Elthusa raynaudii

From Species-ID
Jump to: navigation, search
Notice: This page is derived from the original publication listed below, whose author(s) should always be credited. Further contributors may edit and improve the content of this page and, consequently, need to be credited as well (see page history). Any assessment of factual correctness requires a careful review of the original article as well as of subsequent contributions.

If you are uncertain whether your planned contribution is correct or not, we suggest that you use the associated discussion page instead of editing the page directly.

This page should be cited as follows (rationale):
van der Wal S, Smit N, Hadfield K (2019) Review of the fish parasitic genus Elthusa Schioedte & Meinert, 1884 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of three new species. ZooKeys 841 : 1–37, doi. Versioned wiki page: 2019-04-23, version 175869, https://species-id.net/w/index.php?title=Elthusa_raynaudii&oldid=175869 , contributors (alphabetical order): Pensoft Publishers.

Citation formats to copy and paste

BibTeX:

@article{van2019ZooKeys841,
author = {van der Wal, Serita AND Smit, Nico J. AND Hadfield, Kerry A.},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {Review of the fish parasitic genus Elthusa Schioedte & Meinert, 1884 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of three new species},
year = {2019},
volume = {841},
issue = {},
pages = {1--37},
doi = {10.3897/zookeys.841.32364},
url = {https://zookeys.pensoft.net/articles.php?id=32364},
note = {Versioned wiki page: 2019-04-23, version 175869, https://species-id.net/w/index.php?title=Elthusa_raynaudii&oldid=175869 , contributors (alphabetical order): Pensoft Publishers.}

}

RIS/ Endnote:

TY - JOUR
T1 - Review of the fish parasitic genus Elthusa Schioedte & Meinert, 1884 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of three new species
A1 - van der Wal S
A1 - Smit N
A1 - Hadfield K
Y1 - 2019
JF - ZooKeys
JA -
VL - 841
IS -
UR - http://dx.doi.org/10.3897/zookeys.841.32364
SP - 1
EP - 37
PB - Pensoft Publishers
M1 - Versioned wiki page: 2019-04-23, version 175869, https://species-id.net/w/index.php?title=Elthusa_raynaudii&oldid=175869 , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/zookeys.841.32364

Wikipedia/ Citizendium:

<ref name="van2019ZooKeys841">{{Citation
| author = van der Wal S, Smit N, Hadfield K
| title = Review of the fish parasitic genus Elthusa Schioedte & Meinert, 1884 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of three new species
| journal = ZooKeys
| year = 2019
| volume = 841
| issue =
| pages = 1--37
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.841.32364
| url = https://zookeys.pensoft.net/articles.php?id=32364
| pmc =
| accessdate = 2025-04-05

}} Versioned wiki page: 2019-04-23, version 175869, https://species-id.net/w/index.php?title=Elthusa_raynaudii&oldid=175869 , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Taxonavigation

Ordo: Isopoda
Familia: Cymothoidae
Genus: Elthusa

Name

Elthusa raynaudii (Milne Edwards, 1840) – Wikispecies link – Pensoft Profile

Type material

Type material held at the Museum national d’Histoire naturelle, Paris (syntypes MNHN-IU-2016-9885; MNHN-IU-2016-9884).

Type locality

Cape of Good Hope, South Africa.

Type host

Unknown.

Material examined

(all from South Africa). Syntype. SOUTH AFRICA • 1 ♀ (ovigerous, 26.7 mm TL, 14.1 mm W); south coast of South Africa, Cape of Good Hope; MNHN-IU-2016-9885. Other material. SOUTH AFRICA • 1 ♀ (ovigerous, 26.0 mm TL, 14.0 mm W); Indian Ocean, south coast of South Africa, RV Africana (fish sorting table); 34°38'S, 25°38'E; April 2003; coll. Nico J. Smit; dissected; in the collection of the authors at NWU • 1 ♀ (ovigerous, 26.0 mm TL, 15.0 mm W); Atlantic Ocean, RV Dr Fridtjof Nansen trawl (Station NAN401T062); January 2007; coll. L Atkinson; SAMC-A47881 • 1 ♀ (ovigerous, 20.0 mm TL, 12.0 mm W); Atlantic Ocean, RV Dr Fridtjof Nansen (fish sorting table); 32°17'S, 16°54'E; 269 m; February 2010; coll. KA Hadfield; dissected; SAMC-A089957.

