Eustochomorpha
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Ordo: Hymenoptera
Familia: Mymaridae
Name
Eustochomorpha Girault, 1915 – Wikispecies link – Pensoft Profile
Type species
Eustochomorpha haeckeli Girault, by monotypy.
Diagnosis
Female. Antenna with funicle 8-segmented and clava 2-segmented (Figs 3, 7, 14). Venation about 0.9× fore wing length. Postmarginal vein distinct, uniformly thick and about 0.7× marginal vein length, with at least 2 short, socketed setae (Fig. 8, apical one indicated by arrow, Fig. 20c). Hypochaeta possibly present (Fig. 20a), originating on the parastigma instead of on the costal cell. Ovipositor strongly extending posteriorly beyond gastral apex but not extending anteriorly under mesosoma (Figs 2, 4, 28, 30).
Description
Female. Body ≈1280–1290 μm in length, excluding section of ovipositor exserted beyond apex of hypopygium. Colour. Generally brown (Fig. 4), with a few metallic reflections (an artefact of lighting?) on gaster. Gaster at base and most parts of legs lighter brown to white. Wings hyaline, with venation light brown. Head. Head about 1.7× as wide as long, 1.3× as wide as high and 1.4× as high as long; in lateral view with anterior surface convex, receding strongly from ventral margin of eye to mouth opening, and bulging medially compared to laterally (at preorbital groove); posterior surface slightly, evenly convex (Fig. 4). Face about as wide as high (Fig. 5), without subantennal grooves; preorbital groove ventral to torulus straight then, more ventrally, slightly curving medially to ventrolateral margin of mouth opening. Torulus in slight triangular depression about 2.0× as high as torulus width and separated by 1.5× torulus width from transverse trabecula (Fig. 5). Vertex in lateral view (Fig. 11) sloping obliquely to transverse trabecula to form an obtuse angle with face, and vertex posteriorly merging smoothly into occiput medially but separated laterally from occiput by short transverse vertexal suture. Ocellar triangle raised slightly above level of rest of vertex, the mid ocellus almost vertical, the lateral ocelli oblique, facing posterolaterally; ocelli with POL about 2.0× LOL and about 1.26 OOL. Transverse trabecula separated at each apex from supraorbital trabecula (Fig. 5); preorbital trabecula (apparently absent because not sclerotized) short, extending to inner orbit at about dorsal margin of torulus (Fig. 5); supraorbital trabecula in 2 subequal sections, the posterior sections slightly converging and the sutures outside posterior sections continuing onto occiput as the occipital grooves (short lines each appearing as posterior extension of supraorbital suture and strongly converging towards occipital foramen) (Fig. 6) separating occiput medially from temple dorsolaterally. Occiput separated from vertex by vertexal suture, this only present laterally behind lateral ocelli (Fig. 12). Eye large with numerous (about 100) small facets, in lateral view slightly higher than wide (10:9) and clearly separated dorsally from back of head (temple about 0.3× eye width). Ocular apodeme fairly long, crooked, needle-like. Malar sulcus absent (Fig. 11). Gena at level of ventral margin of eye wider than malar space. Occiput entire, not separated by transverse groove into upper and lower portions (Fig. 6a). Mouthparts. Labrum with 2? setae; mandible with 3 teeth. Antenna. Scape about 2.6× as long as wide, with radicle distinct from rest of scape and about 0.25× total scape length (Fig. 13); pedicel about 2.1× as long as wide, almost as wide and about 0.7× as long as entire scape; funicle 8-segmented; clava 2-segmented (Figs 3, 7, 14), 1.08× as wide as apical funicle segment and 0.32–0.36× as long as entire funicle. Mesosoma. Mesosoma about 2.2× as long as wide, 2.3× as long as high and 1.1× as wide as high. Pronotum entire (Figs 9, 12, 15), in dorsal view clearly visible, medially about 0.3× as long as mesoscutum; collar almost horizontal, with a shallow median longitudinal impression and prominent angular shoulders; neck shorter than collar, posteriorly strongly sloping up to collar; in lateral view, pronotum rectangular, slightly overlapping anterior margin of mesoscutum, with lateral surface merging smoothly into dorsal surface, with a shallow, oblique impression for femur. Spiracle (Figs 17, 18) apparently slightly stalked, at posterolateral angle of pronotum in a slight depression and facing dorsolaterally. Propleura near anterior apex not quite abutting, then gap widening slightly more anteriorly. Prosternum rhomboidal, incompletely divided medially by