Draconarius immensus

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Notice: This page is derived from the original publication listed below, whose author(s) should always be credited. Further contributors may edit and improve the content of this page and, consequently, need to be credited as well (see page history). Any assessment of factual correctness requires a careful review of the original article as well as of subsequent contributions.

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This page should be cited as follows (rationale):
Xin-Ping Wang, Charles E. Griswold, Jeremy A. Miller (2010) Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains. Zootaxa 2593 : -1 – -1, doi. Versioned wiki page: 2014-04-17, version 50912, https://species-id.net/w/index.php?title=Draconarius_immensus&oldid=50912 , contributors (alphabetical order): PlaziBot.

Citation formats to copy and paste

BibTeX:

@article{Xin-Ping2010Zootaxa2593,
author = {Xin-Ping Wang AND Charles E. Griswold AND Jeremy A. Miller},
journal = {Zootaxa},
title = {Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains},
year = {2010},
volume = {2593},
issue = {},
pages = {-1 -- -1},
doi = {TODO},
url = {http://www.mapress.com/zootaxa/2010/f/zt02593p127.pdf},
note = {Versioned wiki page: 2014-04-17, version 50912, https://species-id.net/w/index.php?title=Draconarius_immensus&oldid=50912 , contributors (alphabetical order): PlaziBot.}

}

RIS/ Endnote:

TY - JOUR
T1 - Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains
A1 - Xin-Ping Wang
A1 - Charles E. Griswold
A1 - Jeremy A. Miller
Y1 - 2010
JF - Zootaxa
JA -
VL - 2593
IS -
UR - http://dx.doi.org/TODO
SP - -1
EP - -1
PB -
M1 - Versioned wiki page: 2014-04-17, version 50912, https://species-id.net/w/index.php?title=Draconarius_immensus&oldid=50912 , contributors (alphabetical order): PlaziBot.

M3 - doi:TODO

Wikipedia/ Citizendium:

<ref name="Xin-Ping2010Zootaxa2593">{{Citation
| author = Xin-Ping Wang, Charles E. Griswold, Jeremy A. Miller
| title = Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains
| journal = Zootaxa
| year = 2010
| volume = 2593
| issue =
| pages = -1 -- -1
| pmid =
| publisher =
| doi = TODO
| url = http://www.mapress.com/zootaxa/2010/f/zt02593p127.pdf
| pmc =
| accessdate = 2020-10-22

}} Versioned wiki page: 2014-04-17, version 50912, https://species-id.net/w/index.php?title=Draconarius_immensus&oldid=50912 , contributors (alphabetical order): PlaziBot.</ref>

See also the citation download page at the journal.


Taxonavigation

Familia: Amaurobiidae
Genus: Draconarius

Name

Draconarius immensus Xin-Ping Wang, 2010Wikispecies linkPensoft Profile

  • Draconarius immensus Xin-Ping Wang, 2010, Zootaxa 2593: -1--1.

Description

(Fig. 540)

Materials Examined

China Jiulong County

Diagnosis

Diagnosis: This species is similar to D. incertus Wang 2003 and D. pseudobrunneus Wang 2003 in having a similar epigynum, a bifurcate conductor, and a simple median apophysis, but can be distinguished from D. incertus by the subdistally arising spermathecal heads, from D. pseudobrunneus by the different shapes of atrium in the female, and by the short median apophysis and the embolus with thread starting from posterior margin of embolic base in the male (Xu & Li 2006 b: figs 13 - 19; Wang & Jaeger 2010: figs 6 - 8).

Description

Description: See Xu & Li (2006 b) and Wang & Jaeger (2010).

Distribution

Distribution: China (Sichuan: Jiulong; Yunnan: Zhongdian) (Fig. 540).

Description

(Figs 189 - 195, 540)

Materials Examined

CHINA Bawan village G. Tang CHINA Yunnan China Bawan village G. Tang

Etymology

Etymology: The species name came from Latin " improprius ", which means " improper, unsuitable ", and refers to its provisional generic assignment; adjective.

Diagnosis

Diagnosis: The female of this new species can be easily distinguished from other Coelotinae by the absence of epigynal teeth, the anteriorly protruding anterior atrial margin, and the round spermathecae (Figs 189 - 190).

Description

Description: Female (holotype). Large sized Coelotinae, total length 11.8 (Fig. 193). Dorsal shield of prosoma 5.75 long, 3.80 wide; opisthosoma 6.00 long, 4.18 wide. AME smallest, approximately % the size of ALE; ALE largest; posterior eyes subequal in size, slightly smaller than ALE (AME 0.15, ALE 0.21, PME 0.18, PLE 0.18); anterior eyes equally separated from each other by 2 / s of AME diameter; PME separated from each other by 2 / s their diameter, widely separated from PLE by more than their diameter (AME-AME 0.11, AME-ALE 0.09, PME-PME 0.12, PME-PLE 0.22, AME-PME 0.15) (Fig. 194). Chelicera with 3 promarginal and 2 retromarginal teeth. Labium longer than wide (L / W = 1.14) (Fig. 195). Epigynum without epigynal teeth; atrium large, situated anteriorly, with anterior margin protruding anteriorly to an anteriorly narrow and posteriorly wide atrium; copulatory ducts small, originated anteriorly; spermathecal heads distinct, arising anteriorly, extending laterally; spermathecae large, round, widely separated by slightly less than their width, stalks slightly extending distally (Figs 189 - 192). Male. Unknown.

Distribution

Distribution: China (Yunnan: Tengchong) (Fig. 540).

Description

(Figs 196 - 231, 540)

Materials Examined

China Lushui County China Lushui County CHINA Pianma Township C. Griswold, D. Kavanaugh, K. Guo China Mingguang Township CM. Yin, J. F. Hu, X. H. Yang China Mingguang Township X. J. Peng, X. P. Wang, P. Hu China Cikai Township G. Tang China Cikai Township D. Kavanaugh China Cikai Township D. Kavanaugh, PE. Marek China Cikai Township D. Kavanaugh, P. E. Marek, D. Z. Dong, X. C. Liang China Cikai Township D. Kavanaugh, P. E. Marek China Dulongjiang Township V. F. Lee China Cikai Township H. M. Yan China Cikai Township H. M. Yan China Cikai Township D. Kavanaugh, P. E. Marek China Cikai Township D. Kavanaugh, P. E. Marek, D. Z. Dong, X. C. Liang China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Lishadi subdistrict G. X. Peng China Yaping subdistrict M. Yan, G. X. Peng China Gaoligongshan C. Griswold, D. Kavanaugh China Dali P. Jaeger China Cangshan P. Jaeger China Cangshan P. Jaeger China Cangshan P. Jaeger China Cangshan P. Jaeger

Diagnosis

Diagnosis: This species is similar to D. pseudobrunneus and D. latusincertus sp. nov. in having a similar epigynum and palp. The female can be distinguished by the distally arising spermathecal heads (Figs 196, 198 - 199, 206 - 229), and the male by the more or less small patellar apophysis and slightly convex posterior margin of embolus (Figs 202 - 204).

Description

Description: See Wang (2003). Photos of habitus, eyes, labium and genitalia are provided in this study (Figs 196 - 231).

Distribution

Distribution: China (Yunnan: Dali, Tengchong, Lushui, Fugong, Gongshan) (Fig. 540).

Description

(Fig. 540)

Materials Examined

China Lijiang

Diagnosis

Diagnosis: The female of this species can be easily distinguished from other coelotines by the absence of epigynal teeth, the large, anteriorly situated atrium and copulatory ducts, and the long, coiled spermathecal tubes (Xu & Li 2006 a: figs 18 - 19).

Description

Description: Female. See also Xu & Li (2006 a). Male. Unknown.

Distribution

Distribution: China (Yunnan: Lijiang) (Fig. 540).

Description

(Figs 232 - 239, 540)

Materials Examined

CHINA Jietou Township D. Kavanaugh

Etymology

Etymology: The specific name is a patronym in honor of David Kavanaugh, curator at CAS and the collector of the type specimen; noun in genitive case.

Diagnosis

Diagnosis: The male of this new species is similar to D. agrestis Wang 2003 and D. pseudoagrestis sp. nov. in having a similar palp, but can be distinguished by the large lateral tibial apophysis and the distinct posterior extension of the conductor (Figs 232 - 233).

