Himalcercyon mirus

From Species-ID
Jump to: navigation, search
Notice: This page is derived from the original publication listed below, whose author(s) should always be credited. Further contributors may edit and improve the content of this page and, consequently, need to be credited as well (see page history). Any assessment of factual correctness requires a careful review of the original article as well as of subsequent contributions.

If you are uncertain whether your planned contribution is correct or not, we suggest that you use the associated discussion page instead of editing the page directly.

This page should be cited as follows (rationale):
Jia F, Liang Z, Fikáček M (2020) A review of Himalcercyon stat. nov., with description of a new species from the Chinese Himalaya and an updated key to Asian genera of Megasternini (Coleoptera, Hydrophilidae). Deutsche Entomologische Zeitschrift 67(1) : 35–49, doi. Versioned wiki page: 2020-05-11, version 183411, https://species-id.net/w/index.php?title=Himalcercyon_mirus&oldid=183411 , contributors (alphabetical order): Pensoft Publishers.

Citation formats to copy and paste

BibTeX:

@article{Jia2020DeutscheEntomologischeZeitschrift67,
author = {Jia, Fenglong AND Liang, Zulong AND Fikáček, Martin},
journal = {Deutsche Entomologische Zeitschrift},
publisher = {Pensoft Publishers},
title = {A review of Himalcercyon stat. nov., with description of a new species from the Chinese Himalaya and an updated key to Asian genera of Megasternini (Coleoptera, Hydrophilidae)},
year = {2020},
volume = {67},
issue = {1},
pages = {35--49},
doi = {10.3897/dez.67.50078},
url = {https://dez.pensoft.net/articles.php?id=50078},
note = {Versioned wiki page: 2020-05-11, version 183411, https://species-id.net/w/index.php?title=Himalcercyon_mirus&oldid=183411 , contributors (alphabetical order): Pensoft Publishers.}

}

RIS/ Endnote:

TY - JOUR
T1 - A review of Himalcercyon stat. nov., with description of a new species from the Chinese Himalaya and an updated key to Asian genera of Megasternini (Coleoptera, Hydrophilidae)
A1 - Jia F
A1 - Liang Z
A1 - Fikáček M
Y1 - 2020
JF - Deutsche Entomologische Zeitschrift
JA -
VL - 67
IS - 1
UR - http://dx.doi.org/10.3897/dez.67.50078
SP - 35
EP - 49
PB - Pensoft Publishers
M1 - Versioned wiki page: 2020-05-11, version 183411, https://species-id.net/w/index.php?title=Himalcercyon_mirus&oldid=183411 , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/dez.67.50078

Wikipedia/ Citizendium:

<ref name="Jia2020Deutsche Entomologische Zeitschrift67">{{Citation
| author = Jia F, Liang Z, Fikáček M
| title = A review of Himalcercyon stat. nov., with description of a new species from the Chinese Himalaya and an updated key to Asian genera of Megasternini (Coleoptera, Hydrophilidae)
| journal = Deutsche Entomologische Zeitschrift
| year = 2020
| volume = 67
| issue = 1
| pages = 35--49
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/dez.67.50078
| url = https://dez.pensoft.net/articles.php?id=50078
| pmc =
| accessdate = 2024-12-18

}} Versioned wiki page: 2020-05-11, version 183411, https://species-id.net/w/index.php?title=Himalcercyon_mirus&oldid=183411 , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Taxonavigation

Ordo: Coleoptera
Familia: Hydrophilidae
Genus: Himalcercyon

Name

Himalcercyon mirus (Hebauer, 2002) stat. nov. – Wikispecies link – Pensoft Profile

Type locality

Nepal, Kathmandu district, Sheopuri Mt., 2100–2300 m a.s.l. [GPS ca 27.816672N, 85.400000E].

Material examined

Holotype: NEPAL ● 1 ♂; Kathmandu Distr. Sheopuri Mt.; 2100–2300 m a.s.l.; 25 Jun 1988; W. Schawaller leg.; SMNS.
Paratypes: NEPAL ● 2 ♀♀; same data as for holotype; SMNS ● 1 ♀; same data as for holotype; NMPC ● 1 ♀; Annapurna, Telbrung Danda; 2600–2800 m a.s.l.; 13 Jun 1997; Schmidt leg.; SMNS.

