in numerous overlapping strips; usually bearing a few transverse and many longitudinal cracks (especially on branch flanges and decurrent leaf bases), otherwise firmly attached; margins elongate, sinuous, +/- entire with scarcely any flaking; old bark initially dark brown to grey-brown, soon becoming covered in crustose lichens and sparse liverwort growth; coarsely stringy to tessellated and distinctly corky-coriaceous, usually remaining firmly attached, if detaching, then usually doing so along transverse cracks; flakes usually centrally attached, margins somewhat tabular with entire margins and coarsely frayed apices; upper surface of bark flakes coarsely tessellated, sometimes flaking secondarily as small tabular shards; upper trunk bark breaking into smaller pieces in hand but not crumbling. Branches numerous, usually present from close to or at trunk base, prostrate and widely spreading, new growth subscandent (in rare tree forms this habit is retained resulting in arching, pendulous branches); branchlets numerous, widely spreading to subscandent, often coarsely interwoven, initially red, +/- quadrangular to subterete, leaves usually densely crowded along stems and brachyblasts, though in vigorous new growth sometimes widely spaced; branchlets sericeous, indumentum copious, silky, hairs antrorse-appressed, weakly flexuose up to 0.06 mm long, hyaline to translucent (appearing silvery-white when young, maturing silver-grey). Vegetative buds inconspicuous, usually obscured from view by surrounding leaves; at resting stage 0.3-0.8 mm diam. narrowly ovoid to ellipsoid; scales deciduous; (0.3-)1.2 mm long, pale yellow-brown to reddish brown, broadly to narrowly ovate-lanceolate grading through lanceolate to narrowly lanceolate; midrib +/- keeled, sometimes prolonged to apiculate tip, otherwise apex obtuse to subacute or acute; oil glands inconspicuous, sparse, scattered in irregular lines either side of midrib; lamina initially completely invested by long silky silvery-white hairs, becoming glabrate, with hairs progressively confined to scale margins, midrib, and keel prolongation (if present). Leaves heterophyllous, weakly to strongly spicy-scented when crushed, mostly sessile, sometimes shortly petiolate (up to 1.6 mm long). Seedling and juvenile leaves dark green to glaucous, glabrous up to 25.0 x 3.5 mm, oblanceolate to lanceolate, apex acute, often shortly mucronate, base attenuate, lateral veins evident, especially on seedling leaves, both surfaces distinctly gland-dotted, oil glands up to 480 on either surface. Mature leaves soon developing (depending on degree of exposure), densely crowded along branchlets and brachyblasts, particularly toward apices, initially obliquely ascending, soon suberect to widely spreading usually weakly recurved in distal 30%; lamina (5.6-)14.5(-20.6) x (2.0-)3.2(-4.5) mm, initially appearing silvery-white (due to dense hair covering), maturing silvery-grey to reddish grey (as some hairs are shed), usually paler beneath; lamina broadly lanceolate, elliptic to obovate, rarely oblong-obovate, apex sharply acute, often cuspidate, base attenuate; adaxial lamina surface flat to weakly concave, glandular punctate, with oil glands scarcely evident when fresh due to dense hair covering, becoming more obvious in old leaves and in dried specimens, up to c.380, midrib slightly raised for c. 70% of leaf length; abaxial surface flat to weakly convex, usually densely covered in hairs, sometimes glabrate in old leaves, glandular punctate, oil glands up to 300; midrib raised for entire length; lamina margins distinctly less hairy than lamina surface; hairs of midribs and margins converging at leaf apex. Perules deciduous (shedding very early in inflorescence maturation), (0.8-)1.2(-1.4) x (0.8-)1.0(-1.2) mm, orange brown to amber with a broad pale brown margin (this reducing in thickness toward apex); broadly ovate grading through to ovate-lanceolate, apex cuspidate; lamina 6-8-nerved with poorly defined midrib and bearing up to 10-20 oil glands between nerves; lamina surface initially sparsely covered in deciduous long silky silvery-white hairs, soon becoming glabrate, except for a distinct stout weft on the cuspidate apex. Inflorescence mostly a compact, corymbiform (4-)9(-20)-flowered botryum 7.0-20.0 mm long and usually terminated by a tuft of leaves and a semi-dormant vegetative bud; inflorescences initially present on brachyblasts in the distal one-third of the active branchlets, increasing in abundance and soon dominating all the distal terminal and lateral growth; on occasion inflorescences may extend to elongated botrya on late season's vegetative growth. Inflorescence axis densely invested with antrorse-appressed, weakly flexuose, silky hairs. Pherophylls deciduous (shed early during bud maturation), rarely present at flowering, foliose or squamiform, basal