Description

Description: Male (holotype). Medium sized Coelotinae, total length 7.17 (Fig. 237). Dorsal shield of prosoma 3.73 long, 2.52 wide; opisthosoma 3.42 long, 2.27 wide. AME smallest, 2 / 3 the size of ALE; lateral eyes subequal or with PLE slightly larger; PME slightly smaller than lateral eyes (AME 0.10, ALE 0.16, PME 0.14, PLE 0.17); anterior eyes equally separated by slightly more than half of their diameter; PME separated from each other by slightly less than half of their diameter, from PLE by PLE diameter (AME-AME 0.06, AME-ALE 0.07, PME-PME 0.06, PME-PLE 0.16, AME-PME 0.09) (Fig. 238). Chelicera with 3 promarginal and 2 retromarginal teeth. Labium longer than wide (L / W = 1.12) (Fig. 239). Palp with a small patellar apophysis; RTA more than half of tibial length, with a blunt distal end; lateral tibial apophysis distinctly large; cymbial furrow extending more than half of cymbial length; conductor long, slightly coiled, with a well developed basal lamella and a small dorsal apophysis; median apophysis spoon-shaped, not free-standing along anterior edge; embolus long, filiform, proximal in origin, arising at 6 - o'clock-position, running half an oval, extending posteriorly to middle part of tibia, and coiling anteriorly beyond distal part of bulb (Figs 232 - 236). Female. Unknown.

Distribution

Distribution: China (Yunnan: Tengchong) (Fig. 540).

Description

(Fig. 541)

Materials Examined

China Haoming County

Diagnosis

Diagnosis: The female of this species can be easily identified by the posteriorly situated epigynal hoods, the distinctly posteriorly protruding anterior atrial margin, the presence of small epigynal teeth, and the coiled copulatory ducts (with 3 or more loops) (Liu & Li 2009: figs 7 - 8).

Description

Description: See Liu & Li (2009).

Distribution

Distribution: China (Yunnan: Haoming) (Fig. 541).

Description

(Figs 240 - 248, 541)

Materials Examined

CHINA Lishadi Township G. Tang CHINA Lishadi subdistrict H. M. Yan, G. X. Peng China Gaoligongshan C. Griswold, D. Kavanaugh

Etymology

Etymology: The species name is derived from the Latin words " latus " and " cavus ", meaning " broad, wide " and " cavity " separately, and refers to the large, distinct atrial opening; term in apposition.

Diagnosis

Diagnosis: The female of this new species is similar to D. tibetensis Wang 2003 in having a similar atrium and similar spermathecal tubes, but the epigynal teeth of this new species arise at level anterior to epigynal hoods and distinctly separated from anterior atrial margin (Figs 240 - 241), while in D. tibetensis the epigynal teeth arise at level posterior to epigynal hoods and close to anterior atrial margin.

Description

Description: Female (holotype). Large sized Coelotinae, total length 12.3 (Fig. 246). Dorsal shield of prosoma 6.16 long, 4.24 wide; opisthosoma 6.10 long, 3.92 wide. Eyes subequal in size, with PME slightly smaller (AME 0.22, ALE 0.21, PME 0.18, PLE 0.22); anterior eyes equally separated by half of their size; PME separated from each other by about their diameter, widely separated from PLE by 1.5 times PME diameter (AME-AME 0.10, AME-ALE 0.09, PME-PME 0.20, PME-PLE 0.30, AME-PME 0.25) (Fig. 247). Labium slightly longer than wide (L / W = 1.09) (Fig. 248). Chelicera with 3 promarginal and 2 retromarginal teeth. Epigynal teeth short, arising anteriorly at level anterior to epigynal hoods, distinctly separated from anterior atrial margin; atrium large, but shallow, with distinct, continuous anterior margin, but indistinct lateral margins; atrial ridges indistinct; copulatory ducts small, originating and extending medially between sper-mathecae; spermathecae with bases widely separated by at least their width, stalks extending anteriorly beyond their heads; spermathecal heads distinct, arising from middle part of spermathecae (Figs 240 - 246). Male. Unknown.

Distribution

Distribution: China (Yunnan: Fugong) (Fig. 541).

Description

(Figs 249 - 272, 542)

Materials Examined

CHINA Gaoligongshan C. Griswold, D. Kavanaugh CHINA Lishadi subdistrict H. M. Yan, G. X. Peng China Gaoligongshan C. Griswold, D. Kavanaugh China Lishadi subdistrict H. M. Yan China Lishadi subdistrict H. M. Yan, G. X. Peng China Gaoligongshan C. Griswold, D. Kavanaugh China Lishadi D. Z. Dong China Gaoligongshan C. Griswold, D. Kavanaugh CHINA Diancang Shan D. W. Wrase

Etymology

Etymology: The species name is derived from the Latin word " latus ", which means " broad ", and refers to the broad conductor compared to other Draconarius incertus group species; term in apposition.

Diagnosis

Diagnosis: D. latusincertus is similar to D. pseudobrunneus Wang 2003 and D. incertus Wang 2003 in having a similar epigynum and similar palp, but can be distinguished by the distinctly larger body size, the broad conductor in the male; and the large atrium with atrial ridges extending posteriorly close to the epigastric furrow and the distinct anterior extension of the spermathecae in the female (Figs 249 - 252).

Discussion

Notes: The female from Dali, Diancang Shan (SMF) has a similar epigynum as D. latusincertus sp. nov., but its atrium is wider than long and the spermathecae are much shorter (Figs 263 - 264). This specimen is temporarily listed here as D. latusincertus sp. nov. Further collection of both males and females from this location is badly needed.

Description

Description: Male (holotype). Large sized Coelotinae, total length 11.11 (Fig. 267). Dorsal shield of prosoma 6.30 long, 4.33 wide; opisthosoma 4.81 long, 3.40 wide. AME smallest, % the size of ALE, which are the largest; posterior eyes subequal in size, slightly smaller than ALE (AME 0.17, ALE 0.23, PME 0.19, PLE 0.19); AME separated from each other by their diameter, from ALE by half the size of AME; PME separated from each other by slightly less than their diameter, from PLE by more than their diameter; AME and PME separated by AME diameter (AME-AME 0.16, AME-ALE 0.08, PME-PME 0.17, PME-PLE 0.23, AME-PME 0.16) (Fig. 268). Labium subequal in length and width (L / W = 1.09) (Fig. 269). Chelicerae with 3 promarginal and 2 retromarginal teeth. Palp with a short patellar apophysis; RTA more than half of tibial length, with slightly protruding distal end; lateral tibial apophysis distinct, widely separated from RTA; a short macroseta arising near lateral tibial apophysis; cymbial furrow less than half of cymbial length; conductor short, slightly bifurcate, broad, with a slender dorsal apophysis and a small basal lamella; median apophysis simple, not spoon-shaped; embolus short, filiform, prolateral in origin (Figs 251 - 252, 255 - 258). Female (paratype of CASENT 9011586 measured). Large sized Coelotinae, total length 12.15 (Fig. 270). Dorsal shield of prosoma 6.00 long, 4.05 wide; opisthosoma 6.15 long, 4.60 wide. AME smallest, % the size of ALE, which are the largest; PLE slightly smaller than ALE, PME slightly smaller than PLE (AME 0.18, ALE 0.24, PME 0.20, PLE 0.22); AME separated from each other by their diameter, from PLE by % of AME diameter; PME separated from each other by their diameter, from PLE by more than PLE size; AME and PME separated by AME diameter (AME-AME 0.18, AME-ALE 0.12, PME-PME 0.20, PME-PLE 0.25, AME-PME 0.18) (Fig. 271). Labium slightly longer than wide (L / W = 1.11) (Fig. 272). Promargin of chelicera with 3, retromargin with 2 teeth. Epigynal teeth absent; atrium large, with distinct, posteriorly protruding anterior margin and indistinct lateral margins; atrial ridges distinct, situated on posterior half of atrium, slightly extending posteriorly, but distinctly separated from epigastric furrow; copulatory ducts short, originating and extending between spermathecae; spermathecae broad, slightly elongated, widely separated, anteriorly extending beyond heads; spermathecal heads arising from distal half of spermathecae (Figs 249 - 250, 253254, 259 - 266).

Distribution

Distribution: China (Yunnan: Dali, Fugong) (Fig. 542).

Description

(Figs 273 - 287, 542)

Materials Examined

CHINA Gaoligongshan C. Griswold, H. M. Yan CHINA Yunnan China Dulongjiang Township J. A. Miller, D. Kavanaugh China Dulongjiang Township J. A. Miller, J. Wang China Dulongjiang Township J. A. Miller, D. Kavanaugh

Etymology

Etymology: The specific name is in honour of Gershom Levy (1927 - 2009), who helped the first author on the Near East Coelotinae spiders; noun in genitive case.

