Crassomicrodus

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Taxonavigation

Ordo: Hymenoptera
Familia: Braconidae

Name

Crassomicrodus Ashmead, 1900 – Wikispecies link – Pensoft Profile

• Crassomicrodus Ashmead 1900. Type species Microdus fulvescens Cresson 1865, designated by Ashmead 1900 [Examined].
• Epimicrodus Ashmead 1900. Type species Microdus divisus Cresson 1873, designated by Ashmead 1900 [Examined].
• Crassomicrodus Ashmead= (Epimicrodus Ashmead) synonymized by Bradley 1916.

Diagnosis

Crassomicrodus species can be distinguished from other agathidines with the following combination of characters: simple tarsal claws, without basal lobes, apicolateral pegs of the hind tibia are hair-like, labio-maxillary complex not elongate; mandible with two teeth; and metasomal tergum 1 smooth.

Description

Head. Transverse or triangular; area between antennal sockets with a median pyramidal-shaped elevation or transverse; gena not bulging to distinctly bulging; labio-maxillary complex not elongate, mandible with two teeth, antenna with 25 to 43 flagellomeres.

Mesosoma

Pronotum surface smooth or punctuate; notauli from lacking to impressed; anterolateral edges of scutellum with or without a small acute projection; lateral scutellar depression from smooth to crenulate; dorsal surface of propodeum from rugulose to reticulate rugose; subalar lobe separated from mesopleuron by a wide or narrow groove; metapleuron from smooth to reticulate rugose; inner spur of hind tibia from 0.47 to 0.78 times longer than basitarsus; tarsal claw without basal lobe; outer apex of the hind tibia without flattened pegs; forewing vein R1 0.47–0.70 times longer than RS; crossvein r arising before or beyond middle of stigma. Metasoma. Metasomal median tergite 1 smooth; apical width 1.78–3.92 times longer than basal width; ovipositor sheaths and ovipositor variable in length; metasoma 1.00–1.42 times longer than mesosoma.

Key to the New World species of Crassomicrodus Ashmead

Key to the New World species of Crassomicrodus Ashmead — J. Figueroa , M. Sharkey , J. Nápoles , J. García , A. Martínez , V. López-Martínez , S. Pineda