Redescription

Form and color. Body size 3.1–3.5 mm (3.4 mm in holotype), body width 2.0–2.1 mm (2.0 mm in holotype), widest at anterior third of elytra, weakly narrowing posteriad (Fig. 1A). Dorsum pitchy-brown to black; head with paler clypeus; pronotal margins brown; elytral apices and posterior half of lateral elytral margins brownish; epipleuron pitchy brown laterally, reddish mesally; antenna, maxillary and labial palpi brown to reddish brown; legs reddish brown, with darker femora.
Head. Clypeus with moderately dense fine setiferous semicircular punctures, smooth between punctures. Frons with punctures of the same size and density as those on clypeus, smooth between punctures. Mentum 1.4× wider than long, rugose, with dense punctures (Fig. 2B), slightly concave anteriorly. Antenna with pedicel ca 0.2× as long as scape, pedicel ca. as long as antennomeres 3 and 4 combined, cupule small.
Thorax. Pronotum with punctation similar to that on frons, interstices without microsculpture; lateral marginal bead shortly overlapping to anterior margin but not to posterior margin, stopping at posterior angle. Scutellar shield smooth, with five to seven punctures. Elytral striae sharply impressed (Fig. 1A), striae 6, 8, and 9 not reaching base; intervals with finer and sparser punctures than on pronotum, each puncture bearing a fine short seta, interstices between punctures smooth. Epipleuron with bare outer and pubescent inner portion delimited from each other by a fine ridge, inner pubescent part narrower than the outer part, reaching the level of posterior part of metaventrite (Fig. 1A). Mesoventral elevation arrowhead-shaped, ca 1.5× longer than wide, sparsely pubescent (Fig. 2F). Metaventrite with large median elevation, finely and sparsely punctate (Fig. 2E), interstices without microsculpture; lateral portions microsculptured, with sparse coarse punctures and dense pubescence. Legs with trochanters densely pubescent, femora with sparse and moderately coarse punctures, interstice between punctures with fine microsculpture consisting of transverse lines.
Male genitalia. Middle lobe of abdominal sternite IX narrow, shorter than lateral struts (Fig. 1D). Aedeagus (Fig. 1C) with median lobe ca as long as tegmen; paramere ca 1.5× as long as phallobase. Paramere gradually narrowed from base to apex, obliquely truncate apically, widened inwards to form a process with a few setae. Median lobe ca as wide as paramere, gradually narrowing in apical third, apex narrowly rounded, gonopore subapical.

Distribution

Known from two localities in central Nepal (Fig. 4). Key to Eastern Palaearctic and Oriental genera of the Megasternini The following key is mainly based in the ventral characters, namely the form of prosternum and meso- and metaventrite, which are illustrated in Figures 5–8. The concept of some of the genera will likely be modified in the future; the key reflects the current status. The key includes all genera occurring east of Iran, the Black Sea, and the Ural Mountains. (i.e. it does not cover the Near East and the Arabian Peninsula); eastwards it includes all regions west of New Guinea. See Table 1 for the number of described species and references to the most important keys or taxonomic treatments for each genus. Remarks and numbers of species only refer to those from the Eastern Palaeartic and Oriental Regions.