Diagnosis

Diagnosis: The male of this new species can be easily recognized by the large patellar apophysis, the slender RTA that does not reach the distal tibia, and the long, elongated, prolaterally originating base of the embolus, and the female by the absence of epigynal teeth, the distinct atrium, the large, round spermathecae, and the long, ventrally extending spermathecal tubes (Figs 273 - 276).

Description

Description: Female (holotype). Medium sized Coelotinae, total length 5.85 (Fig. 282). Dorsal shield of prosoma 3.10 long, 1.96 wide; opisthosoma 2.75 long, 1.95 wide. AME smallest, less than half the size of ALE; ALE largest; posterior eyes subequal, slightly smaller than ALE (AME 0.06, ALE 0.14, PME 0.12, PLE 0.13); AME separated from each other by slightly more than their diameter, from ALE by slightly less than AME diameter; PME separated from each other and from PLE by 2 / s of AME diameter (AME-AME 0.08, AME-ALE 0.03, PME-PME 0.08, PME-PLE 0.10, AME-PME 0.10) (Fig. 283). Labium with length and width subequal (Fig. 284). Promargin of chelicera with 3, retromargin with 2 teeth. Epigynal teeth absent; atrium distinct, large, length and width subequal, with indistinct anterior margin and distinct, parallel extending lateral margins; copulatory ducts indistinct from dorsal view; spermathecae large, round, slightly separated, with spermathecal tubes originating posteriorly and extending anteriorly on ventral side of large spermathecal balls, converging and close together anteriorly; spermathecal head distinct, situated distally, extending laterally (Figs 273 - 274, 277 - 278). Male (paratype). Medium sized Coelotinae, total length 5.55 (Fig. 285). Dorsal shield of prosoma 2.80 long, 2.30 wide; opisthosoma 2.75 long, 1.70 wide. AME smallest, half the size of lateral eyes; lateral eyes subequal in size, largest; PME slightly smaller than lateral eyes (AME 0.07, ALE 0.13, PME 0.11, PLE 0.13); AME separated from each other by about their diameter, from ALE by half of AME diameter; PME separated from each other by 2 / s of their diameter, from PLE by PME diameter (AME-AME 0.06, AME-ALE 0.03, AME-PME 0.10, PME-PME 0.08, PME-PLE 0.10) (Fig. 286). Labium with equal length and width (L / W = 1.0) (Fig. 287). Chelicerae with 3 promarginal and 2 retromarginal teeth. Palp with a large patellar apophysis; RTA more than half of tibial length, but relatively small and slender, with distal end not protruding beyond distal tibia; lateral tibial apophysis short, broad; cymbial furrow less than half of cymbial length; conductor long, slender, with a small lamella and a broad dorsal apophysis; median apophysis spoon-shaped, elongate, free-standing along anterior edge; embolus short, filiform, prolateral in origin, with elongated base (Figs 275 - 276, 279 - 281).

Distribution

Distribution: China (Yunnan: Fugong, Gongshan) (Fig. 542).

Description

(Fig. 541)

Materials Examined

China Kunming

Diagnosis

Diagnosis: The male is similar to D. paraepisomos Wang & Martens 2009 in having a short conductor and a proximally originating embolus, but can be distinguished by the relatively broad patellar apophysis, the lateral tibial apophysis that is broad and situated close to RTA, and the cymbial furrow that slightly longer than half of cymbial length (Liu & Li 2009: figs 22 - 24). The female is similar to D. yani sp. nov. in having a similar epigynum, but the atrium of D. lini is round, with indistinct anterior margin (Liu & Li 2009: fig. 25).

Description

Description: See Liu & Li (2009).

Distribution

Distribution: China (Yunnan: Yiliang, Kunming) (Fig. 541).

Description

(Figs 288 - 295, 542)

Materials Examined

CHINA Bawan village G. Tang CHINA Xiaoheishan village (National 320 Road) G. Tang China Bawan village G. Tang

Etymology

Etymology: The specific name is derived from one of the type localities, Longling County, Yunnan, China; adjective.

Diagnosis

Diagnosis: Males of this new species can be easily recognized by the long, broad patellar apophysis, the bifurcate conductor, the broad, round, simple median apophysis, and the prolaterally originating embolus (Figs 288 - 289).

Description

Description: Male (holotype). Medium sized Coelotinae, total length 9.01 (Fig. 293). Dorsal shield of prosoma 5.01 long, 3.36 wide; opisthosoma 4.00 long, 2.50 wide. AME smallest, ALE largest; PME and PLE subequal in size, slightly smaller than ALE (AME 0.14, ALE 0.19, PME 0.16, PLE 0.17); AME separated from each other and from ALE by about half of AME diameter; PME separated from each other by 2 / s of their diameter, widely separated from PLE by about 1.5 times PME diameter (AME-AME 0.07, AME-ALE 0.08, PME-PME 0.10, PME-PLE 0.22, AME-PME 0.15) (Fig. 294). Labium longer than wide (L / W = 1.11) (Fig. 295). Promargin of chelicera with 3 teeth, retromargin 2. Palpal patellar apophysis large, with broad base and slender distal end; RTA more than half of tibial length, with blunt distal end; lateral tibial apophysis distinctly separated from RTA; cymbial furrow less than half of cymbial length; conductor short, slightly bifurcate, with a dorsal apophysis and a small basal lamella; median apophysis broad, round, simple (not spoon-shaped); embolus short, filiform, prolateral in origin (Figs 288 - 292). Female. Unknown.

Distribution

Distribution: China (Yunnan: Tengchong, Longling) (Fig. 542).

Description

(Figs 296 - 302, 542)

Materials Examined

CHINA Cikai township D. Kavanaugh

Etymology

Etymology: The species name is derived from the Greek words " mikro " and " ommatos ", meaning " small " and " eye " separately, and refers to the relatively small eyes; term in apposition.

Diagnosis

Diagnosis: The female is similar to D. episomos Wang 2003 in having a small, posteriorly situated atrium, anteriorly broad, round spermathecae, and medially arising spermathecal heads, but can be distinguished by the broader atrium and the distinctly separated speramthecae (Figs 296 - 299).

Description

Description: Female (holotype). Large sized Coelotinae, total length likely more than 10.00 (Fig. 300). Dorsal shield of prosoma 5.00 long, 3.48 wide; abdomen damaged. Eyes small and widely separated. Median eyes subequal in size, small; ALE largest, 1.5 times median eyes in size; PLE slightly larger than median eyes (AME 0.09, ALE 0.15, PME 0.08, PLE 0.11); AME separated from each other by more than AME diameter, widely separated from ALE by more than 2 times AME diameter; PME separated from each other by about 2 - 3 times their diameter, from PLE by 4 times PME diameter (AME-AME 0.12, AME-ALE 0.19, AME-PME 0.16, PME-PME 0.22, PME-PLE 0.31) (Fig. 301). Labium longer than wide (L / W = 1.29) (Fig. 302). Chelic-era with 3 promarginal and 2 retromarginal teeth. Epigynal teeth short; atrium small, wider than long, close to epigastric furrow, with distinct atrial ridges; copulatory ducts small, originating medially and posteriorly; spermathecae large, round, bases distinctly separated by about their width, anteriorly converging and slightly separated by less than half of their width; spermathecal heads distinct, situated distally from inner margins of spermathecae (Figs 296 - 299). Male. Unknown.

Distribution

Distribution: China (Yunnan: Gongshan) (Fig. 542).

Description

(Fig. 543)

Materials Examined

China Baoxing County Sichuan Wenchuan County China Weixi County

Diagnosis

Diagnosis: The male is similar to those of D. pseudobrunneus Wang 2003 in having similar epigynum and palp. In D. mupingensis, the dorsal branch of the bifurcate conductor is about the same length as the ventral one (distinctly longer in D. pseudobrunneus) and the embolus thread starts from the anterior margin of the embolic base (from between anterior and posterior margins in D. pseudobrunneus). The female can only be distinguished by the slightly different spermathecae (Xu & Li 2006 b: figs 20 - 26).

Description

Description: See Xu & Li 2006.

Distribution

Distribution: China (Sichuan: Baoxing, Wenchuan; Yunnan: Weixi) (Fig. 543).

Description

(Fig. 543)

Materials Examined

Coelotes noctulus Wang et al. 1990: 226, figs 113 - 114 (female holotype, in HNU, examined). - Song et al. 1999: 377, figs 220 M-N. Wang & Jaeger 2007: 31, figs 33 - 35.