faint longitudinal groove posteriorly. Mesoscutum about 1.7× as long as scutellum, in dorsal view with shallow, thin, slightly diverging notauli apparently absent posteriorly (Figs 9, 15, 16), in lateral view mesoscutum (Fig. 4) almost flat except anteriorly. Scutellum slightly wider than long; anterior scutellum about 3× as long as narrowly trapezoidal, transverse frenum and separated from it by a shallow, curved frenal depression (Figs 9, 16); campaniform sensilla small, slightly closer to lateral margins of anterior scutellum than to each other, with apex of large, cone-shaped fenestra projecting anteriorly between them (Fig. 9). Axilla slightly advanced, the transscutal articulation laterally forming an obtuse angle with median section (Fig. 16); axillula long and narrow, separated from anterior scutellum by posteriorly recurved axillular groove; mesophragma widely convex posteriorly, extending to posterior apex of propodeum. Prepectus probably spindle-shaped (Fig. 17, partly hidden by fore leg); mesopleuron somewhat rectangular, with shallow depression separating mesepisternum from mesepimeron (Fig. 17). Metanotum with distinct lens-shaped dorsellum (only visible Fig. 9, a cleared, slide mount) and lateral panel length (toward hind wing articulation) about half length of dorsellum (Fig. 16). Metapleuron triangular, with a narrow anterior extension almost to base of hind wing, the margin at junction with mesopleuron deeply concave and posterior margin straight and vertical. Propodeum with a complete longitudinal groove medially, in lateral view weakly sloping, almost in same plane as dorsellum, in dorsal view with faint median longitudinal groove. Propodeal spiracle facing dorsolaterally in a slight depression and separated from anterior margin of propodeum by about its own diameter. Wings. Fore wing (Figs 4, 8, 19) wide, with completely transparent membrane sparsely scattered with minute microtrichia. Venation complete; submarginal vein with 1 subbasal seta and about 16 bullae, the most apical one at level of proximal macrochaeta (Fig. 8); parastigma, 0.5× submarginal vein length; marginal vein present, its length (from distal macrochaeta) about 2.0× length of parastigma; stigmal vein short but distinct, curving away from wing margin (Fig. 20) then closely paralleling it, with 4 apical campaniform sensilla (Fig. 21b); postmarginal vein present, about 0.8× as long and as thick as marginal vein, with 4 microchaetae along anterior margin; hypochaeta absent (Fig. 8) but one similar to it originating on parastigma (Fig. 20a); proximal campaniform sensillum near posterior margin of marginal vein just beyond apical macrochaeta (far removed from the 4 campaniform sensilla on stigmal vein). Hind wing (Fig. 8) with membrane not extending to base of wing, relatively wide medially, tapering gradually almost to a point apically. Fore wing membrane with one irregular row of microtrichiae basally and two or three distally. Legs. Profemur and metafemur narrow (Fig. 17), metafemur widest, about 2× as wide as mesofemur. Tarsi 5-segmented. Calcar (moveable protibial spur) apparently without setae along outer margin, and with the inner tine about 0.5× as long as outer tine. Middle and hind legs with tarsomere 1 shorter than tarsomere 2. Metasoma. About 2.2× as long as wide, 2.3× as long as high and 0.9× as wide as high; its length, excluding exserted part of ovipositor, about 2.0× that of mesosoma (Fig. 4). Petiole ring-like, about 0.3× as long as wide. Gastral terga apparently with fine longitudinal wrinkles except gt7; gt1 shorter than remaining terga and apparently gt6 the longest tergum, gt7 apically acutely triangular. Cercus a distinctly raised, longer than wide, peg-like plate, with 4 setae (Figs 22, 25). Gt6 with spiracle minute (apparently absent). Hypopygium extending posteriorly well beyond level of cercus (Figs 4, 23, 25). Ovipositor sheath exserted beyond gastral apex by about one third of its total length and abruptly upturned apically, apparently without subapical seta (Figs 22, 24).
Male. Unknown.
Hosts and habitat
Hosts are unknown. The holotype was collected in forest (Girault 1915[1]).
Taxon Treatment
- Huber, J; 2017: Eustochomorpha Girault, Neotriadomerus gen. n., and Proarescon gen. n. (Hymenoptera, Mymaridae), early extant lineages in evolution of the family Journal of Hymenoptera Research, (57): 1-87. doi
Images
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Other References
- ↑ Girault A (1915) Australian Hymenoptera Chalcidoidea — II. Second supplement. Memoirs of the Queensland Museum 3: 154–169.