Key to Eastern Palaearctic and Oriental genera of the Megasternini — F. Jia , Z. Liang , M. Fikáček
1
Antennal grooves large, reaching to the lateral margin of hypomeron (Fig. 5A, B, D). 
  2
–
Antennal grooves absent or small, not reaching to the lateral margin of the hypomeron (Figs 5E, F, 6, 7, 8A–C) 
  5
2
Metaventrite with complete femoral lines reaching from posteriomesal portion to anterolateral corner (Fig. 5A, D) 
  3
–
Metaventrite without complete femoral lines, at most with short vestiges anterolaterally. (Fig. 5B, C) 
  4
3
Mesoventral plate wider than long. Prosternum with wide plate without median carina (Fig. 5A). Mentum with sharply pointed anterolateral corners (Fig. 8D) 
  Cryptopleurum Mulsant
–
Mesoventral plate approximatley as long as wide. Prosternal plate approximately as long as wide, with more or less distinct median carina (Fig. 5D). Mentum with bluntly rounded anterolateral corners 
  Pachysternum Motschulsky
4
Median portion of prosternum roof-like, high (Fig. 5C). Mesoventral plate longer than wide. Metaventrite without any traces of femoral lines (Fig. 5C). Anterior tibia without anterolateral excision 
  Pacrillum d'Orchymont
–
Median portion of prosternum with flat hexagonal plate, not carinate medially (Fig. 5B). Mesoventral plate slightly wider than long. Metaventrite with vestiges of femoral lines in anterolateral corners (Fig. 5B). Anterior tibia anterolaterally with emargination 
  Megasternum Mulsant
5
Metaventrite with postcoxal ridge widely diverging from posterior margin of coxal cavity and forming an arcuate ridge reaching lateral margin of metaventrite (Figs 5E, F, 6A, B) 
  6
–
Metaventrite with postcoxal ridge parallel to posterior margin of coxal cavity or nearly so, reaching anterolateral corner of metaventrite and not forming any arcuate ridge (Figs 6C–F, 7, 8A–C) 
  9
6
Metaventrite with complete femoral lines crossing the arcuate postcoxal ridge and X-shape in form (Fig. 5E). Mesoventral elevation narrowly elongate or narrow but widely contacting metaventrite 
  Peltocercyon d'Orchymont
–
Metaventrite without X-shaped structure, femoral lines absent or short, not crossing with arcuate postcoxal ridge (Figs 5F, 6A, B). 
  7
7
Mesoventral plate widely contacting metaventrite (Fig. 6A, B). Median portion of prosternum at least weakly delimited from lateral portions 
  8
–
Mesoventral plate separated from metaventrite by a wide deep gap (Fig. 5F). Median portion of prosternum simply carinate, not delimited from lateral portions 
  Armostus Sharp
8
Metaventrite with deep triangular impression along its lateral margin (Fig. 6A) 
  Morastus d'Orchymont
–
Metaventrite without such impression (Fig. 6B) 
  Oosternum Sharp
9
Median portion of prosternum highly elevated and/or delimited from lateral portions by sharp ridges (Figs 6C–F, 7A–D) 
  10
–
Median portion of prosternum finely carinate, not delimited from lateral portions (Figs 7E, F, 8A, B) 