Diagnosis

Diagnosis: The female of this species can be easily recognized by the absence of epigynal teeth, the large, deep, anteriorly situated atrium, and the broad, posteriorly extending spermathecae (Wang & Jaeger 2007: figs 33 - 34). Chelicerae with 4 - 5 promarginal and 5 - 6 retromarginal teeth (Wang & Jaeger 2007: fig. 35).

Description

Description: Female. See Wang & Jaeger (2007). Male. Unknown.

Distribution

Distribution: China (Yunnan: Jinhong) (Fig. 543).

Description

(Figs 303 - 306, 543)

Materials Examined

China Tengchong County

Diagnosis

Diagnosis: The male is similar to D. catillus sp. nov. in having a short conductor, a proximally originating embolus, and the absence of a patellar apophysis, but can be distinguished by the short cymbial furrow, the slightly spoon-shaped median apophysis, and the short embolus (Figs 303 - 305).

Description

Description: Male. See Wang (2003). Photos of habitus and palp are provided in this study. Female. Unknown.

Distribution

Distribution: China (Yunnan: Tengchong) (Fig. 543).

Description

(Figs 307 - 315, 543)

Materials Examined

CHINA Mingguang Township X. J. Peng, X. P. Wang, P. Hu

Etymology

Etymology: The species name came from Latin word " olor ", meaning " swan ", and refers to the swanshaped vulva; adjective.

Diagnosis

Diagnosis: The female of this new species can be easily recognized by the absence of epigynal teeth, the distinctly separated atria, and the long, posteriorly originating, anteriorly diverging copulatory ducts (Figs 307 - 310).

Description

Description: Female (holotype). Medium sized Coelotinae, total length 5.93 (Fig. 311). Dorsal shield of prosoma 2.69 long, 2.45 wide; opisthosoma 3.24 long, 2.26 wide. AME smallest, 2 / s size of other eyes, which are subequal (AME 0.08, ALE 0.13, PME 0.14, PLE 0.13); anterior eyes equally separated by half of AME diameter; PME close together, separated from each other by of their diameter, from PLE by of PME diameter; PME separated from each other by of their diameter, from PLE by about of PME diameter (AME-AME 0.04, AME-ALE 0.04, PME-PME 0.04, PME-PLE 0.09, AME-PME 0.08) (Fig. 312). Labium length and width subequal (Fig. 313). Cheliceral pro- and retromargins with 2 teeth. Epigynum without epigy-nal teeth; atria distinctly separated by a broad septum, which is as broad as atrial width; copulatory ducts large, originating medially and posteriorly, anteriorly extending and diverging; spermathecal bases large, round, widely separated by about 1.5 times their width; spermathecal stalks narrow, extending anteriorly; spermathecal heads distinct, arising distally (Figs 307 - 310). Male. Unknown.

Biology and Ecology

Notes: The specimen was collected from a small funnel web of a road cut on the mountain top at a border station between China and Myanmar (Figs 314 - 315).

Distribution

Distribution: China (Yunnan: Tengchong) (Fig. 543).

Description

(Figs 316 - 321, 543)

Materials Examined

China Kunming

Diagnosis

Diagnosis: Similar to D. terebratus (Peng & Wang 1997) by the absence of epigynal teeth, presence of posteriorly situated epigynal hoods, anteriorly extending copulatory ducts, and long spermathecal heads. The female can be distinguished by the slightly elongated atrium, and the male by the broad, distinctly bifurcate conductor and the broad embolus (Figs 316 - 318).

Description

Description: See Wang et al. (1990) and Wang (2003). Photos of male habitus, eyes, labium and palp are provided in this study.

Distribution

Distribution: China (Yunnan: Kunming) (Fig. 543).

Description

(Fig. 544)

Materials Examined

China Xiangcheng China Xiangcheng County

Diagnosis

Diagnosis: Similar to D. latusincertus sp. nov. in having a similar epigynum, but can be distinguished by the short anterior extensions of the spermathecae (Xu & Li 2006 b: figs 28 - 30).

Description

Description: Female. See Xu & Li (2006 b). Male. Unknown.

Distribution

Distribution: China (Sichuan: Xiangcheng, Yajiang, Litang; Yunnan: Zhongdian; Tibet: Zayu) (Fig. 544).

Description

(Figs 322 - 325, 544)

Materials Examined

CHINA Gaoligongshan C. Griswold, D. Kavanaugh, C. L. Long CHINA Yunnan

Etymology

Etymology: The specific name refers to its similarity to D. spiralis sp. nov.; term in apposition. Diagnosis: This new species is similar to D. spiralis sp. nov. and D. pseudospiralis sp. nov. in having a large atrium, absence of epigynal teeth, coiled spermathecal tubes, distally coiled conductor, and simple median apophysis, but can be distinguished by the absence of patellar apophysis, the small conductor, and the prolaterally originating embolus in the male, and by the presence of diverticula that arise from spermathecal bases in the female (Figs 322 - 325).

Description

Description: Female (holotype). Medium sized Coelotinae, total length 7.20. Dorsal shield of prosoma 3.80 long, 2.60 wide; opisthosoma 3.40 long, 2.40 wide. AME smallest, approximately half the size of ALE; ALE largest; posterior eyes subequal in size, slightly smaller than ALE (AME 0.10, ALE 0.22, PME 0.20, PLE 0.20); AME separated from each other by their diameter, from ALE by slightly less than AME diameter; PME separated from each other by half of their diameter, from PLE by % of PME diameter (AME-AME 0.10, AME-ALE 0.07, PME-PME 0.10, PME-PLE 0.16, AME-PME 0.15). Promargin of chelicera with 3 and ret-romargin with 2 teeth. Epigynal teeth absent; atrium large, but shallow, wider than long, with anterior margin distinctly extending posteriorly to a short septum, which occupies less than half of atrial length; copulatory ducts short, originating anteriorly; spermathecal bases widely separated by about 2 times their width, with long, anteriorly extending diverticula; spermathecal stalks long, anterior and median extending with two coils; spermathecal heads small, but distinct (Figs 322 - 323). Male (paratype measured). Medium sized Coelotinae, total length 8.62. Dorsal shield of prosoma 4.42 long, 3.12 wide; opisthosoma 4.20 long, 3.14 wide. AME smallest, approximately % the size of other eyes, which are subequal (AME 0.15, ALE 0.21, PME 0.21, PLE 0.20); AME separated from each other by 2 / s of their diameter, from ALE by of AME diameter; PME separated from each other by half of their diameter, from PLE by slightly less than their diameter (AME-AME 0.11, AME-ALE 0.05, PME-PME 0.10, PME-PLE 0.18, AME-PME 0.18). Promargin of chelicera with 3 and retromargin with 2 teeth. Palp without patellar apophysis; RTA approximately half of tibial length or slightly longer, with distinctly protruding distal end; lateral tibial apophysis small; cymbial furrow short, slightly less than half of cymbial length; conductor short, with a slightly coiling apex, a slender dorsal apophysis, and a small basal lamella; median apophysis simple, not spoon-shaped; embolus filiform, short, prolateral in origin (Figs 324 - 325).

Distribution

Distribution: China (Yunnan: Lushui) (Fig. 544).

Description

(Figs 326 - 330, 545)

Materials Examined

China Tengchong County

Diagnosis

Diagnosis: The female of this species can be easily recognized by the large atrium, the posteriorly situated epigynal teeth, and the presence of long diverticula on the spermathecal bases (Figs 326 - 327).

Description

Description: Female. See Wang (2003). Photos of habitus, eyes, labium and epigynum are provided in this study (Figs 326 - 330). Male. Unknown.

Distribution

Distribution: China (Yunnan: Tengchong) (Fig. 545).

