Iberomaimetsha nihtmara
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Ordo: Hymenoptera
Familia: Maimetshidae
Genus: Iberomaimetsha
Name
Iberomaimetsha nihtmara Ortega-Blanco, Delclòs & Engel sp. n. – Wikispecies link – ZooBank link – Pensoft Profile
Type material
Holotype MCNA 13049 (?); paratypes MCNA 8758(♀?), MCNA 8790 (♂), MCNA 9373 (?), MCNA 9918 (♀), and MCNA 10732 (?). All specimens deposited in the Museo de Ciencias Naturales de Álava, Vitoria-Gasteiz, Spain.
Type locality
Peñacerrada I, Moraza, Burgos Province, Basque-Cantabrian Basin, Spain.
Stratigraphic horizon
Escucha Formation, Lower Cretaceous, Lower Albian.
Diagnosis
Mandibles distinctly large, exserted, asymmetrical, right 4-toothed, left 3-toothed, arched inwards (teeth pointing toward teeth of opposite mandible); basal flagellomeres (from F1 to F3–4) distinctly larger than apical flagellomeres (F1–F4 about 2–1.5× the length of F7–F14); protibial distal margin with a comb of stiff setae; cell [1M] square or rhomboidal, distinctly smaller than [2Cu] which has a short but distinct connection between 1M+Cu and M (then, [2Cu] composed by five aristae); 2Rs+M longer than 1m-cu and longer than cell [1Rs]; apicalmost sector of Cu arising from almost apical posterior margin of [2Cu]; metasoma moderately short, as long as mesosoma.
Description
Body length (from paratypes, as incomplete metasoma in holotype) around 2.55–2.61 mm. Head rounded in anterior view, length 0.41 mm, width 0.68 mm with a short occipital carina and apparently some polygonal dorsal sculpture; compound eyes not bulging, slightly oval, separated dorsally by 0.39 mm; toruli about at same distance to inner margin of compound eyes and each other; antennae 2.54 mm in length, inserted on frons below midpoint of compound eyes; scape and pedicel short and globular, flagellomeres cylindrical; lengths of antennomeres as follows (all in mm): scape 0.12, pedicel 0.09, flagellomeres 0.29, 0.28, 0.27, 0.23, 0.20, 0.18, 0.15, 0.14, 0.12, 0.12, 0.11, 0.11, 0.10, 0.12; malar space short, length approximately 0.11 mm. Mesosoma (dorsal view) 0.95 mm in length; pronotum not distinct dorsally; mesoscutal length 0.27 mm, with parallel notauli and longitudinal medial line, all three present as linear series of grooves more than well-delimited lines (Fig. 9); axillae wide and long, well distinct, separating completely mesoscutellum and mesoscutum; mesoscutellum distinctly convex, length 0.22 mm; propodeum 0.29 mm in length, areolate (seen in paratype MCNA 8790). Legs distinctly elongate; division of trochantellus not visible; tarsi with no plantar lobes visible; leg measurements (all in mm): profemur 0.82, protibia 0.46, protarsomeres I–V 0.34/0.15/0.10/0.09/0.10; mid-legs very poorly preserved but apparently with mesofemur 0.68, mesotibia ?, mesotarsomeres 0.24/0.12/0.10/?/?; metafemur 1.0, metatibia 0.83, metatarsomeres 0.46/0.20/0.15/0.09/0.12. Forewing hyaline, length 2.1 mm, maximum width 0.80 mm, densely covered by microtrichia; all veins tubular except for nebulous 2rs-m and 2m-cu, thus, only cells [2Rs] and [2M] pseudo-opened apically (Figs 9, 10); 1rs-m not aligned with r-rs; 2m-cu curved S-like; veins M and Cu reaching wing margin (although M somewhat nebulous), with apicalmost sector of Cu slightly angled posteriorly; cell [1R1] longer than both cells [1Rs]+[2Rs]; cell [1Rs] almost ¾ the surface of cell [1M] and almost half of [2Rs]; vein r-rs emerging at two-thirds from base of pterostigma; cell [2R1] short, with vein Rs slightly anteriorly bent distad 2rs-m. Hind wing difficult to observe, hyaline and densely covered by microtrichia, length approximately 1.22 mm, maximum width approximately 0.41 mm; three strong hamuli present on costal vein in distal half of wing. Metasoma (checked on paratype MCNA 8790) pedunculate, short and slightly flattened, length 0.98 mm, greatest height 0.41 mm; sterna hard, convex; no apparent metasomal armature on sternum II or III.
Male: Parameres spatulate, with apparently three short stiff setae apically (Fig. 10).
Etymology
The specific epithet is a noun in apposition and is the Anglo-Saxon word for nightmare, in reference to the terrifying appearance of the species in frontal view and with its enlarged mandibles.
Original Description
- Perrichot, V; Ortega-Blanco, J; McKellar, R; Delclòs, X; Azar, D; Nel, A; Tafforeau, P; Engel, M; 2011: New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae) ZooKeys, 130: 421-453. doi
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