  18
10
Pronotum with deep longitudinal grooves (Fig. 8E). Bare portion of metaventrite very wide (Fig. 6C). Tiny beetles: length ca 1.2 mm 
  Emmidolium d'Orchymont
–
Surface of pronotum without distinct longitudinal depressions. Bare portion of metaventrite confined to medial part only. Tiny to moderately large beetles 
  11
11
Median portion of prosternum in form of very small triangular, very highly elevated projection. Antennal grooves absent (Fig. 6D). Abdomen with apical emargination 
  Chimaerocyon Fikáček, Maruyama, Vondráček & Short
–
Median portion of prosternum never so tiny and not so highly elevated. Antennal grooves present, even though sometimes rather small. Abdomen never with apical emargination 
  12
12
Prosternal elevation with lateral margins deeply excised (Fig. 6E, F) 
  13
–
Prosternal elevation with lateral margins or ridges straight (Fig. 7A–D) 
  14
13
Tiny species, 1.2–1.6 mm. Metaventrite with complete femoral lines (Fig. 6E). Antennal grooves present 
  Paroosternum Scott
–
Large species, ca 3.0 mm. Metaventrite without femoral lines (Fig. 6F). Antennal grooves absent 
  Oreosternum nom. nov.
14
Elytral series deeply impressed with the impressions contiguous to anterior margin of each elytron (Fig. 8F, G). Mesoventral elevation longer than wide, rhomboid to suboval (Fig. 7A, B) 
  15
–
Elytral series not impressed or impressions of elytral striae series not reaching anterior margin of each elytron. Mesoventral elevation elongate or as long as wide 
  16
15
Pronotum highly bulged in lateral view, not forming a continuous curve with elytra. Anterior margin of prosternal elevation strongly projecting anteriad (Fig. 7A). Mesoventral elevation subrhomboid 
  Bolbonotum Hansen
–
Pronotum not highly bulged in lateral view, forming a continuous curve with elytra. Anterior margin of prosternal elevation straight (Fig. 7B). Mesoventral elevation suboval 
  Kahanga Hansen
16
Grooves for reception of procoxae ending far before the anterior margin of mesocoxal cavities (Fig. 8C). Mesoventral plate elongate 
  Gillisius d'Orchymont (part)*
–
Grooves for reception of proxocae reaching nearly the mesocoxal cavities (Fig. 7C, D). Mesoventral elevation approximately as wide as long 
  17
17
Mesoventral elevation nearly semi-elliptical (Fig. 7C), with wide marginal rim. Postcoxal ridges on the metaventrite meeting mesally and forming a short median longitudinal ridge. Metatibiae densely pubescent ventrally (Fig. 8H). Large species: 2.5–3.3 mm 
  Australocyon Hansen
–
Mesoventral elevation more less pentagonal, without any marginal rim (Fig. 7D). Postcoxal ridges mesally bending posteriad, remaining separate, forming two short median longitudinal ridges (in one species largely obsolete). Metatibie without dense ventral pubescence. Medium sized to tiny species: 2.0–2.9 mm 
  Nipponocercyon Satô
18