Description

(Figs 331 - 351, 545)

Materials Examined

China Gongshan County CHINA Bingzhongluo Township Hu Peng China Bingzhongluo Township Hu Peng China Bingzhongluo Township J. A. Miller, D. Kavanaugh China Bingzhongluo Township J. A. Miller, D. Kavanaugh China Cikai Township G. Tang China Cikai Township D. Kavanaugh China Dulongjiang Township G. Tang China Bingzhongluo Township J. A. Miller, D. Kavanaugh China Bingzhongluo Township J. A. Miller, D. Kavanaugh China Bingzhongluo Township Hu Peng China Bingzhongluo Township J. A. Miller, D. Kavanaugh China Lumadeng Township, S. of Shibali Yakou D. Kavanaugh China Bingzhongluo Township P. Hu China Cikai Township H. M. Yan China Cikai Township D. Kavanaugh, P. E. Marek China Cikai Township D. Kavanaugh, P. E. Marek, D. Z. Dong, X. C. Liang China Cikai Township D. Kavanaugh, P. E. Marek China Cikai Township H. M. Yan China Cikai Township H. M. Yan China Cikai Township D. Kavanaugh China Cikai Township D. Kavanaugh, P. E. Marek, D. Z. Dong China Cikai Township D. Kavanaugh China Lishadi P. Paquin China Pihe Township G. Tang China Lishadi P. Paquin China Lishadi Township G. Tang China Pianma Township K. J. Guo China Feng Xue Yakou C. Griswold China Lushui County D. Kavanaugh, P. E. Marek, H. B. Liang

Diagnosis

Diagnosis: The female of this species is similar to D. dubius Wang 2003 by lacking epigynal teeth and having broad spermathecae, but can be distinguished by the presence of distinct anteriorly diverging, posteriorly converging atrial ridges and the broad spermathecae (Figs 332, 337 - 351). The male can be easily recognized by the broad, bifurcate patellar apophysis, the simple median apophysis, and the short, broad, prolaterally originating embolus (Figs 334 - 336).

Description

Description: See Wang (2003). Photos of habitus, eyes, labium and genitalia are provided in this study.

Distribution

Distribution: China (Yunnan: Lushui, Fugong, Gongshan) (Fig. 545).

Description

(Fig. 545)

Materials Examined

China Kunming

Diagnosis

Diagnosis: The female can be easily recognized by the anteriorly situated copulatory ducts and the laterally extending spermathecae, and the male by the long RTA, the round, spoon-shaped median apophysis, and the proximally originated embolus (Wang 2003: figs 50 A-E).

Description

Description: See Wang et al. (1990) and Wang (2003).

Distribution

Distribution: China (Yunnan: Kunming) (Fig. 545).

Description

(Figs 352 - 360, 547)

Materials Examined

CHINA Bawan village G. Tang CHINA Bawan village G. Tang China Feng Xue Yakou C. Griswold, D. Kavanaugh, K. J. Guo

Etymology

Etymology: The species name is derived from its similarity to D. agrestis; term in apposition.

Diagnosis

Diagnosis: This species seems to be similar to D. agrestis Wang 2003 differing only in minor features in male palp: the long conductor that extends posteriorly near the base of embolus from ventral view, the presence of denticles on distal conductor, and the distinctly large membraneous extension on the dorsal edge of the conductor (Figs 352 - 357).

Description

Description: Male (holotype). Large sized Coelotinae (Fig. 358). Total length could be more than 10.00. Dorsal shield of prosoma 6.45 long, 4.26 wide; abdomen damaged. AME smallest, about % the size of other eyes, which are subequal, or with PLE slightly smaller (AME 0.15, ALE 0.22, PME 0.22, PLE 0.18); AME separated from each other by of their diameter, from ALE by AME diameter; PME separated from each other by about half of their diameter, widely separated from PLE by more than their diameter (AME-AME 0.10, AME-ALE 0.16, ALE-PLE 0.07, PME-PME 0.09, PME-PLE 0.26, AME-PME 0.13) (Fig. 359). Labium longer than wide (L / W = 1.25) (Fig. 360). Chelicerae with 3 promarginal and 2 retromarginal teeth. Palp with a small patellar apophysis; RTA more than half of tibial length, with distinctly protruding distal end; lateral tibial apophysis large, close to RTA; cymbial furrow more than 2 / s of cymbial length; conductor long, extending posteriorly near base of embolus, with a large basal lamella and a dorsal apophysis; median apophysis spoon-shaped, elongate, free-standing along anterior edge; embolus long, filiform, proximal in origin, arising at 6: 30 o-clock-position, running half an oval, extending posteriorly to proximal part of tibia and anteriorly coiling beyond distal part of bulb (Figs 352 - 357). Female. Unknown.

Distribution

Distribution: China (Yunnan: Tengchong, Lushui) (Fig. 547).

Description

(Figs 361 - 406, 548)

Materials Examined

China Gongshan County CHINA Feng Xue Yakou C. Griswold China Pianma Township K. J. Guo China Pianma Township K. J. Guo China Pianma Township K. J. Guo China Cikai Township H. M. Yan China Cikai Township D. Kavanaugh, P. E. Marek, D. Z. Dong, X. C. Liang China Cikai Township H. M. Yan China Cikai Township D. Kavanaugh, P. E. Marek, D. Z. Dong, X. C. Liang China Cikai Township H. M. Yan China Lishadi subdistrict H. M. Yan, G. X. Peng China Gaoligongshan C. Griswold, H. M. Yan China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, H. M. Yan China Gaoligongshan C. Griswold, D. Kavanaugh China Lishadi subdistrict H. M. Yan, G. X. Peng China Yaping subdistrict H. M. Yan, G. X. Peng China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Shibali Forest Station C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gao-ligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan

Diagnosis

Diagnosis: D. pseudobrunneus is similar to D. incertus Wang 2003 and D. latusincertus sp. nov. in having similar epigynum and palp, but can be distinguished from D. latusincertus sp. nov. by the small body size, the small, anteriorly extending conductor in the male (Figs 363 - 364, 367 - 369), and the slightly anteriorly extending spermathecae in the female; from D. incertus by the more or less long patellar apophysis and the smooth posterior margin of embolus in the male, and anteriorly extending spermathecae in the female (Figs 361 - 362, 365, 375 - 406).

Description

Description: Female. See Wang (2003). Photos of habitus, eyes, labium and epigynum are provided in this study (Figs 365 - 366, 373 - 374, 375 - 406). Male (one male of CASENT 9020464 measured). The male is here described for the first time. Medium sized Coelotinae, total length 6.28 (Fig. 370). Dorsal shield of prosoma 3.28 long, 2.00 wide; opisthosoma 3.00 long, 2.15 wide. AME smallest, half the size of ALE; ALE largest, PLE slightly smaller than ALE, PME slightly smaller than PLE (AME 0.06, ALE 0.14, PME 0.13, PLE 0.11); anterior eyes close together; AME separated from each other by half of their diameter, from ALE by of AME diameter; PME separated from each other by l A of their diameter, from PLE by half of PME diameter (AME-AME 0.03, AME-ALE 0.02, PME-PME 0.04, PME-PLE 0.07, AME-PME 0.06) (Fig. 371). Labium length and width subequal (L / W = 1.0) (Fig. 372). Chelicerae with 3 promarginal and 2 retromarginal teeth. Palp with patellar apophysis moderately long, slender; RTA slightly less than or about half of tibial length, with moderately protruding distal end; lateral tibial apophysis large, distinctly separated from RTA; cymbial furrow small, less than half of cymbial length; conductor moderately long, with a slightly bifurcate apex, a small basal lamella, and a dorsal apophysis; median apophysis simple, not spoon-shaped; embolus short, filiform, prolateral in origin (Figs 363 - 364, 367 - 369).

Distribution

Distribution: China (Yunnan: Lushui, Fugong, Gongshan) (Fig. 548).

Description

(Figs 407 - 427, 547)

Materials Examined

China Gongshan County CHINA Cikai Township H. M. Yan China Cikai Township H. M. Yan China Bing Zhong Luo Township P. Hu China Pihe Township G. Tang China Lishadi Township G. Tang China Lishadi P. Paquin China Lishadi P. Paquin China Lishadi P. Paquin, D. Kavanaugh China Lishadi P. Paquin China Lishadi P. Paquin, D. Kavanaugh China Lishadi P. Paquin China Lishadi P. Paquin China Lishadi P. Paquin China Gaoligongshan C. Griswold, H. M. Yan China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, H. M. Yan China Gaoligongshan C. Griswold China LiShadi subdistrict H. M. Yan China Gaoligongshan C. Griswold, D. Kavanaugh China Pihe Township G. Tang China Gaoligongshan C. Griswold, H. M. Yan China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Gaoligongshan C. Griswold, D. Kavanaugh China Yaping subdistrict H. M. Yan, G. X. Peng China Gaoligongshan C. Griswold, D. Kavanaugh

Diagnosis

Diagnosis: This species is similar to D. capitulatus Wang 2003 in having similar spermathecae, absence of epigynal teeth, and anteriorly extending conductor, and prolaterally originated embolus. The female of this species has the atrial ridges extending laterally, ending posteriorly close to the epigastric furrow, forming a round-shaped median area (Figs 407 - 408), whereas the atrial ridges of D. capitulatus extend relatively medially, ending posteriorly anterior to the epigastric furrow, and forming a triangular-shaped median area (Fig. 44). The male can be recognized by the slender patellar apophysis, the slightly bifurcate, smoothly curving conductor, and the absence of a median apophysis (Figs 409 - 410).