1		Head triangular in frontal view (Figs 2a, 4a, 5a, 6a, 10a, 12a, 14a); gena not bulging; area between antennal sockets with a median pyramidal-shaped elevation (Figs 2a, 4a, 5a, 6a, 10a, 12a, 14a); length of ventrolateral margin of clypeus similar to diameter of tentorial pit (Fig. 1a, 8a)    2
–		Head transverse in frontal view (Figs 1a, 3a, 7a, 8a, 9a, 11a, 13a), if somewhat triangular then length of ventrolateral margin of clypeus longer than diameter of tentorial pit (Fig. 3a); gena bulging or at least slightly bulging (Figs 1a, 3a, 7a, 8a, 9a, 11a, 13a); area between antennal sockets variable in shape    10
2	(1)	Malar space at least 0.8 times as long as eye height (Figs 5a, 14a)    3
–		Malar space at most 0.6 times as long as eye height (Figs 2a, 4a, 6a, 10a, 12a)    4
3	(2)	Scutellar sulcus with 3 or 4 carinae; second submarginal cell quadrangular; fore and middle legs black; body length 6.95–8.60 mm    Crassomicrodus divisus (Cresson)
–		Scutellar sulcus with 1 carina; second submarginal cell triangular; fore and middle legs yellowish-orange; body length 4.20–6.48 mm    Crassomicrodus pallens (Cresson)
4	(2)	Forewing vein R1 at most half the length of RS; head yellowish-orange (Fig. 14a)    Crassomicrodus pallens (Cresson)
–		Forewing vein R1 at least 0.6 times longer than RS; head black (Fig. 4a)    5
5	(4)	Hind wing vein 1M less 1.0 times as long as 1r-m; body black; wings strongly infumate (Fig. 4e); body length at least 8.1 mm    Crassomicrodus costaricensis sp. n.
–		Hind wing vein 1M at least 1.4 times as long as 1r-m; body at least with some areas yellowish-orange or yellowish-red (Figs 2d, 6e, 12e), if body dark then body length less than 7.9 mm; wings at most slightly infumate (Fig. 10e)    6
6	(5)	Head and mesosoma black (Figs 2d, 12e); posterior surface of antennal sockets at least slightly rugulose    7
–		Head and mesosoma usually with some areas yellowish-orange or yellowish-red (Figs 6cde, 10cde), if entirely black then the posterior surface of antennal sockets smooth (Fig. 10b)    9
7	(6)	Subalar lobe separated from mesopleuron by a wide groove (Fig. 6c); body length 7.10–7.65 mm; 38–40 flagellomeres    8
–		Subalar lobe separated from mesopleuron by a narrow groove (Fig. 2c); body length 5.20–5.50 mm; 31–33 flagellomeres    Crassomicrodus azteca sp. n.
8	(7)	Hind legs black (Fig. 12e); wings hyaline; posterior area of antennal sockets rugulose (Fig. 12b)    Crassomicrodus oaxaquensis sp. n.
–		Hind femora yellowish-orange (Fig. 6e); wings slightly infumate; posterior area of antennal sockets rugose (Fig. 6b)    Crassomicrodus jalisciensis sp. n.
9	(6)	Setae length on body surface slightly similar to setae length at base of mandible (6ac); posterior surface of antennal sockets rugose (Fig. 6b); frons deeply excavated; pronotum punctulate (Fig. 6c); propodeum reticulate rugulose    Crassomicrodus jalisciensis sp. n.
–		Setae length on body surface distinctly smaller than setae length at base of mandible (Figs 10ac); posterior surface of antennal sockets smooth (Fig. 10b); frons excavated; pronotum smooth (Fig. 10c); propodeum rugose    Crassomicrodus nigriceps (Cresson)
10	(1)	Area between antennal sockets with a median pyramidal-shaped elevation (Figs 3a, 7a)    11
–		Area between antennal sockets at least weakly transverse or with a median elevation in trapezoidal shape (Figs 1a, 8a, 9a, 11a, 13a)    14
11	(10)	Frons at least slightly excavated (Figs 3ab, 8ab); notauli impressed (Figs 3d, 8d); forewing M+CU pigmented over most its length; 34–41 flagellomeres    12
–		Lead statement missing!    Frons not excavated (Figs 7ab); notauli not impressed (Fig. 7d); forewing M+CU unpigmented over most its length; 25–28 flagellomeres C. mariae sp. n.
12	(11)	Length of ventrolateral margin of clypeus similar to diameter of tentorial pit (Fig. 8a)    13
–		Length of ventrolateral margin of clypeus distinctly longer than diameter of tentorial pit (Fig. 3a)    Crassomicrodus clypealis sp. n.
13	(12)	Inner spur of hind tibia distinctly longer than half of basitarsus length (0.60–0.72); gena bulging (Fig. 8a); lateral depression of scutellum smooth, only its ventral edge with small punctures    Crassomicrodus melanopleurus Ashmead
–		Inner spur of hind tibia less than half length of basitarsus (a few specimens with the spur at most reaching 0.54 times); gena distinctly bulging; lateral depression of scutellum rugose and fovelate    Crassomicrodus fulvescens (Cresson)
14	(10)	Head and mesosoma black; anterolateral edges of scutellum lacking small acute projection    15
–		Head and mesosoma at least with some areas yellowish-orange or yellowish-red (Figs 1acde); anterolateral edges of scutellum with small acute projection (Fig. 1df)    Crassomicrodus apicipennis Muesebeck
15	(14)	Notauli not impressed over most of mesoscutum (Fig. 9c); metasoma black; setae at base of mandible similar in size to setae on rest of body surface (9ad); length of setae in scutellar disk 0.18 to 0.20 mm; ovipositor sheaths of female at least 1.83 mm in length    Crassomicrodus muesebecki Marsh
–		Notauli impressed over most of mesoscutum (Fig. 11c, 13d); metasoma yellowish-orange (Figs 11e, 13e); setae at base of mandible distinctly longer than setae on rest of body surface (11ad, 13ac); length of setae in scutellar disk less than 0.17 mm; ovipositor sheaths of female at most 0.22 mm in length    16
16	(15)	Area between antennal sockets with a median elevation in trapezoidal shape (Fig. 13a); gena distinctly bulging (Fig. 13a); malar space 0.54–0.59 times longer than eye height; 31–34 flagellomeres; body length 6.70–7.08 mm    Crassomicrodus olgae sp. n.
–		Area between antennal sockets with a median transverse elevation (Fig. 11a); gena bulging (Fig. 11a); malar space 0.38–0.47 times longer than eye height; 28–31 flagellomeres; body length 3.95–5.35 mm    Crassomicrodus nigrithorax Muesebeck

Species descriptions of Crassomicrodus

Taxon Treatment

• Figueroa, J; Sharkey, M; Nápoles, J; García, J; Martínez, A; López-Martínez, V; Pineda, S; 2011: Revision of the new world genus Crassomicrodus Ashmead (Hymenoptera, Braconidae, Agathidinae), with an identification key to species ZooKeys, 142: 27-75. doi