Description

(ovigerous ♀). Figs 1–3. Body ovoid, slightly twisted to the left, 1.7 times as long as greatest width; dorsal surfaces smooth and polished in appearance, widest at pereonite 5, most narrow at pereonite 1; pereonite lateral margins mostly posteriorly ovate, medially indented. Cephalon 0.9 times longer than wide, visible in dorsal view, sub-truncate with blunt anterior margin. Frontal margin thickened, ventrally folded. Eyes oval with distinct margins; one eye 0.2 times width of cephalon, 0.4 times length of cephalon. Pereonite 1 smooth; anterior border medially straight, curved laterally; anterolateral angle narrowly rounded, extending to the medial region of eyes. Posterior margins of pereonites smooth, slightly curved laterally. Coxae 2–3 wide, with posteroventral angles rounded; coxae 4–7 with rounded point, not extending past pereonite posterior margin. Pereonites 2–5 subequal, becoming more progressively rounded posteriorly; pereonites 6 and 7 slightly narrower. Pleon 0.4 times as long as total body length, with pleonite 1 largely concealed by pereonite 7, slightly visible in dorsal view; pleonites posterior margin mostly concave. Pleonite 2 partially overlapped by pereonite 7. Pleonites 3–5 similar in form to pleonite 2; pleonites subequal in length, with posterolateral angles narrowly rounded, posterior margin straight. Pleotelson 0.6 times as long as anterior width, dorsal surface smooth; lateral margins weakly convex; posterior margin evenly rounded. Antennula shorter than antenna, consisting of eight articles; antennula peduncle articles I and II distinct and articulated, extending to anterior of pereonite 1. Antenna consists of eleven articles, extending to middle of pereonite 1.
Pereopod 1 basis 1.6 times as long as greatest width; ischium 0.7 times as long as basis; merus proximal margin without bulbous protrusion; carpus with rounded proximal margin; propodus 1.4 times as long as wide; dactylus slender, 1.6 times as long as propodus, 2.9 times as long as basal width. All pereopods without robust or simple setae. Pereopod 7 basis with carina, 2.5 times as long as greatest width; ischium without protrusions, 0.5 times as long as basis; merus 0.7 times as long as wide, 0.4 times as long as ischium; carpus without bulbous protrusion, 0.7 times as long as wide, 0.3 times as long as ischium; propodus 0.8 times as long as wide, 0.3 times as long as ischium; dactylus slender, 2.3 times as long as propodus, 3.5 times as long as basal width.
Pleopods simple, exopod larger than endopod. Pleopod 1 exopod 1.3 times as long as wide, lateral margin weakly convex, distally narrowly rounded, mesial margin straight; peduncle 2.3 times as wide as long.
Uropod more than half the length of pleotelson; peduncle 0.5 times longer than rami, lateral margin without setae; rami not extending beyond pleotelson, apices broadly rounded. Endopod apically rounded, 2.7 times as long as greatest width, terminating without setae. Exopod extending to end of endopod, 2.2 times as long as greatest width, apically rounded, terminating without setae.
Variations. Intra-specific variations can cause difficulty in identification and should be taken into consideration. One of the more obvious variations is the overall body shape of examined individuals, as seen from the dorsal view. While the syntype (MNHN–IU–2016–9885) has weakly convex, symmetrical lateral margins, specimen SAMC-A089957 is not as symmetrical, with the right margin being strongly convex and that of the left margin, weakly convex. The latter specimen therefore appears to be less symmetrical. Bruce (1990)[40] mentioned this occasional asymmetrical body shape as an observed variation, as a result of slightly twisted individuals. The body shape of the South African specimen (SAMC-A089957) accords to the shape of individuals illustrated and described by Bruce (1990)[40]. In addition, the widest part of this species may vary between pereonite 4 and pereonite 5. This variation may also cause individual body shapes to appear dissimilar. The anterior margin of the cephalon of the syntype (MNHN–IU–2016–9885) appears to be rounded rather than subtruncate. The posterior margin of pleonite 5 can be roughly straight (AM G2181 from Bruce 1990[40]), have a slight medial point, or be weakly concave (Bruce 1990[40], present study). Although Bruce (1990)[40] described the uropods as being short, most measure more than half the length of the pleotelson.
Size. Ovigerous females 20.0–26.7 mm TL, 14.0–15.0 mm W. Other material: ovigerous females 22.0–67.0 mm TL (average 30.83 mm TL) (Bruce 1990[40]).

