Epanerchodus pinguis
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Ordo: Polydesmida
Familia: Polydesmidae
Genus: Epanerchodus
Name
Epanerchodus pinguis Golovatch & Mikhaljova & Chang, 2011 sp. n. – Wikispecies link – ZooBank link – pinguis Pensoft Profile
Type material:
Holotype ♂ (head missing) (NMNS-6558-001), Taiwan, Nantou County, Huisun timber land, 15.04.2002, leg. S.H. Wu. Paratype ♂ (NMNS-6558-002), same locality, 24.03.1998, leg. S.H. Wu.
Name:
To emphasize the stout gonopod telopodite.
Diagnosis:
Differs from the other Epanerchodus species, in particular from the apparently especially similar Epanerchodus orientalis, in the mostly square, broader and slightly upturned paraterga, coupled with the gonopod showing an unusually densely setose coxa and a remarkably stout telopodite (see also Key below).
Description:
Length ca 16 (paratype) or 20 mm (holotype); width of midbody pro- and metazona 1.5 and 2.9 mm (paratype) or 1.5 and 3.0 mm (holotype), respectively. Coloration in alcohol uniformly light red-brown to red-brown; venter and legs yellowish (Figs 37–39).
All characters as in Epanerchodus orientalis except as follows.
Antennae rather long, slender, only slightly clavate, reaching behind segment 3 dorsally; antennomere 3 longest, clearly longer than highest 5th; antennomeres 5 and 6 each with an evident, compact, distodorsal group of bacilliform sensilla; antennomere 7 with a minute dorsoparabasal cone and a distodorsal group of microscopic sensilla.
In width, collum < head ≤ segment 2 ≤ 3 < 4 < = 5–16, thereafter body gradually tapering towards telson (Fig. 39). Paraterga strongly developed, starting from collum, slightly upturned, set high, mostly level to a very faintly convex dorsum, slightly below dorsum only on collum and segment 2; paraterga on collum small, subtriangular, a small lateral incision in front of a narrowly rounded caudal corner; front shoulders drawn forward only paraterga 2 and 3, straight and subrectangular on paraterga 4, onward straight but directed increasingly caudolaterad; caudal edge nearly straight on paraterga 2–7, thereafter caudal corner increasingly acutangular, nearly always narrowly rounded, only on segment 19 spiniform and nearly pointed, mostly lying nearly level to rear tergal contour until segment 15, onward extending increasingly beyond it (Figs 37–39). Paraterga 2 with 3–4 small lateral incisions, all following poreless segments with three, all pore-bearing ones with four, small but evident incisions, each usually bearing a small seta on top at lateral margin. Metatergal sculpture typical, rather obliterate, with three indistinct transverse rows of setiferous, polygonal bosses (Figs 37–39). Tergal setae very short, mostly retained, a little longer only on collum and in rear row on metatergum 19 (Figs 37, 39). Stricture between pro- and metazona wide and smooth. Limbus very thin, microdenticulate. Epiproct rather short, conical (Fig. 39), preapical papillae prominent. Hypoproct semi-circular; caudal, paramedian, setiferous papillae evident and well-separated.
Sterna without modifications, very densely setose (Figs 40, 41). Legs long and slender, evidently enlarged (Figs 42), ca 1.6–1.7 times as long as midbody height, prefemora not swollen dorsally, acropodite (femur+postfemur+tibia+tarsus) with sphaerotrichomes ventrally (Fig. 42).
Gonopods (Figs 40, 43, 44) with large, subquadrate, medially fused coxae carrying numerous long setae ventromedially. Telopodite mostly hidden inside gonocoel, unusually stout, subfalcate, prefemoral (densely setose) portion about half as long as entire telopodite; endomere (en) short, simple, with a subapical knob laterally and a rounded tip, as well as with two subunciform processes (s and p); hairy pulvillus very evident, exomere totally suppressed.
Remarks
This species is apparently very local in distribution (Map 2) and seems to be allopatric with Epanerchodus orientalis.
Original Description
- Golovatch, S; Mikhaljova, E; Chang, H; 2011: The millipede family Polydesmidae in Taiwan, with descriptions of five new species (Polydesmida, Diplopoda) ZooKeys, 93: 9-42. doi
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