portion tightly clasping pedicel base; (1.0-)1.2 x (0.2-)0.4 mm; foliose pherophylls pale green to red-green, oblong to oblong-lanceolate, very rarely broadly spathulate, cuspidate, deeply concave in cross section, with the abaxial surface copiously invested in sericeous, antrorse-appressed hairs; oil glands scarcely evident, up to 10 (usually less); midrib not evident; squamiform pherophylls tightly clasping pedicels, 0.3-1.0 x 0.4-0.8 mm, red-brown to brown, broadly to narrowly ovate or lanceolate, apex acute, subacute to obtuse, weakly keeled, margins and distal portion of keel finely ciliate. Pedicels (2.8-)5.7(-7.3) mm long at anthesis, scarcely elongating after anthesis, terete, initially invested with silky, antrorse-appressed, weakly flexuose, hairs becoming glabrate. Flower buds (2.3-)3.8(-4.9) x (2.1-) 3.1(-4.2) mm, ovoid to pyriform, apex flat to weakly domed prior to bud burst with calyx lobes held flat across surface, rarely meeting. Fresh flowers when fully expanded (5.7-)8.1(-10.2) mm diam. Hypanthium (1.9-)2.6(-3.6) x (2.1-)3.1(-4.2) mm, with free portion 0.4-0.7 mm long, silvery-white to silvery grey or reddish-grey due to copious covering of hairs; narrowly obconic to obconic or cupular, terminating in a scarcely defined chartaceous rim bearing five persistent calyx lobes; hypanthium surface smooth when fresh becoming irregularly wrinkled when dry, somewhat finely glandular punctate, oil glands scarcely evident due to dense covering of long, silky, antrorse-appressed silvery hairs, ribs not evident. Calyx lobes 5, initially erect to suberect, sometimes spreading, submembranous, (1.1-)1.3(-1.6) x (0.9-)1.2(-1.8) mm, broadly obtuse, red-green to pale green with a white or pink membranous margin, not obviously keeled, sparsely and finely gland-dotted, oil glands +/- colourless; lobe margins finely ciliate, hairs eglandular, central portion of lobes densely covered in short silky, antrorse-appressed hairs. Receptacle greenish pink or pink at anthesis, darkening to crimson after fertilisation. Petals 5(-6), (2.0-)2.9(-3.6) x (2.1-)2.7(-3.3) mm, white, very rarely basally flushed pink, broadly ovate, suborbicular to orbicular, rarely +/- cuneate-truncate, spreading, upper third often weakly recurved, apex rounded, margins +/- finely and irregularly crumpled or frayed, oil glands not evident in fresh or dried material. Stamens 18-38(-46) in 1-2 weakly defined whorls, arising from receptacular rim, filaments white. Antipetalous stamens (2-)3(-4), antisepalous (2-)4(-6). Outermost antipetalous stamens outcurved, widely spreading, more rarely slightly incurved, on filaments 2.0-3.6 mm long, inner stamen if present, 0.4-0.9 mm, outcurved or incurved. Antisepalous stamens usually shorter than outermost antipetalous stamens, 0.6-3.6 mm, weakly incurved or outcurved, usually in mixtures of both. Anthers dorsifixed, 0.06-0.1 x 0.06-0.09 mm, broadly ellipsoid to scutiform, latrorse. Pollen white (11.9-)15.4(-19.9) μm. Anther connective gland pale pink when fresh, drying pale orange, spheroidal, coarsely papillate. Ovary (3-)4(-5) locular, each with 18-22(-34) ovules in two rows on each placental lobe. Style 1.8-2.5(-3.0) mm long at anthesis, elongating slightly after anthesis, white basally flushed pink or pale pink; stigma narrowly capitate, as wide as or scarcely wider than style, +/- flat, greenish-pink or pink, flushing red after anthesis, surface finely granular-papillate. Fruits long persistent, copiously covered in short, silky, antrorse-appressed hairs; (2.2-)3.0(-3.6) x (2.7-)3.2(-3.9) mm, initially graphite grey, maturing to charcoal at dehiscence, and in old dehisced capsules fading to greyish-white; narrowly obconic to obconic, rarely cupular, calyx valves persistent, incurved, somewhat chartaceous, splits concealed by dried, erect, free portion of hypanthium. Seeds 0.52-1.04(-1.09) x 0.38-0.58(-0.72) mm, obovoid, oblong, or oblong-ellipsoid, usually curving toward apex, laterally compressed, 2-3-angled with convex to flattened faces, apex rounded to subacute; base oblique, +/- flattened; testa semi-glossy, orange-brown to dark brown, surface coarsely reticulate. FL: (Sep-)Nov-Jan(-Mar). FT: Feb-May(-Jul). Chromosome Number n = 11II, 2n = 22 (see de Lange and Murray 2004).
Distribution
Distribution (Fig. 7). Endemic, New Zealand, Aotea (Great Barrier Island) (20-510 m a.s.l.). On Aotea (Great Barrier Island) found mostly on the rhyolite outcrops of the central high points and western slopes (de Lange and Norton 2004).
Taxon Treatment
- de Lange, Peter J.; 2014: A revision of the New Zealand Kunzeaericoides (Myrtaceae) complex, PhytoKeys 40: 1-1. doi
This treatment was originally uploaded by Plazi, compare this treatment on Plazi. Unless this treatment has been substantially changed on Species-ID, Plazi requests to maintain a link back to the original repository.