Distribution

Records listed from west to east. North Pacific Ocean: Bering Sea (Kensley 1976[44]). South America: Punta Quillaipe (Menzies 1962[36]) and Chile (Nierstrasz 1931[29]); Uruguay (Mañé-Garzón 1979[39]). South Atlantic Ocean: Saint Helena and Tristan da Cunha (Sivertsen and Holthuis 1980[24]). South Africa: Table Bay (Barnard 1920[30]); Cape of Good Hope (Milne Edwards 1840[1]); Durban (Barnard 1955[21]). India: Travancore (Pillai 1954[31]). Southern Indian Ocean: Amsterdam Island (Kensley 1976[44]). Australia: southern and south-eastern Australia (Schioedte and Meinert 1884[4], Hale 1926[6], Bruce 1990[40], Whitelegge 1901[45]). Japan: Yokohama (Schioedte and Meinert 1884[4]). New Zealand (Filhol 1885[12], Chilton 1909[15], Nierstrasz 1915[28], Hurley 1961[22], Bruce 1990[40]).

Hosts

Elthusaraynaudii has been recorded from various fish hosts of multiple orders and families. These hosts are: Chelidonichthyskumu (Cuvier, 1829) (see Avdeev 1978[33]); Chorisochismusdentex (Pallas, 1769) (see Barnard 1920[30]); Cyttusaustralis (Richardson, 1843) (see Avdeev 1978[33], 1984[46], Bruce 1990[40]); Cyttusnovaezelandiae (Arthur, 1885) (see Avdeev 1978[33], 1984[46]); Cyttustraversi Hutton, 1872, previously Cyttoidopsmccullochi (Whitley, 1947) (see Avdeev 1984[46], Bruce 1990[40]); Genypterusblacodes (Bloch and Schneider, 1801) (see Hewitt and Hine 1972[23]); Gnathanacanthusgoetzeei Bleeker, 1855 (see Bruce 1990[40]); Hyporhamphusintermedius (Cantor, 1842) (see Powell 1959[47], Stephenson 1969[48]); Latrislineata (Forster, 1801) (see Kensley 1976[44]); Meuscheniafreycineti (Quoy and Gaimard, 1824) (see Bruce 1990[40]); Mustelusantarcticus Günther, 1870 (see Hewitt and Hine 1972[23]); Nemadactylusmonodactylus (Carmichael, 1819), previously Acantholatrismonodactylus (Carmichael, 1819) (see Sivertsen and Holthuis 1980[24]); Nematalosanasus (Bloch, 1795) (see Ghani 2003[43]); Notacanthussexspinis Richardson, 1846 (see Avdeev 1978[33], 1984[46]); Nototheniamicrolepidota Hutton, 1875, previously Nototheniacolbecki (see Chilton 1909[15], Hewitt and Hine 1972[23], Avdeev 1978[33], 1984[46]); Notolabrustetricus (Richardson, 1840), previously Pseudolabrustetricus (see Bruce 1990[40]); Paranototheniamagellanica (Forster, 1801), previously Nototheniamacrocephala (see Avdeev 1978[33]); Ilishamelastoma (Bloch and Schneider, 1801) previously Pellonabrachysoma (see Pillai 1954[31]); Pelotretisflavilatus Waite, 1911 (see Chilton 1911[16]); Pseudophycisbachus (Forster, 1801), previously Physiculusbachus (see Hewitt and Hine 1972[23]); Physiculus sp. (see Bruce 1990[40]); Pseudophycisbarbata Günther, 1863, previously Physiculusbarbatus (Günther, 1863) (see Bruce 1990[40]); Pseudolabrusmiles (Schneider and Forster, 1801) (see Poore 1981[35], Bruce 1990[40]); Pseudophycisbachus (Forster, 1801) (see Chilton 1911[16], Bruce 1990[40]); Rexeasolandri (Cuvier, 1832) (see Bruce 1990[40]); Rhombosolea sp. (see Hewitt and Hine 1972[23]); Sardinopssagax (Jenyns, 1842), previously Clupeaneopilchardus Steindachner, 1879 (see Chilton 1911[16]); Scorpaenacardinalis Solander and Richardson, 1842 (see Poore 1981[35]); Sebastescapensis (Gmelin, 1789), previously Sebastichthyscapensis (Gmelin, 1789) (see Sivertsen and Holthuis 1980[24]); Stolephoruscommersonnii Lacepède, 1803 (see Pillai 1954[31]); Thyrsitesatun (Euphrasen, 1791) (see Sivertsen and Holthuis 1980[24]); Zenopsisnebulosa (Temminck and Schlegel, 1845), previously Zenopsisnebulosus (see Bruce 1990[40]); Zeusfaber Linnaeus, 1758 (see Hale 1926[6], Avdeev 1984[46]). Unidentified by scientific names: banded perch (Serranidae), flathead (Platycephalidae) (see Bruce 1990[40]).

