Helle
Notice: | This page is derived from the original publication listed below, whose author(s) should always be credited. Further contributors may edit and improve the content of this page and, consequently, need to be credited as well (see page history). Any assessment of factual correctness requires a careful review of the original article as well as of subsequent contributions.
If you are uncertain whether your planned contribution is correct or not, we suggest that you use the associated discussion page instead of editing the page directly. This page should be cited as follows (rationale):
Citation formats to copy and paste
BibTeX: @article{Winterton2012ZooKeys172, RIS/ Endnote: TY - JOUR Wikipedia/ Citizendium: <ref name="Winterton2012ZooKeys172">{{Citation See also the citation download page at the journal. |
Ordo: Diptera
Familia: Acroceridae
Name
Helle Osten Sacken, 1896 – Wikispecies link – Pensoft Profile
- Helle Osten Sacken, 1896: 16 – Hutton 1901[1]: 28; Paramonov 1955[2]: 21; Schlinger and Jefferies 1989[3]: 376. Type species: Acrocera longirostris Hudson, 1892: 56 by monotypy.
Diagnosis
Body length: 4.0–6.0 mm [male], 6.0–7.0 mm [female]. Body shape strongly arched; colouration non-metallic (brown or black); head size slightly narrower than thorax width, shape sub-spherical; postocular ridge and occiput rounded; three ocelli, anterior ocellus reduced in size; posterior margin of eye rounded; eye apilose; position of antennae on head near middle of frons; eyes contiguous above antennal base, not contiguous below antennal base; palpus present; proboscis greater than head length; flagellum stylate, apex with terminal seta; postpronotal lobes enlarged, medially contiguous to form collar; subscutellum enlarged; legs not elongated; wing markings absent; costa ending near wing apex, costal margin straight; humeral crossvein absent; radial veins straight or curved towards wing anterior margin; R1 inflated distally at pterostigma; pterostigma and cell r1 membranous, not ribbed; R2+3 present; R4+5 angled anteriorly approximately midway; cell r4+5 bisected by 2r-m, basal cell very narrow elongate, closed; 2r-m joining M1 to R4+5; cell r4+5 present, narrow elongate, closed (open apically when 2r-m rarely absent); crossvein 2r-m present (rarely absent);R4 without spur vein; medial vein compliment with M1, M2 and M3 present (M3 fused with CuA1); discal cell closed completely; medial veins not reaching wing margin; CuA1 joining M3, petiolate to margin; CuA2 fused to A1 before wing margin, petiolate; wing microtrichia absent; anal lobe well developed; alula well developed; abdominal tergites smooth, rounded; abdomen shape elongate, narrow cylindrical or conical (male), or rounded and inflated (female).
Included species
Helle longirostris (Hudson, 1892); Helle rufescens Brunetti, 1926.
Comments
Helle is an endemic genus to New Zealand that is closely related to Schlingeriella, the only other philopotine genus in the region (Gillung and Winterton 2011[4]; Winterton et al. 2007[5]). Characteristics supporting this closerelationship include thickening of wing vein R1 at the pterostigma, elongate mouthparts, apilose eyes, 2r-m absent (rarely in Helle) and R4+5 angled anteriorly approximately half way along vein. Helle can be differentiated from all other philopotine genera based on the relatively complete wing venation, inflated R1 at pterostigma, palpi present and apilose eyes.
Key to Helle species
Taxon Treatment
- Winterton, S; 2012: Review of Australasian spider flies (Diptera, Acroceridae) with a revision of Panops Lamarck ZooKeys, 172: 7-75. doi
Other References
- ↑ Hutton F (1901) Synopsis of the Diptera Brachyera of New Zealand. Trans. Proc. N. Z. Inst. 33: 1-95.
- ↑ Paramonov S (1955) New Zealand Cyrtidae (Diptera) and the problem of the Pacific island fauna. Pacific Science 9: 16-25.
- ↑ Schlinger E, Jefferies M (1989) Family Acroceridae. In: Evenhuis N (Ed). Catalog of Diptera of the Australasian and Oceanian regions. Bishop Museum Special Publication. Bishop Museum Press, 86: 375-37.
- ↑ Gillung J, Winterton S (2011) New genera of philopotine spider flies (Diptera, Acroceridae) with a key to living and fossil genera. ZooKeys 127: 15-27. doi: 10.3897/zookeys.127.1824
- ↑ Winterton S, Wiegmann B, Schlinger E (2007) Phylogeny and Bayesian divergence time estimations of small-headed flies (Diptera: Acroceridae) using multiple molecular markers. Molecular Phylogenetics and Evolution 43: 808-832.
Images
|