Description

Description: Female. See Wang (2003). Photos of habitus, eyes, labium and epigynum are provided in this study (Figs 411 - 414, 418 - 424). Male (male of CASENT 9025826 measured). The male is here described for the first time. Large sized coe-lotine, total length 12.04 (Fig. 425). Dorsal shield of prosoma 5.41 long, 3.63 wide; opisthosoma 6.63 long, 4.89 wide. AME smallest, half the size of ALE; ALE largest, PLE slightly smaller than ALE, PME slightly smaller than PLE (AME 0.11, ALE 0.19, PME 0.16, PLE 0.18); AME separated from each other by their diameter, from PLE by slightly less than AME diameter; PME separated from each other by slightly less than their diameter, from PLE by slightly more than PLE diameter (AME-AME 0.11, AME-ALE 0.09, PME-PME 0.13, PME-PLE 0.20, AME-PME 0.14) (Fig. 426). Labium length and width subequal (L / W = 1.0) (Fig. 427). Chelicerae with 3 promarginal and 2 retromarginal teeth. Palp with patellar apophysis long, slender; RTA short, less than or about half of tibial length, with distinctly protruding distal end; lateral tibial apophysis large; cymbial furrow small, less than half of cymbial length; conductor moderately long, anteriorly extending, with a slightly bifurcate apex, a small basal lamella, and a dorsal apophysis; median apophysis absent; embolus short, filiform, prolateral in origin (Figs 409 - 410, 415 - 417).

Distribution

Distribution: China (Yunnan: Fugong, Gongshan) (Fig. 547).

Description

(Figs 428 - 448, 548)

Materials Examined

CHINA Luoshuidong C. Griswold, D. Kavanaugh, C. L. Long CHINA Yunnan China Bawan village G. Tang China Bawan village G. Tang China Bawan village G. Tang China Pass over Gaoligongshan at 2100 m C. Griswold, D. Kavanaugh, C-L. Long CHINA Nankang (National 320 Road) G. Tang China Nankang (National 320 Road) G. Tang China Nankang (National 320 Road) G. Tang China Nankang (National 320 Road) G. Tang China Nankang (National 320 Road) G. Tang China Nankang (National 320 Road) G. Tang China Longjiang Township H. M. Yan, K. J. Guo China 40 km of the road Bawan-Tengchong G. Tang China Bawan Township H. M. Yan, K. J. Guo China 39 - 41 km of the road Bawan-Tengchong G. Tang China Henghe village G. Tang China Bawan village G. Tang China Bawan village G. Tang China Bawan village G. Tang China Dahaoping village C. Griswold

Etymology

Etymology: The specific name refers to its similarity to D. spiralis; name in apposition.

Diagnosis

Diagnosis: This new species is similar to D. spiralis sp. nov. and D. paraspiralis sp. nov. in having coiled spermathecal tubes, a large atrium, the absence of epigynal teeth, a distally coiled conductor, and a simple median apophysis, but can be distinguished by the distally broad conductor and the proximally originating embolus in the male, and by the anterior atrial margin that extends posteriorly to at least half of the atrial length and the broad copulatory ducts in the female (Figs 428 - 431).

Description

Description: Male (holotype). Large sized Coelotinae (Fig. 444). Total length 10.2. Dorsal shield of prosoma 5.00 long, 3.14 wide; opisthosoma 5.20 long, 3.60 wide. AME smallest, 2 / s size of ALE; ALE largest; PLE slightly smaller than ALE, PME slightly smaller than PLE (AME 0.20, ALE 0.28, PME 0.24, PLE 0.26); anterior eyes equally separated by half of AME diameter; PME separated from each other by slightly more than half of PME diameter, from PLE by 4 / 5 of PME diameter (AME-AME 0.10, AME-ALE 0.09, PME-PME 0.13, PME-PLE 0.20, AME-PME 0.15) (Fig. 445). Labium slightly longer than wide (L / W = 1.09) (Fig. 446). Promargin of chelicera with 3, retromargin with 2 teeth. Palpal patellar apophysis tiny, barely visible; RTA approximately half of tibial length or slightly longer; lateral tibial apophysis present; cymbial furrow less than half of cymbial length; conductor broad, with a distinctly coiled apex, a well developed basal lamella, and a slender dorsal apophysis; median apophysis simple, not spoon-shaped; embolus filiform, long, proximal in origin, arising at 6 - o'clock-position, running half an oval, extending posteriorly to distal part of tibia and anteriorly coiling beyond distal part of bulb (Figs 430 - 431, 434 - 436). Female (paratype measured). Large sized Coelotinae, total length 10.8 (Fig. 441). Dorsal shield of prosoma 5.20 long, 3.80 wide; opisthosoma 5.60 long, 4.00 wide. AME smallest, 4 / 5 size of ALE; ALE largest; posterior eyes subequal in size, slightly smaller than ALE (AME 0.19, ALE 0.25, PME 0.23, PLE 0.24); AME separated from each other by % of their diameter, from ALE by half of AME diameter; PME separated from each other by half of their diameter, widely separated from PLE by slightly more than PME diameter (AME-AME 0.15, AME-ALE 0.09, PME-PME 0.12, PME-PLE 0.28, AME-PME 0.20) (Fig. 442). Labium slightly longer than wide (L / W = 1.06) (Fig. 443). Promargin of chelicera with 3, retromargin with 2 teeth. Epigynal teeth absent; atrium large, but shallow, with anterior margin extending posteriorly to a septum that extends half or more than half of atrial length; copulatory ducts large, originating anteriorly, extending posteriorly and then curving back anteriorly; spermathecae with bases widely separated by more than two times their width, stalks long, anteriorly extending and slightly converging; spermathecal heads arising distally (Figs 428 - 429, 432 - 433, 437 - 440).

Distribution

Distribution: China (Yunnan: Tengchong, Longling, Longyang, Luxi) (Fig. 548).

Description

(Figs 449 - 453, 546)

Materials Examined

China Tengchong

Diagnosis

Diagnosis: The female can be easily recognized by the presence of a tongue-shaped, weakly sclerotized median piece that arises from the anterior half of atrium, the large, anteriorly extending copulatory ducts, and the broad, long spermathecae (Figs 449 - 450).

Description

Description: Female. See Wang (2003). Photos of habitus, eyes, labium and epigynum are provided in this study (Figs 449 - 453). Male. Unknown.

Distribution

Distribution: China (Yunnan) (Fig. 546).

Description

(Figs 454 - 460, 548)

Materials Examined

CHINA Daxinde Township G. Tang

Etymology

Etymology: The species name came from Latin " quattour ", which means " the fourth ", and refers to the presence of four retromarginal cheliceral teeth; adjective.

Diagnosis

Diagnosis: The female of this new species is similar to D. yani sp. nov. in having epigynal teeth that are widely separated from each other and from the atrial margins, and spermathecae that extend anteriorly beyond the spermathecal heads, but can be distinguished by the indistinct anterior atrial margin and the relatively small spermathecae (Figs 454 - 455). The chelicerae have four retromarginal teeth in this new species, but two in D. yani sp. nov.

Description

Description: Female (holotype). Medium sized Coelotinae (Fig. 458). Total length 7.30. Dorsal shield of prosoma 3.80 long, 2.48 wide; opisthosoma 3.50 long, 2.40 wide. Median eyes subequal in size, slightly smaller than lateral eyes, which are subequal (AME 0.14, ALE 0.16, PME 0.14, PLE 0.17); AME separated from each other by 2 / s of AME diameter, from ALE by slightly less than 1 / 2 of AME diameter; PME separated from each other by slightly less than their diameter, from PLE by PLE diameter (AME-AME 0.09, AME-ALE 0.06, PME-PME 0.12, PME-PLE 0.16, AME-PME 0.16) (Fig. 459). Labium slightly longer than wide (L / W = 1.10) (Fig. 460). Chelicera with 3 promarginal and 4 retromarginal teeth. Epigynal teeth short, arising anterolaterally of atrium, distinctly separated from atrial margins, widely separated from each other by at least atrial width; atrium small, close to epigastric furrow, without distinct anterior margin and lateral margins; cop-ulatory ducts small, originating posteriorly; spermathecae with bases widely separated by at least their width, stalks broad, anteriorly extending and converging anterior of heads; spermathecal heads distinct, arising from anterior half of spermathecae (Figs 454 - 457). Male. Unknown.