Remarks

Elthusaraynaudii can be distinguished by the cephalon having a narrowly truncate rostrum; pereonite 1 with anterior margin straight; pleonites subequal in shape and width; and broadly rounded uropod apices that extend to more than half the length of the pleotelson.
Originally described in 1840, from the Cape of Good Hope in South Africa, from an unknown host, Elthusaraynaudii has been recorded numerous times from a wide range of localities within the Indo-Pacific region. It is the only species of Elthusa that has been described from sub-Saharan Africa. It has been recorded from an unknown host from the Cape of Good Hope (see Milne Edwards 1840[1]); from the rocksucker, Chorisochismusdentex (Pallas, 1769) near Cape Town (Table Bay) (see Barnard 1920[30]); from a wrasse in Durban (see Barnard 1955[21]); as well as from the striped trumpeter, Latrislineata (Forster, 1801) (see Kensley 1976[44]).
Elthusasigani Bruce, 1990, which is only known from its type locality in Queensland, Australia, seems to be most similar to E.raynaudii. Elthusasigani can be distinguished from E.raynaudii by having an evenly concave pereonite 1 anterior margin; a flat, straight cephalon anterior margin; and coxae 7 that extend past the posterior margin of pereonite 7. In addition, E.sigani is a much smaller species in overall body length range (9.0–13.0 mm), compared to E.raynaudii (20.0–26.7 mm).

Table 1. Interspecific character states between Elthusaraynaudii (Milne Edwards, 1840), Elthusaxena sp. n., Elthusaacutinasa sp. n., and Elthusarotunda sp. n. from sub-Saharan African marine waters.
Morphological feature Elthusaraynaudii (Milne Edwards, 1840) Elthusaxena sp. n. Elthusaacutinasa sp. n. Elthusarotunda sp. n.
Body shape Ovoid Elongate ovoid Elongate ovoid Round
Shape of cephalon and anterior margin Sub-truncate, blunt anterior margin Sub-triangular, bluntly pointed anterior margin Sub-triangular, pointed anterior margin Sub-triangular, blunt anterior margin
Pereonite 1 anterior margin Straight Medially indented Medial projection Concave
Coxae 7 posterior margin Not extending past posterior margin of pereonite 7 Not extending past posterior margin of pereonite 7 Extending past posterior margin of pereonite 7 Not extending past posterior margin of pereonite 7
Pereopod 7 protrusions Absent Absent Absent Present on merus and carpus
Pleonite length Pleonites 1–5 sub-equal Pleonite 5 longest and indented Pleonite 1 longest Pleonite 5 longest, medially convex
Pleonite 1 width Narrower than other pleonites As long as other pleonites As wide as pleotelson Narrower than other pleonites
Pleonite 5 lateral margins Visible Visible Largely concealed by pleonite 4 Slightly concealed by pleonite 4
Pleotelson shape Evenly rounded Roughly quadrate and curved upwards Rounded Broadly rounded
Pleopod 5 endopod Slightly smaller than exopod Smaller than exopod (not dissected) Half the size of exopod Smaller than exopod (not dissected)
Uropods Broadly rounded, more than half the length of pleotelson Apices narrowly rounded, more than half the length of pleotelson Short, pointed, less than half the length of pleotelson Broadly rounded, half the length of pleotelson