Abdominal ventrite 1 without median carina. Mesoventral elevation narrowly laminar (Fig. 7E) 
  Cycreon d'Orchymont
–
Abdominal ventrite 1 carinate medially. Mesoventral elevation in form of a lamina or an elongate plate 
  19
19
Ventral face of meso- and metatibiae with dense, long pubescence. Ventral morphology similar to Figure 7F 
  Pilocnema Hansen
–
Ventral face of meso- and metatibiae never densely pubescent, at most with sparse short setae. Ventral morphology similar to Figures 2, 3, and 8A, B 
  20
20
Mesoventral elevation laminar or forming an oval elongate plate; posterior part of the plate rounded or acute (as in Fig. 8A, B) 
  21
–
Mesoventral elevation elongate, but sharply cut off posteriorly, contacting metaventrite more or less in a straight line (as in Figs 2F, 8C) 
  22
21
Median portion of prosternum with a pair of transverse ridges partly delimiting prosternal process (Fig. 8A) 
  Pseudocercyon d'Orchymont
–
Median portion of prosternum without such ridges, only simply carinate (Fig. 8B) 
  Cercyon Leach
22
Mesoventral elevation arrowhead-shaped, with lateral angulate lobes (Figs 2F, 3C) 
  Himalcercyon Hebauer
–
Mesoventral elevation elongate oval (as in Fig. 8C); if small lateral lobes are present, they are below the plate 
  23
23
India, continental Southeast Asia and China. 
  Gillisius d'Orchymont (part)
–
Islands of the Malay Archipelago. 
  Pelosoma Mulsant*
Table 1. List of Eastern Palaearctic and Oriental general of the Megasternini, with number of described species and references to the most important keys or taxonomic treatments.
Genus Described species Keys or original descriptions
Armostus 11 d’Orchymont 1942[1]; Hebauer 2002a[2]; Hoshina and Satô 2006[3]
Australocyon 7 Hansen 2003[4]; Fikáček et al. 2012[5]
Bolbonotum 3 Hansen 1999a[6]
Cercyon 148 Shatrovskiy 1992[7]; Hansen 1999b[8]; Short and Hebauer 2006[9]; Hoshina 2008[10]; Jia et al. 2011[11]; 2019[12]; Ryndevich et al. 2017[13]; 2019[14]; Ryndevich and Prokin 2017[15]
Chimaerocyon 2 Fikáček et al. 2013[16]
Cryptopleurum 7 d’Orchymont 1926[17]; Jia and Zhang 2017[18]
Cycreon 4 + 1 ssp. Arriaga-Varela et al. 2018b[19]
Emmidolium 1 Fikáček 2007[20]
Gillisius 2 d’Orchymont 1925a[21]; 1926[17]
Himalcercyon 2 this paper
Kahanga 1 Hansen 1999a[6]
Megasternum 4 Shatrovskiy 1989[22]; Fikáček et al. 2012[5]; Ryndevich 2017[23]
Morastus 1 d’Orchymont; 1926[17]
Nipponocercyon 3 Hoshina and Fikáček 2010[24]; Fikáček et al. 2012[5]; 2015a[25]
Oosternum 9 Hebauer 2002a[2]; Hoshina and Satô 2004b[26]; 2005[27]
Oreosternum 1 Hebauer 2002
Pachysternum 11 Fikáček et al. 2012[5]
Pacrillum 5 Hoshina and Satô 2004a[28]; Fikáček and Hebauer 2005[29]; Shatrovskiy 1989[22] as Agnaeformia
Paroosternum 5 Hebauer 2006[30]
Pelosoma 2 d’Orchymont 1925b[31]; 1932[32]
Peltocercyon 4 d’Orchymont 1925a[21]; Hoshina 2016[33]; 2018[34]
Pilocnema 1 Hansen 2003[4]
Pseudocercyon 1 d’Orchymont 1926[17]