Distribution

Distribution: China (Yunnan: Lushui) (Fig. 548).

Description

(Figs 461 - 467, 548)

Materials Examined

CHINA Jietou Township C. M. Yin, J. F. Hu, X. H. Yang

Etymology

Etymology: The species name came from Latin " renalis ", which means " of the kidneys ", and refers to the kidney-shaped spermathecal tubes; adjective.

Diagnosis

Diagnosis: The female of this new species can be easily distinguished from other Coelotinae by the laterally extending and expanding, kidney-shaped spermathecae (Figs 461 - 462). Both cheliceral pro- and retro-margins have three teeth.

Description

Description: Female (holotype). Large sized Coelotinae (Fig. 465). Total length 13.5. Dorsal shield of prosoma 6.25 long, 4.27 wide; opisthosoma 7.26 long, 4.95 wide. AME largest; ALE slightly smaller than AME; posterior eyes subequal, slightly smaller than ALE (AME 0.19, ALE 0.17, PME 0.16, PLE 0.16); anterior eyes equally separated by about half of AME diameter; PME separated from each other by their diameter, widely separated from PLE by 1.5 times PME diameter (AME-AME 0.08, AME-ALE 0.09, PME-PME 0.17, PME-PLE 0.25, AME-PME 0.19) (Fig. 466). Labium slightly longer than wide (Fig. 467). Chelicera with 3 promarginal and 3 retromarginal teeth. Epigynal teeth long, situated anterolaterally of atrium, widely separated from each other by more than atrial width; atrium large, but shallow, longer than wide, with distinct, continuous anterior margin, but indistinct lateral margins; epigynal hoods distinct, situated at level posterior to epigynal teeth; copulatory ducts large, originating anteriorly, closely set; spermathecae with small, slightly separated bases and long, slender, anteriorly extending, laterally diverging and enlarged stalks, which are broad, kidney-shaped; spermathecal heads small, arising from distal half of spermathecae (Figs 461 - 464). Male. Unknown.

Distribution

Distribution: China (Yunnan: Tengchong) (Fig. 548).

Description

(Figs 468 - 472, 549)

Materials Examined

China Tengchong County

Diagnosis

Diagnosis: The female of this new species can be easily recognized by the epigynal hoods that are situated relatively posteriorly at a level posterior to the epigynal teeth and the copulatory ducts that originate posteriorly between spermathecae, extending and coiling anteriorly and laterally (Figs 468 - 469).

Description

Description: Female. See Wang (2003). Photos of habitus, eyes, labium and epigynum are provided in this study (Figs 468 - 472). Male. Unknown.

Distribution

Distribution: China (Yunnan: Tengchong) (Fig. 549).

Description

(Figs 473 - 480, 550)

Materials Examined

China Tengchong County China Jietou Township X. J. Peng, X. P. Wang, P. Hu China Houqiao Township X. P. Wang, P. Hu China Gaoligongshan C. Griswold, D. Kavanaugh China Lishadi P. Paquin, D. Kavanaugh

Diagnosis

Diagnosis: The female can be easily recognized by the absence of epigynal teeth, the presence of broad spermathecae, the long atrium, and the subdistally arising spermathecal heads (Figs 473, 475 - 477).

Description

Description: Female. See Wang (2003). Photos of habitus, eyes, labium and epigynum are provided in this study (Figs 473 - 480). Male. Unknown.

Distribution

Distribution: China (Yunnan: Tengchong, Fugong) (Fig. 550).

Description

(Figs 481 - 484, 549)

Materials Examined

CHINA Gaoligongshan C. Griswold, D. Kavanaugh, C. L. Long CHINA Yunnan China Gaoligongshan C. Griswold, D. Kavanaugh, C. L. Long China Gaoligongshan C. Griswold, D. Kavanaugh, C. L. Long

Etymology

Etymology: The specific name refers to the coiled conductor apex and spermathecal tubes; adjective. Diagnosis: This species is similar to C. pseudospiralis sp. nov. and C. paraspiralis sp. nov. The female can be distinguished from C. pseudospiralis by the slightly protruding anterior atrial margin and from C. paraspiralis sp. nov. by the presence of broad diverticula on the anterior spermathecae and the less coiled spermathecae; the male by the long cymbial furrow, which is almost as long as the cymbium, the slender conductor apex, and the extremely long, slender embolus (Figs 481 - 484).

Description

Description: Male (holotype). Medium sized Coelotinae, total length 8.40. Dorsal shield of prosoma 4.00 long, 3.00 wide, opisthosoma 4.40 long, 3.00 wide. AME smallest, about % of the other eyes, which are about equal in size (AME 0.14, ALE 0.23, PME 0.20, PLE 0.20); AME separated from each other by half of their diameter, from PLE by about l A of AME diameter; PME separated from each other by slightly less than their diameter, from PLE by about % of PME diameter (AME-AME 0.07, AME-ALE 0.04, AME-PME 0.14, PME-PME 0.08, PME-PLE 0.15). Promargin of chelicera with 3, retromargin with 2 teeth. Palp with a tiny patellar apophysis; RTA slightly longer than half of tibial length, with broad, distinctly protruding distal end; lateral tibial apophysis large; cymbial furrow long, almost reaching distal cymbium; conductor broad, long, with a coiled, slender apex, a large basal lamella, and a slender dorsal apophysis; median apophysis simple, not spoon-shaped; embolus filiform, long, proximal in origin, arising at 6 - o'clock-position, running half an oval, extending posteriorly to patella, and coiling anteriorly beyond distal part of bulb; embolus with a small base (Figs 483 - 484). Female (paratype). Large sized Coelotinae, total length 11.4. Dorsal shield of prosoma 5.60 long, 3.50 wide, opisthosoma 5.80 long, 4.20 wide. AME smallest, 3 / 5 size of lateral eyes, which are subequal and the largest; PME slightly smaller than lateral eyes (AME 0.15, ALE 0.25, PME 0.21, PLE 0.26); AME separated from each other by slightly less than AME diameter, from ALE by half of AME diameter; PME separated from each other and from PLE by approximately AME diameter (AME-AME 0.13, AME-ALE 0.07, AME-PME 0.20, PME-PME 0.22, PME-PLE 0.20). Promargin of chelicera with 3, retromargin with 2 teeth. Epig-ynal teeth absent; atrium large, but shallow, with anterior margin extending posteriorly to less than half of atrial length; lateral atrial margins distinct, extending posteriorly and reaching epigastric furrow; copulatory ducts large, originating and extending medially; spermathecae with bases widely separated, stalks long, anteriorly extending and converging, then curved posteriorly and formed ring-shaped vulva; anterior portion of spermathecae slightly expanded to broad diverticula; spermathecal heads distinct, arising distally from posterior part of epigynum (Figs 481 - 482).

Distribution

Distribution: China (Yunnan: Lushui) (Fig. 549).

Description

(Figs 485 - 492, 550)

Materials Examined

CHINA Nankang (National 320 Road) G. Tang

Etymology

Etymology: The specific name is in honour of Guo Tang at HNU who collected the specimen; noun in genitive case.

Diagnosis

Diagnosis: Males of this new species can be easily recognized by the broad, distally extending conductor, the broad, slightly bifurcate patellar apophysis, and the slightly spoon-shaped median apophysis (Figs 485489).

Description

Description: Male (holotype). Medium sized Coelotinae (Fig. 490). Total length 9.15. Dorsal shield of prosoma 4.65 long, 3.26 wide; opisthosoma 4.50 long, 2.96 wide. Anterior eyes subequal in size, largest; PLE slightly smaller, PME slightly smaller than PLE (AME 0.17, ALE 0.17, PME 0.12, PLE 0.14); anterior eyes equally separated by about of their diameter; PME separated from each other by their diameter, from PLE by about 1.5 times PME diameter (AME-AME 0.05, AME-ALE 0.04, PME-PME 0.13, PME-PLE 0.20, AME-PME 0.17) (Fig. 491). Labium slightly longer than wide (L / W = 1.08) (Fig. 492). Promargin of chelicera with 3, retromargin with 2 teeth. Palp with a broad, slightly bifurcate patellar apophysis; RTA almost as long as tibia, with distinctly protruding distal end; lateral tibial apophysis large, close to RTA; cymbial furrow short, 1 / 3 of cymbial length; conductor broad, extending distally, with a small apophysis on dorsal edge, a less developed lamella, and a large dorsal apophysis; median apophysis simple, slightly spoon-shaped; embolus short, more or less broad, prolateral in origin (Figs 485 - 489). Female. Unknown.