Taxon Treatment

  • van der Wal, S; Smit, N; Hadfield, K; 2019: Review of the fish parasitic genus Elthusa Schioedte & Meinert, 1884 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of three new species ZooKeys, 841: 1-37. doi

Images

Figure 1. Elthusaraynaudii (Milne Edwards, 1840) ♀ (ovigerous, 20.0 mm TL, 12.0 mm W) (SAMC-A089957) from Dr Fridtjof Nansen research vessel A dorsal body B lateral body C oostegites D dorsal view of cephalon and pereonite 1 E uropod F ventral cephalon G pleopod 1 H dorsal view of pleon I pereopod 1 J pereopod 7.
Figure 1. Elthusaraynaudii (Milne Edwards, 1840) ♀ (ovigerous, 20.0 mm TL, 12.0 mm W) (SAMC-A089957) from Dr Fridtjof Nansen research vessel A dorsal body B lateral body C oostegites D dorsal view of cephalon and pereonite 1 E uropod F ventral cephalon G pleopod 1 H dorsal view of pleon I pereopod 1 J pereopod 7. 
Figure 2. Photos of Elthusaraynaudii (Milne Edwards, 1840) ♀ (ovigerous, 26.0 mm TL, 15.0 mm W) (SAMC-A47881) from Dr Fridtjof Nansen research vessel A dorsal view B ventral view C lateral view.
Figure 2. Photos of Elthusaraynaudii (Milne Edwards, 1840) ♀ (ovigerous, 26.0 mm TL, 15.0 mm W) (SAMC-A47881) from Dr Fridtjof Nansen research vessel A dorsal view B ventral view C lateral view. 
Figure 3. Photos of syntype material Livonecaraynaudii Milne Edwards, 1840 ♀ (ovigerous, 26.7 mm TL, 14.1 mm W) (MNHN–IU–2016–9885) A dorsal view B ventral view C lateral view.
Figure 3. Photos of syntype material Livonecaraynaudii Milne Edwards, 1840 ♀ (ovigerous, 26.7 mm TL, 14.1 mm W) (MNHN–IU–2016–9885) A dorsal view B ventral view C lateral view. 

