Kadua nukuhivensis (Wagner, Warren L. & Lorence, David H. 2011)
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Ordo: Gentianales
Familia: Rubiaceae
Genus: Kadua
Name
Kadua nukuhivensis (Lorence & J. Florence) W. L. Wagner & Lorence, Syst. Bot. 30: 832, 2005. – Wikispecies link – Pensoft Profile
- Kadua nukuhivensis Wagner, Warren L., 2011, PhytoKeys 4: 125-125.
Description
Description. Glabrous shrubs or small trees 2.5-6 m tall, trunk to 20 cm in diam., leafy twigs cylindrical, 7-9 mm in diam., internodes compressed, bark grayish- to blackish-brown. Leaves of a pair equal to sometimes unequal, blade obovate to obovate elliptic or oblong, 8-18 x 6-10.5 cm, chartaceous to subcoriaceous, drying brown or blackish-brown, base cuneate, decurrent, apex obtuse to rounded, secondary veins 7-9 pairs, loosely brochidodromous, prominulous above, tertiary veins slightly prominulous, higher order venation obscure, midrib sulcate above, rounded below, margin entire, plane to slightly revolute; petioles 8-13 x 2-3 mm in diam., sulcate; stipules inter- and intrapetiolar, sheath truncate, cupuliform, 2.5-3 x 5-6 mm, caducous but becoming thickened and horseshoe-like on adaxial surface of petioles. Inflorescences terminal and rarely also axillary in upper leaf axils, cymose, corymbiform, 7-13 cm long including corollas, on a stout, compressed peduncle 3-6 cm long or sessile with a pair of basal branches, often subtended by a pair of reduced subsessile leaves, bracts foliaceous, 1.5-3 cm long, caducous. Flowers 12-30, possibly dimorphic, fragrant when fresh, subtended by caducous triangular bracts c. 1 mm long, hypanthium obconical, 3 mm long, calyx limb cupuliform, 2 mm long, calyx lobes broadly triangular, 0.3-0.5 mm long; corolla fleshy, white or pinkish white when fresh, apex flat to slightly depressed in bud, at anthesis salverform, tube 26-28 x 2 mm in diam. distally, lobes 4, linear-oblong, recurved, 10-15 x 3-5 mm, apex with a recurved hook-like appendage; stamens with anther tips exserted for 1.5 mm, mucronulate, linear, 3-4 x 1 mm, attached 1.5-2 mm below apex of tube; style 17-19 mm long, included, stigmatic lobes 2, adnate, 3 mm long. Capsules on stout pedicels to 6 mm long, turbinate to obpyriform, 15-25 x 7-14 mm, strongly compressed, bisulcate, strongly 8-ribbed, 2/3 inferior, apex (beak portion above the calyx) obtuse, 5-6 mm long, smooth, dehiscence loculicidal then septicidal, old capsules disintegrating leaving network of vascular bundles enclosing the persistent endocarp. Seeds 1-1.3 x 0.5-0.9 mm, irregularly angulate to broadly ellipsoid, trigonous, margins thin, testa dull, dark brown, surface finely papillose.
Distribution
Distribution. Marquesas Islands, Nuku Hiva, where known only from the island's central mountain crest, on the leeward side of the Terre Deserte in the upper Tapuaehu Valley, between 1000 and 1065 m elevation.
Etymology
Etymology. The specific epithet refers to the only known island of occurrence for this species.
Discussion
Discussion. Kadua nukuhivensis resembles Kadua tahuatensis, and molecular evidence places these two as sister species in the same clade within the larger clade of Hawaiian and French Polynesian species (Motley 2003)
Taxon Treatment
- Wagner, Warren L.; Lorence, David H.; 2011: Revision of Kadua (Rubiaceae) in the Marquesas Islands, French Polynesia, with description of the new species K. lichtlei, PhytoKeys 4: 125-125. doi
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