New replacement name

Taxon Treatment

  • Jia, F; Liang, Z; Fikáček, M; 2020: A review of Himalcercyon stat. nov., with description of a new species from the Chinese Himalaya and an updated key to Asian genera of Megasternini (Coleoptera, Hydrophilidae) Deutsche Entomologische Zeitschrift, 67(1): 35-49. doi

Images

Figure 1. Habitus and genital morphology of Himalcercyon species. A–D.H. mirus (A, B. Paratypes; C, D. Holotype). E–H.H. franzi sp. nov. (E, F. Paratypes; G, H. Holotype). A, E. Dorsal view; B, F. Lateral view; C, G. Aedeagus in dorsal view; D, H. Sternite IX in dorsal view.
Figure 1. Habitus and genital morphology of Himalcercyon species. A–D.H. mirus (A, B. Paratypes; C, D. Holotype). E–H.H. franzi sp. nov. (E, F. Paratypes; G, H. Holotype). A, E. Dorsal view; B, F. Lateral view; C, G. Aedeagus in dorsal view; D, H. Sternite IX in dorsal view. 
Figure 2. Morphology of Himalcercyon mirus (Hebauer, 2002). A. Complete ventral view. B. Mentum. C. Prosternal carina in ventrolateral view. D. Prosternum and hypomeron. E. Meso- and metaventrite. F. Details of mesoventral plate.
Figure 2. Morphology of Himalcercyon mirus (Hebauer, 2002). A. Complete ventral view. B. Mentum. C. Prosternal carina in ventrolateral view. D. Prosternum and hypomeron. E. Meso- and metaventrite. F. Details of mesoventral plate. 
Figure 3. Morphology of Himalcercyon franzi sp. nov. A. Mentum. B. Prosternum and hypomeron. C. Mesoventral elevation. D. Antenna. E. Mesotarsus, ventral view. F. Median portion of metaventrite. G. Elytral punctation. H. Elytral apex.
Figure 3. Morphology of Himalcercyon franzi sp. nov. A. Mentum. B. Prosternum and hypomeron. C. Mesoventral elevation. D. Antenna. E. Mesotarsus, ventral view. F. Median portion of metaventrite. G. Elytral punctation. H. Elytral apex. 
Figure 4. Known distribution of Himalcercyon: Circles.H. mirus (Hebauer); Square.H. franzi sp. nov. Color shading of the map indicated altitude: green = lowest, brown = highest.
Figure 4. Known distribution of Himalcercyon: Circles.H. mirus (Hebauer); Square.H. franzi sp. nov. Color shading of the map indicated altitude: green = lowest, brown = highest. 
Figure 5. Ventral view of thorax of eastern Palaearctic and Oriental genera of the Megasternini. A.Cryptopleurum ferrugineum. B.Megasternum concinnum. C.Pacrillum manchuricum. D.Pachysternum nigrovittatum. E.Peltocercyon coomani. F.Armostus schenklingi.
Figure 5. Ventral view of thorax of eastern Palaearctic and Oriental genera of the Megasternini. A.Cryptopleurum ferrugineum. B.Megasternum concinnum. C.Pacrillum manchuricum. D.Pachysternum nigrovittatum. E.Peltocercyon coomani. F.Armostus schenklingi. 
Figure 6. Ventral view of thorax of eastern Palaearctic and Oriental genera of the Megasternini. A.Morastus gracilipalpis. B.Oosternum sp. (O. soricoides group). C.Emmidolium excavatum. D.Chimaerocyon shimadai. E.Paroosternum sp. F.Oreosternum frigidum.
Figure 6. Ventral view of thorax of eastern Palaearctic and Oriental genera of the Megasternini. A.Morastus gracilipalpis. B.Oosternum sp. (O. soricoides group). C.Emmidolium excavatum. D.Chimaerocyon shimadai. E.Paroosternum sp. F.Oreosternum frigidum. 
Figure 7. Ventral view of thorax of eastern Palaearctic and Oriental Megasternini. A.Bolbonotum sp. B.Kahanga inconspicua, holotype. C.Australocyon sp. (A. pilocnemoides group). D.Nipponocercyon shibatai. E.Cycreon floricola. F.Pilocnema sp.
Figure 7. Ventral view of thorax of eastern Palaearctic and Oriental Megasternini. A.Bolbonotum sp. B.Kahanga inconspicua, holotype. C.Australocyon sp. (A. pilocnemoides group). D.Nipponocercyon shibatai. E.Cycreon floricola. F.Pilocnema sp. 
Figure 8. Ventral view of thorax and additional diagnostic characters of eastern Palaearctic and Oriental genera of the Megasternini. A–C. Ventral view: A.Pseudocercyon andrewesi. B.Cercyon sp. C.Gillisius madurensis, holotype. D–H. Other characters: D.Cryptopleurum coomani, mentum. E.Emmidolium excavatum, pronotum. F.Bolbonotum sp., base of elytra. G.Kahanga inconspicua, holotype, base of elytra. H.Australocyon sp. (A. pilocnemoides group), hind femur.
Figure 8. Ventral view of thorax and additional diagnostic characters of eastern Palaearctic and Oriental genera of the Megasternini. A–C. Ventral view: A.Pseudocercyon andrewesi. B.Cercyon sp. C.Gillisius madurensis, holotype. D–H. Other characters: D.Cryptopleurum coomani, mentum. E.Emmidolium excavatum, pronotum. F.Bolbonotum sp., base of elytra. G.Kahanga inconspicua, holotype, base of elytra. H.Australocyon sp. (A. pilocnemoides group), hind femur. 