Other References

  1. ↑ 1.0 1.1 1.2 Milne Edwards H (1840) Histoire Naturelle des Crustacés III, Comprenent l’anatomie, la physiologie et la classification de ces animaux, vol. 3.Librairie encyclopédique de Roret, Paris, 638 pp. https://doi.org/10.5962/bhl.title.6234
  2. ↑ Krauss F (1843) Die Siidafrikanischen Crustaceen. Eine Zusammenstellung aller bekannten Malacostraca, Bemerkungen iiber deren Lebensweise und geographische Verbreitung, nebst Beschreibung und Abbildung mehrer neuen Arten, Stuttgart, 1–68. https://doi.org/10.5962/bhl.title.4825
  3. ↑ Bleeker P (1857) Recherches sur les Crustacés de L’Inde Archipelagique. II. Sur les Isopodes Cymothoadiens de L’Archipel Indien.Natuurkundige vereeniging in Nederlandsche-Indie, Batavia, Verhandelingen2: 20–40. https://doi.org/10.5962/bhl.title.9908
  4. ↑ 4.0 4.1 4.2 Schioedte J, Meinert F (1884) Symbolæ ad monographium Cymothoarum crustaceorum isopodum familiæ. IV. Cymothoidæ Trib. II. Cymothoinæ. Trib. III: Livonecinæ.Naturhistorisk tidsskrift, Kjøbenhavn14: 221–454. https://doi.org/10.5962/bhl.title.10300
  5. ↑ Thielemann M (1910) Beitriige zur Naturgeschichte Ostasiens. Herausgegeben von F. Doflein. Band H, no. 9. Beitrage zu Kenntnis der isopodenfauna Ostasiens. Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse der K. Bayer. Akademia der Wissenschaften 2, 3: 1–109. https://doi.org/10.1002/ange.19390522011
  6. ↑ 6.0 6.1 6.2 6.3 Hale H (1926) Review of the Australian isopods of the cymothoid group. Part II.Transactions of the Royal Society of South Australia50: 201–234.
  7. ↑ White A (1847) List of the specimens of Crustacea in the collection of the British Museum.British Museum (Natural History), Department of Zoology, London, 143 pp. https://doi.org/10.5962/bhl.title.1708
  8. ↑ Miers E (1874) Descriptions of some new species of Crustacea, chiefly from New Zealand. Annals and Magazine of Natural History series 4, 17: 218–229. https://doi.org/10.1080/00222937608681934
  9. ↑ Miers E (1877) On a collection of Crustacea, Decapoda and Isopoda, chiefly from South America, with descriptions of new species.Proceedings of the Zoological Society (London)43: 653–679.
  10. ↑ Ellis I (1981) Some type specimens of Isopoda (Flabellifera) in the British Museum (Natural History) and the isopods in the Linnaean collection.Bulletin of the British Museum of Natural History (Zoology)40: 121–128. https://doi.org/10.5962/bhl.title.25418
  11. ↑ 11.0 11.1 Gerstaecker A (1882) Sechste Ordnung: Isopoda–Asseln.Bronn, HG, Klassen und Ordnungen des Thier-Reichs5: 8–278. https://doi.org/10.5962/bhl.title.2054
  12. ↑ 12.0 12.1 Filhol H (1885) Considerations relative it la faune des Crustaces de la Nouvelle Zelande.Bibliotheque de l’Ecole des Hautes it etudes, Section des Sciences naturelles30: 1–60. https://doi.org/10.1093/ww/9780199540884.013.u212849
  13. ↑ Thomson G, Chilton C (1886) Critical list of the CrustaceaMalacostraca of New Zealand.Transactions of the New Zealand Institute18: 141–159.
  14. ↑ Whitelegge T (1902) Crustacea. Part II. Isopoda. Part 1. In: Scientific Results of the Trawling Expedition of HMCS Thetis, Vol. 1. Memoirs of the Australian Museum4: 201–246.
  15. ↑ 15.0 15.1 15.2 Chilton C (1909) The Crustacea of the subantarctic Islands of New Zealand.Subantarctic Islands of New Zealand26: 601–671.
  16. ↑ 16.0 16.1 16.2 16.3 Chilton C (1911) Scientific Results of the New Zealand Government Trawling Expedition, 1907. Crustacea.Records of the Canterbury Museum1: 285–312. https://doi.org/10.5962/bhl.title.14022
  17. ↑ Chilton C (1912) Miscellaneous notes on some New Zealand Crustacea.Transactions of the New Zealand Institute44: 128–135.