Images

Figure 1. Crassomicrodus apicipennis. Female a anterior view of head, arrow indicates a median transverse elevation with two weakly defined lateral tubercles, ventrolateral margin of clypeus, and tentorial pit b dorsal view of head, arrow indicates groove between lateral ocelli with small foveolae c lateral view of mesosoma, arrows indicate pronotum and metapleuron d dorsal view of mesosoma, arrows indicate anterolateral edge of scutellum and propodeum e female habitus f dorsal view of scutellum, arrow indicates anterolateral edge of scutellum with a small acute projection.  ' Figure 2. Crassomicrodus azteca sp. n.Female a anterior view of head, arrow indicates a median pyramidal-shaped elevation b dorsal view of head and mesosoma, arrow indicates posterior surface of antennal sockets slightly rugulose c lateral view of mesosoma, arrow indicates subalar lobe separated from mesopleuron by narrow groove d female habitus, arrow indicates triangular-shaped second submarginal cell.  Figure 3. Crassomicrodus clypealis sp. n. Female a anterior view of head, arrows indicate a median pyramidal-shaped elevation with two weakly defined tubercles, ventrolateral margin of clypeus, and tentorial pit b dorsal view of head, arrow indicates frons deeply excavated with a pair of microfoveolate grooves c female habitus d dorsal view of mesosoma, arrows indicate impressed notauli e lateral view of mesosoma, arrow indicates subalar lobe separated from mesopleuron by narrow groove.  Figure 4. Crassomicrodus costaricensis sp. n. Female a anterior view of head, arrow indicates a median pyramidal-shaped elevation b dorsal view of head c lateral view of mesosoma, arrow indicates subalar lobe separated from mesopleuron by wide groove d dorsal view of mesosoma e female habitus.  Figure 5. Crassomicrodus divisus. Female a anterior view of head, arrow indicates a median pyramidal-shaped elevation b dorsal view of head c. dorsal view of mesosoma d lateral view of mesosoma, arrow indicates subalar lobe separated from mesopleuron by wide groove e. female habitus.  Figure 6. Crassomicrodus jalisciensis. Female a anterior view of head, arrow indicates a median pyramidal-shaped elevation b dorsal view of head, arrow indicates posterior surface of antennal sockets rugulose c lateral view of mesosoma, arrows indicate pronotum and subalar lobe separated from mesopleuron by wide groove d dorsal view of mesosoma e female habitus.  Figure 7. Crassomicrodus mariae. Female a anterior view of head, arrow indicates a median pyramidal-shaped elevation b dorsal view of head, arrow indicates frons not excavated c lateral view of mesosoma, arrow indicates punctate pronotum d dorsal view of mesosoma, arrow indicates notauli not impressed e female habitus.  Figure 8. Crassomicrodus melanopleurus. Female a anterior view of head, arrows indicate a median pyramidal-shaped elevation with two weakly defined tubercles ventrolateral margin of clypeus, and tentorial pit b dorsal view of head, arrow indicates frons excavated with a pair of microfoveolate grooves c lateral view of mesosoma d dorsal view of mesosoma, arrow indicates impressed notauli e female habitus, arrow indicates inner spur of hind tibia.  Figure 9. Crassomicrodus muesebecki. Female a anterior view of head, arrows indicate a median transverse elevation with two weakly defined tubercles b dorsal view of head c dorsal view of mesosoma, arrow indicates notauli not impressed d lateral view of mesosoma e female habitus.  Figure 10. Crassomicrodus nigriceps. Female a anterior view of head, arrows indicate a median pyramidal-shaped elevation with two weakly defined tubercles b dorsal view of head, arrow indicates posterior surface of antennal sockets smooth c lateral view of mesosoma, arrow indicates pronotum smooth d dorsal view of mesosoma e female habitus.  Figure 11. Crassomicrodus nigrithorax. Female a anterior view of head, arrows indicate a median transverse elevation with two weakly defined tubercles b dorsal view of head c dorsal view of mesosoma, arrow indicates notauli impressed d lateral view of mesosoma e female habitus.  Figure 12. Crassomicrodus oaxaquensissp. n. Female a anterior view of head, arrows indicate a median pyramidal-shaped elevation with two weakly defined tubercles b dorsal view of head, arrow indicates posterior surface of antennal sockets rugulose c lateral view of mesosoma, arrow indicates subalar lobe separated from mesopleuron by wide groove d dorsal view of mesosoma e female habitus.  Figure 13. Crassomicrodus olgaesp. n. Female a anterior view of head, arrows indicate a median elevation in trapezoidal shape with two weakly defined tubercles b dorsal view of head c lateral view of mesosoma d dorsal view of mesosoma, arrow indicates notauli impressed e female habitus.  ' Figure 14. Crassomicrodus pallens. Female a anterior view of head, arrows indicate a median pyramidal-shaped elevation with two weakly defined tubercles b dorsal view of head c lateral view of mesosoma d dorsal view of mesosoma e female habitus.