Other References

  1. ↑ d’Orchymont A (1942) Palpicornia (Coleoptera). Notes diverses et espèces nouvelles. III.Bulletin du Musée Royal d’Histoire Naturelle de Belgique18(26): 1–20.
  2. ↑ 2.0 2.1 Hebauer F (2002a) Hydrophilidae of northern India and southern Himalaya (Coleoptera: Hydrophilidae).Acta Coleopterologica18(1): 3–72.
  3. ↑ Hoshina H, Satô M (2006) A taxonomic study of the genus Armostus (Coleoptera: Hydrophilidae) from Japan.The Coleopterists Bulletin60: 95–104. https://doi.org/10.1649/840.1
  4. ↑ 4.0 4.1 Hansen M (2003) Discovery of Australocyon Hansen and Pilocnema Hansen (Coleoptera, Hydrophilidae) outside the Australian region. In: Cuccodoro G Leschen R (Eds) Systematics of Coleoptera: papers celebrating the retirement of Ivan Löbl.Memoirs of Entomology International Associate Publisher17: 53–84.
  5. ↑ 5.0 5.1 5.2 5.3 Fikáček M, Jia F, Prokin A (2012a) A review of the Asian species of the genus Pachysternum (Coleoptera: Hydrophilidae: Sphaeridiinae).Zootaxa3219(1): 1–53. https://doi.org/10.11646/zootaxa.3219.1.1
  6. ↑ 6.0 6.1 Hansen M (1999a) Fifteen new genera of Hydrophilidae (Coleoptera), with remarks on the generic classification of the family.Entomologica Scandinavica30: 121–172. https://doi.org/10.1163/187631200X00228
  7. ↑ Shatrovskiy A (1992) New and little known Hydrophiloidea (Coleoptera) from Southern Primorye territory and adjacent regions.Entomological Review71(2): 359–371.
  8. ↑ Hansen M (1999b) Hydrophiloidea (s.str.) (Coleoptera). World catalogue of insects Volume 2: Hydrophiloidea (Coleoptera).Apollo Books, Stenstrup, 416 pp.
  9. ↑ Short A, Hebauer F (2006) World catalogue of the Hydrophiloidea – additions and corrections, 1 (1999–2005).Koleopterologische Rundschau76: 315–359.
  10. ↑ Hoshina H (2008) A new species of the genus Cercyon (Coleoptera: Hydrophilidae) from Fukui Pref., Japan.Memoirs of the Research and Education Center for Regional Environment, University of Fukui15: 1–7.
  11. ↑ Jia F, Fikáček M, Ryndevich S (2011) Taxonomic notes on Chinsese Cercyon: description of a new species, new synonyms, and additional faunistic records (Coleoptera: Hydrophilidae: Sphaeridiinae).Zootaxa3090: 41–56. https://doi.org/10.11646/zootaxa.3090.1.3
  12. ↑ Jia F, Liang Z, Ryndevich S, Fikáček M (2019) Two new species and additional faunistic records of Cercyon Leach, 1817 from China (Coleoptera: Hydrophilidae).Zootaxa4565: 501–514. https://doi.org/10.11646/zootaxa.4565.4.4
  13. ↑ Ryndevich S, Jia F, Fikáček M (2017) A review of the Asian species of the Cercyon unipunctatus group (Coleoptera: Hydrophilidae: Sphaeridiinae).Acta Entomologica Musei Nationalis Pragae57(2): 535–576. https://doi.org/10.1515/aemnp-2017-0089
  14. ↑ Ryndevich S, Hoshina H, Prokin A (2019) Review of species of the genus Cercyon of Russia and adjacent regions. VI. Subgenus Cercyon, the C. shinanensis group (Coleoptera: Hydrophilidae).Zoosystematica Rossica28(2): 258–266. https://doi.org/10.31610/zsr/2019.28.2.258
  15. ↑ Ryndevich S, Prokin A (2017) Two new species of Cercyon (Clinocercyon) from Russian Far East (Coleoptera: Hydrophilidae).Zootaxa4300(1): 125–134. https://doi.org/10.11646/zootaxa.4300.1.7
  16. ↑ Fikáček M, Maruyama M, Vondráček D, Short A (2013) Chimaerocyon gen. nov., a morphologically aberrant myrmecophilous genus of water scavenger beetle (Coleoptera: Hydrophilidae: Sphaeridiinae).Zootaxa3716(2): 277–288. https://doi.org/10.11646/zootaxa.3716.2.8
  17. ↑ 17.0 17.1 17.2 17.3 d’Orchymont A (1926) Contribution à l’étude des hydrophilides VI.Bulletin et Annales de la Société Entomologique de Belgique66: 201–248.