  18. ↑ Stebbing T (1910) General catalogue of South African Crustacea Part V of SA Crustacea, for the Marine Investigations in South Africa.Annals of the South African Museum6: 281–599. https://doi.org/10.5962/bhl.part.15558
  19. ↑ Young M (1926) Marine biological notes no. 2. Fecundity of Livonecaraynaudii Milne-Edw. (synonym: Livonecanovae-zeelantjiae Miers).New Zealand Journal of Science and Technology8: 282–286.
  20. ↑ Barnard K (1940) Contributions to the crustacean fauna of South Africa, XII: Further additions to the Tanaidacea, Isopoda and Amphipoda, together with keys for the identification of the hitherto recorded marine and freshwater species.Annals of the South African Museum33: 381–543. https://doi.org/10.5962/bhl.part.22318
  21. ↑ 21.0 21.1 21.2 Barnard K (1955) Additions to the fauna-list of South African Crustacea and Pycnogonida.Annals of the South African Museum43(1): 1–107.
  22. ↑ 22.0 22.1 Hurley D (1961) A checklist and key to the Crustacean Isopoda of New Zealand and Subantarctic Is.Transactions of the Royal Society of New Zealand (Zoology)1: 239–292.
  23. ↑ 23.0 23.1 23.2 23.3 23.4 23.5 Hewitt G, Hine P (1972) Checklist of parasites of New Zealand fishes and of their hosts.New Zealand journal of marine and freshwater research6: 69–114. https://doi.org/10.1080/00288330.1977.9515410
  24. ↑ 24.0 24.1 24.2 24.3 24.4 Sivertsen E, Holthuis L (1980) The marine isopod Crustacea of the Tristan da Cunha Archipelago.Gunneria33: 1–128.
  25. ↑ Beumer J, Ashburner L, Burbury M, Jette E, Latham D (1982) A checklist of the parasites of fishes from Australia and its adjacent Antarctic territories. Technical Communication no. 48. St Albans: Commonwealth Institute of Parasitology.Commonwealth Agricultural Bureaux, Farnham Royal, Slough, 107 pp. https://doi.org/10.1017/s0022149x00028091
  26. ↑ Richardson H (1909) Isopods collected in the northwest Pacific by the U.S. bureau of fisheries steamer “Albatross” in 1906.Proceedings of the United States National Museum37: 75–129. https://doi.org/10.5479/si.00963801.37-1701.75
  27. ↑ Kussakin O (1979) Marine and brackish isopods (Isopoda) of cold and temperate waters of the northern hemisphere, vol. 1. Suborder Flabellifera.Opredeliteli po faune SSSR, izdavaemye Zoologicheskim institutom Akademii nauk SSSR122: 1–470.
  28. ↑ 28.0 28.1 Nierstrasz H (1915) Die Isopoden-Sammlung im Naturhistorischen Reichsmuseum zu Leiden – 1. Cymothoidae.Zoologische Mededelingen (Leiden)1: 71–108.
  29. ↑ 29.0 29.1 29.2 Nierstrasz H (1931) Isopoda genuina. II. Flabellifera. In: Weber M, De Beaufort LF (Eds) Die Isopoden der Siboga-Expedition Siboga Expeditie (Uitkomsten op Zoölogisch, Botanisch, Oceanographisch en Geologisch Gebied verzameld in de Oost-Indische 1899–1900 aan boord HM Siboga onder commando van Luitenant ter zee 1e kl GF Tydeman) EJ Brill, Leiden, 123–233. https://doi.org/10.5962/bhl.title.10641
  30. ↑ 30.0 30.1 30.2 30.3 Barnard K (1920) Contributions to the crustacean fauna of South Africa, 6: Further additions to the list of marine Isopoda.Annals of the South African Museum17: 319–438. https://doi.org/10.5962/bhl.part.22318
  31. ↑ 31.0 31.1 31.2 31.3 Pillai N (1954) A preliminary note on the Tanaidacea and Isopoda of Travancore.Bulletin of the Central Research Institute, University of Travancore, Trivandrum3: 1–22.
  32. ↑ Avdeev V (1975) Two representatives of parasitic isopods of the genus Lironeca (Cymothoidae) from the region of Australia and New Zealand.Parasitologia3: 247–251.