Distribution

Distribution: China (Yunnan: Longling) (Fig. 550).

Description

(Figs 493 - 500, 550)

Materials Examined

CHINA Pianma Township C. Griswold

Etymology

Etymology: The species name is derived from the Latin words " tri " and " dens ", meaning " three " and " tooth " respectively, and refers to the presence of three promarginal and three retromarginal cheliceral teeth; term in apposition.

Diagnosis

Diagnosis: This new species can be easily recognized by the absence of a patellar apophysis, the short RTA, the absence of a lateral tibial apophysis, the long conductor, and the absence of a median apophysis (Figs 493 - 497). Both the cheliceral promargin and retromargin have 3 teeth.

Description

Description: Male (holotype). Medium sized Coelotinae (Fig. 498). Total length 6.15. Dorsal shield of prosoma 3.15 long, 1.82 wide; opisthosoma 3.00 long, 1.87 wide. AME smallest, about % size of other eyes, which are subequal (AME 0.08, ALE 0.12, PME 0.12, PLE 0.12); anterior eyes equally separated by half of AME diameter; posterior eyes equally separated by slightly less than their diameter (AME-AME 0.04, AME-ALE 0.04, PME-PME 0.09, PME-PLE 0.09, AME-PME 0.10) (Fig. 499). Labium length and width subequal (L / W = 1.0) (Fig. 500). Cheliceral pro- and retromargins with 3 teeth. Palp without patellar apophysis; RTA short, less than half of tibial length; lateral tibial apophysis absent; cymbial furrow short, less than half of cymbial length; conductor long, extending posteriorly to distal tibia, with a small basal lamella; conductor dorsal apophysis absent; median apophysis absent; embolus short, filiform, prolateral in origin (Figs 493497). Female. Unknown.

Distribution

Distribution: China (Yunnan: Lushui) (Fig. 550).

Description

(Figs 501 - 504, 550)

Materials Examined

China Kunming

Diagnosis

Diagnosis: The female is similar to D. ornatus (Wang, Yin, Peng & Xie 1990) in having a large atrium and broad copulatory ducts, but this species has epigynal teeth and epigynal hoods that are posteriorly situated close to the epigastric furrow (Figs 502 - 503).

Description

Description: Female. See Wang & Jaeger (2007). Photos of habitus, eyes and epigynum are provided in this study (Figs 501 - 504). Male. Unknown.

Distribution

Distribution: China (Yunnan: Kunming) (Fig. 550).

Description

(Fig. 549)

Materials Examined

China Luoping County

Diagnosis

Diagnosis: This species is similar to D. degenaratus (Liu & Li 2009) in having a long patellar apophysis and a short, slender conductor in the male, and a similar female epigynum, but can be distinguished by the proximally originated embolus in the male and the slender epigynal teeth in the female (Liu & Li 2009: figs 10 - 14).

Description

Description: See Liu & Li (2009).

Distribution

Distribution: China (Yunnan: Luoping) (Fig. 549).

Description

(Fig. 550)

Materials Examined

China Zhongdian County

Diagnosis

Diagnosis: The female of this species has a similar epigynum to D. incertus Wang 2003 and related species, but can be easily distinguished by the posteriorly arising spermathecal heads (Wang & Jaeger 2010: figs 3 - 5).

Description

Description: Female. See Wang & Jaeger (2010). Male. Unknown.

Distribution

Distribution: China (Yunnan: Zhongdian) (Fig. 550).

Description

(Figs 505 - 511, 550)

Materials Examined

CHINA along Dulong Valley Road about 38 - 41 km X. Xu, C. M. Yin

Etymology

Etymology: The specific name is in honour of Xiang Xu at HNU who collected the holotype specimen; noun in genitive case.

Diagnosis

Diagnosis: Females can be easily recognized by the posteriorly situated epigynal hoods, the distinct atrium, and the broad, closely set copulatory ducts that cover most of the spermathecal tubes in dorsal view (Figs 505 - 508).

Description

Description: Female (holotype). Large sized Coelotinae (Fig. 509). Total length 12.45. Dorsal shield of prosoma 6.25 long, 4.60 wide; opisthosoma 6.20 long, 4.50 wide. Eyes subequal in size, with AME and PME slightly smaller than lateral eyes (AME 0.20, ALE 0.22, PME 0.19, PLE 0.22); anterior eyes equally separated by slightly more than half of AME diameter; PME separated from each other by approximately their diameter, widely separated from PLE by about 1.5 times PME diameter (AME-AME 0.13, AME-ALE 0.13, PME-PME 0.20, PME-PLE 0.32, AME-PME 0.22) (Fig. 510). Labium slightly longer than wide (L / W = 1.10) (Fig. 511). Promargin of chelicera with 3 teeth, retromargin with 2. Epigynal teeth short, arising anterolaterally of atrium and widely separated from each other by slightly more than atrial width; atrium small, but distinct, wider than long, with a small tongue-shaped piece in anterior part of atrium; anterior atrial margin distinct; epigynal hoods situated posteriorly close to epigastric furrow, at same level as anterior atrial margin and epig-ynal teeth; copulatory ducts broad, originating posteriorly between spermathecae, extending anteriorly and covering most of spermathecae in dorsal view; spermathecae with small, round bases, which are widely separated by at least their width; spermathecal stalks extending, converging, and closely set anteriorly; spermathe-cal head arising anteriorly and extending laterally (Figs 505 - 508). Male. Unknown.

Distribution

Distribution: China (Yunnan: Gongshan) (Fig. 550).

Description

(Figs 512 - 532, 549)

Materials Examined

CHINA Gaoligongshan C. Griswold, H. M. Yan CHINA Bing Zhong Luo Township Hu Peng China Bingzhongluo Township J. A. Miller, D. Kavanaugh China Lishadi Township G. Tang China Lumadeng P. Paquin China Lishadi P. Paquin China Gaoligongshan C. Griswold China Lishadi P. Paquin, D. Kavanaugh

Discussion

Notes: The female specimen CASENT 9022569 (Figs 529 - 530) has a similar epigynum in ventral view, but very different spermathecal tubes. Due to the limited number of specimens examined, we temporarily list it here as D. yani sp. nov.

Etymology

Etymology: The specific name is a patronym in honor of one of the collectors of the type specimen, Hengmei Yan at HNU; noun in genitive case.

Diagnosis

Diagnosis: The female of this new species is similar to D. quattour sp. nov. in having epigynal teeth that are widely separated from each other and from the atrial margin, and spermathecae that extend anteriorly of the spermathecal heads, but can be distinguished by the presence of a distinct anterior atrial margin and atrial ridges (Figs 512 - 515).

Description

Description: Female (holotype). Medium sized Coelotinae, total length 9.46 (Fig. 516). Dorsal shield of prosoma 4.26 long, 3.00 wide; opisthosoma 5.20 long, 3.50 wide. AME smallest, half the size of ALE; ALE largest; PME subequal to AME in size, PLE slightly larger than PME (AME 0.11, ALE 0.20, PME 0.12, PLE 0.14); AME separated from each other by slightly more than their diameter, from ALE by AME diameter; PME separated from each other by slightly more than their diameter, widely separated from PLE by about 2 times PME diameter (AME-AME 0.14, AME-ALE 0.12, PME-PME 0.15, PME-PLE 0.22, AME-PME 0.12) (Fig. 517). Labium longer than wide (L / W = 1.20) (Fig. 518). Promargin of chelicera with 3, retromargin with 2 teeth. Epigynal teeth short, widely separated from each other by about atrial width, situated anteriorly of atrium and distinctly separated from anterior atrial margin; atrium small, wider than long, close to epigastric furrow, with anterior margin slightly extending posteriorly; copulatory ducts small, originating posteriorly between spermathecae; spermathecae broad, with bases distinctly separated, anteriorly extending and closely set; spermathecal heads small, arising from distal half of spermathecae, with spermathecae extending anterior to their heads (Figs 512 - 515, 519 - 532). Male. Unknown.

Distribution

Distribution: China (Yunnan: Fugong, Gongshan) (Fig. 549).

Taxon Treatment

  • Xin-Ping Wang; Charles E. Griswold; Jeremy A. Miller; 2010: Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains, Zootaxa 2593: -1--1. doi
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