  18. ↑ Jia F, Zhang R (2017) A review of the genus Cryptopleurum from China (Coleoptera: Hydrophilidae).Acta Entomologica Musei Nationalis Pragae57(2): 577–592. https://doi.org/10.1515/aemnp-2017-0090
  19. ↑ Arriaga-Varela E, Wong S, Kirejtshuk A, Fikáček M (2018b) Review of the flower-inhabiting water scavenger beetle genus Cycreon (Coleoptera, Hydrophilidae), with descriptions of new species and comments on its biology.Deutsche Entomologische Zeitschrift65: 99–115. https://doi.org/10.3897/dez.65.26261
  20. ↑ Fikáček M (2007) Emmidolium excavatum Orchymont (Coleoptera: Hydrophilidae: Sphaeridiinae) confirmed in Africa and the Arabian Peninsula.Acta Entomologica Musei Nationalis Pragae47: 117–122.
  21. ↑ 21.0 21.1 d’Orchymont A (1925a) Contribution à l’étude des hydrophilides III.Bulletin et Annales de la Société Entomologique de Belgique65: 261–295.
  22. ↑ 22.0 22.1 Shatrovskiy A (1989) 12. Sem. Hydrophilidae – Vodolyubi. [12. Fam. Hydrophilidae – Water scavenger beetles]. In: Ler P (Ed.) Opredelitel’ Nasekomykh Dal’ Nego Vostoka SSSR v Shesti Tomakh.Tom 3. Zhestkokrylye, ili zhuky. Chast 1. [Keys to the insects of the Far East of the USSR in six volumes (Vol. 3). Coleoptera or beetles. Part 1]. Nauka, Leningrad, 264–293. [in Russian]
  23. ↑ Ryndevich S (2017) New faunistic records of hydrophilid beetles (Coleoptera: Hydrophiloidea: Hydrophilidae) from Eurasia. BarSU Herald.Series of Biological Sciences (General Biology), Agricultural Sciences (Agronomy)5: 65–70.
  24. ↑ Hoshina H, Fikáček M (2010) Morphological study and taxonomic revision of the genus Nipponocercyon (Coleoptera: Hydrophilidae: Sphaeridiinae).Acta Entomologica Musei Nationalis Pragae50: 117–130.
  25. ↑ Fikáček M, Angus R, Gentili E, Jia F, Minoshima Y, Prokin A, Przewozny M, Ryndevich S (2015a) Helophoridae, Hydrochidae, Hydrophilidae. In: Löbl I Löbl D (Eds) Catalogue of Palaearctic Coleoptera (Vol.2). Brill, Leiden/Boston, 25–76.
  26. ↑ Hoshina H, Satô M (2004b) Two new species of the genus Oosternum (Coleoptera: Hydrophilidae) from Japan.Memoirs of the Research and Education Center for Regional Environment, University of Fukui11: 9–15.
  27. ↑ Hoshina H, Satô M (2005) Taxonomic notes of the genus Oosternum (Coleoptera: Hydrophilidae) from Japan.Memoirs of the Research and Education Center for Regional Environment Fukui University12: 1–9.
  28. ↑ Hoshina H, Satô M (2004a) First record of the genus Pacrillum (Coleoptera: Hydrophilidae) from Japan, with a redescription of P. manchuricum.The Entomological Review of Japan59: 233–239.
  29. ↑ Fikáček M, Hebauer F (2005) A new aberrant species of the genus Pacrillum from Nepal (Hydrophilidae: Sphaeridiinae: Megasternini).Acta Entomologica Musei Nationalis Pragae45: 59–64.
  30. ↑ Hebauer F (2006) Revision of the genus Paroosternum Scott, 1913 (Coleoptera: Hydrophilidae: Sphaeridiinae).Acta Coleopterologica22(2): 38–58.
  31. ↑ d’Orchymont A (1925b) Contribution à l’étude des hydrophilides IV.Bulletin et Annales de la Société Entomologique de Belgique65: 298–305.
  32. ↑ d’Orchymont A (1932) Zur Kenntnis der Kolbenwasserkäfer (Palpicornia) von Sumatra, Java und Bali. Archiv für Hydrobiologie, Supplement Band XI (Tropische Binnengewässer II): 623–714. [pls XIV–XVII]
  33. ↑ Hoshina H (2016) Taxonomic report of the genus Peltocercyon (Coleoptera: Hydrophilidae: Sphaeridiinae).Japanese Journal of Systematic Entomology22(1): 131–133.
  34. ↑ Hoshina H (2018) Discovery of second Japanese species of the genus Peltocercyon (Coleoptera: Hydrophilidae: Sphaeridinae) with description of a new species.Japanese Journal of Systematic Entomology24(2): 293–295.
Retrieved from "https://species-id.net/w/index.php?title=Himalcercyon_mirus&oldid=183411"