  33. ↑ 33.0 33.1 33.2 33.3 33.4 33.5 33.6 Avdeev V (1978) Notes on the distribution of the marine Cymothoidae (Isopoda, Crustacea) in the Australian–New Zealand region.Folia Parasitologica25: 281–283.
  34. ↑ Trilles J (1976) Les Cymothoidae (Isopoda, Flabellifera) des collections du Muséum National d’Histoire Naturelle de Paris. IV. Les Lironecinae Schioedte & Meinert, 1884. Bulletin du Muséum National d’Histoire Naturelle, 3e serie (Zoologie) 390: 773–800. https://doi.org/10.5962/bhl.title.58212
  35. ↑ 35.0 35.1 35.2 Poore G (1981) Marine Isopoda of the Snares Islands, New Zealand–1. Gnathiidea, Valvifera, Anthuridea, and Flabellifera.New Zealand Journal of Zoology8: 331–348. https://doi.org/10.1080/03014223.1981.10430613
  36. ↑ 36.0 36.1 Menzies R (1962) The zoogeography, ecology, and systematics of the Chilian marine isopods. The Lund University Chile Expedition, 1948–49, no. 42. Lunds Universitets Arskriffter, 2, Bund 57: 1–162.
  37. ↑ Kensley B (1978) Guide to the marine isopods of southern Africa. The Rustica Press (Pty.) Ltd., Trustees of the South African Museum, Wynberg, Cape Town, 173 pp.
  38. ↑ Moreira P, Sadowsky V (1978) An annotated bibliography of parasitic lsopoda (Crustacea) of Chondrichthyes.Boletim do Instituto Oceanográfico27: 95–152. https://doi.org/10.1590/s0373-55241978000200005
  39. ↑ 39.0 39.1 Mañé-Garzón F (1979) Una nueve especie del genero Lironeca Leach, 1818 (IsopodaCymothoidae) de la costa oceanica del Uruguay.Revista de Biologia del Uruguay12: 11–22.
  40. ↑ 40.00 40.01 40.02 40.03 40.04 40.05 40.06 40.07 40.08 40.09 40.10 40.11 40.12 40.13 40.14 40.15 40.16 40.17 40.18 40.19 40.20 Bruce N (1990) The genera Catoessa, Elthusa, Enispa, Ichthyoxenus, Idusa, Livoneca and Norileca n. gen. (Isopoda, Cymothoidae), crustacean parasites of marine fishes, with descriptions of eastern Australian species.Records of the Australian Museum42: 247–300. https://doi.org/10.3853/j.0067-1975.42.1990.118
  41. ↑ Bruce N, Lew Ton H, Poore G (2002) Cymothoidae Leach, 1814. In: Poore G (Ed.) Crustacea: Malacostraca: Syncarida and Peracarida: Isopoda, Tanaidacea, Mictacea, Thermosbaenacea, Spelaeogriphacea.CSIRO Publishing, Melbourne, 168–183. https://doi.org/10.1163/156854008x361012
  42. ↑ Williams E, Bunkley-Williams L, Ebert D (2010) An accidental attachment of Elthusaraynaudii (Isopoda, Cymothoidae) in Etmopterus sp. (Squaliformes, Etmopteridae).Acta Parasitologica55: 99–101. https://doi.org/10.2478/s11686-010-0006-6
  43. ↑ 43.0 43.1 Ghani N (2003) Isopod parasites of marine fishes of Pakistan.Proceedings of Pakistan Congress of Zoology23: 217–221.
  44. ↑ 44.0 44.1 44.2 44.3 Kensley B (1976) Isopodan and Tanaidacean Crustacea from St. Paul and Amsterdam Islands, southern Indian Ocean.Annals of the South African Museum69: 261–323.
  45. ↑ Whitelegge T (1901) Crustacea. Part II. ln: Scientific Results of the trawling Expedition of H.M.C.S. “Thelis” of the Coast of New South Wales in February and March, 1898.Australian Museum, Sydney4: 203–246. https://doi.org/10.3853/j.0067-1967.4.1901.471
  46. ↑ 46.0 46.1 46.2 46.3 46.4 46.5 Avdeev V (1984) Localization of isopods of the family Cymothoidae in the gill and oral cavities of fishes.Parasitology18: 23–29.
  47. ↑ Powell A (1959) Native Animals of New Zealand. Auckland Museum Handbook of Zoology.Unity Press, Auckland, 96 pp.
  48. ↑ Stephenson A (1969) Ironamelanosfiicta (Isopoda, Cymothoidae). A new record for New Zealand waters, with descriptions of male, female and larval states.Records of the Auckland Institute and Museum6: 427–34.
Retrieved from "https://species-id.net/w/index.php?title=Elthusa_raynaudii&oldid=175869"