Chlaenius (Lithochlaenius) formosensis

Notice:

This page is derived from the original publication listed below, whose author(s) should always be credited. Further contributors may edit and improve the content of this page and, consequently, need to be credited as well (see page history). Any assessment of factual correctness requires a careful review of the original article as well as of subsequent contributions. If you are uncertain whether your planned contribution is correct or not, we suggest that you use the associated discussion page instead of editing the page directly. This page should be cited as follows (rationale): Liu Y, Kavanaugh D, Shi H, Liang H (2011) A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species. ZooKeys 128 : 15–52, doi. Versioned wiki page: 2011-09-09, version 14099, https://species-id.net/w/index.php?title=Chlaenius_(Lithochlaenius)_formosensis&oldid=14099 , contributors (alphabetical order): Pensoft Publishers. Citation formats to copy and paste BibTeX: @article{Liu2011ZooKeys128, author = {Liu, Ye AND Kavanaugh, David H. AND Shi, Hongliang AND Liang, Hongbin}, journal = {ZooKeys}, publisher = {Pensoft Publishers}, title = {A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species}, year = {2011}, volume = {128}, issue = {}, pages = {15--52}, doi = {10.3897/zookeys.128.1804}, url = {http://www.pensoft.net/journals/zookeys/article/1583/abstract}, note = {Versioned wiki page: 2011-09-09, version 14099, https://species-id.net/w/index.php?title=Chlaenius_(Lithochlaenius)_formosensis&oldid=14099 , contributors (alphabetical order): Pensoft Publishers.} } RIS/ Endnote: TY - JOUR T1 - A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species A1 - Liu Y A1 - Kavanaugh D A1 - Shi H A1 - Liang H Y1 - 2011 JF - ZooKeys JA - VL - 128 IS - UR - http://dx.doi.org/10.3897/zookeys.128.1804 SP - 15 EP - 52 PB - Pensoft Publishers M1 - Versioned wiki page: 2011-09-09, version 14099, https://species-id.net/w/index.php?title=Chlaenius_(Lithochlaenius)_formosensis&oldid=14099 , contributors (alphabetical order): Pensoft Publishers. M3 - doi:10.3897/zookeys.128.1804 Wikipedia/ Citizendium: <ref name="Liu2011ZooKeys128">{{Citation | author = Liu Y, Kavanaugh D, Shi H, Liang H | title = A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species | journal = ZooKeys | year = 2011 | volume = 128 | issue = | pages = 15--52 | pmid = | publisher = Pensoft Publishers | doi = 10.3897/zookeys.128.1804 | url = http://www.pensoft.net/journals/zookeys/article/1583/abstract | pmc = | accessdate = 2025-03-31 }} Versioned wiki page: 2011-09-09, version 14099, https://species-id.net/w/index.php?title=Chlaenius_(Lithochlaenius)_formosensis&oldid=14099 , contributors (alphabetical order): Pensoft Publishers.</ref> See also the citation download page at the journal.

Taxonavigation

Ordo: Coleoptera
Familia: Carabidae
Genus: Chlaenius

Name

Chlaenius formosensis Lorenz – Wikispecies link – Pensoft Profile

• Chlaenius noguchii formosanus Habu, 1965: 86 (nec Jedlička, 1935:5)
• Chlaenius formosanus Morita, 1993:161
• Chlaenius formosensis Lorenz, 1998:339 (replacement name); Kirschenhofer, 2005:491

Specimens examined

Total 24 specimens: China: 8 males and 12 females (CCCC), "Taiwan Prov., Hsinchu, Wufeng, 1996.08.03, Chen C.C. coll.”; 3 males and 1 female (CCCC), "Taiwan Prov., Hsinchu, Wufeng, 1998.4.11, Chen C.C. coll.

Diagnosis

Antennomere 1 cylindrical (Fig. 56); antennomeres 1–3 brown; intervals 1–7 convex, glabrous medially, with one row of pubescence laterally (Fig. 84); apex of sternum VII rounded (Figs 101–102); lamella of aedeagus slightly truncate at apex (Fig. 125), thick, bent ventrally (Fig. 126).

Description

Total length = 16.0–17.0 mm (mean = 16.45), width = 5.87–6.40 mm (mean = 5.99); HW = 2.90–3.00 mm (mean = 2.96), EYL = 1.10–1.15 mm (mean = 1.12), ratio Ant3/Ant1 = 1.71–1.88 (mean = 1.76), PL/PW = 0.81–0.88 (mean = 0.85), EL/EW = 1.68–1.86 (mean = 1.76), EL/ PL = 1.32–1.48 (mean = 1.39).
Head and pronotum black, with green or coppery metallic luster; elytra black; ventral surface black; mandibles, trochanters, and tarsomeres dark brown; antennae, palpomeres, femora, and tibiae yellow to brown.
Head with vertex nearly glabrous, very sparsely punctate and pubescent near eyes; Antennomere 1 cylindrical (Fig. 56); labrum slightly emarginate at apex. Pronotum with disk convex, nearly smooth, glabrous; basal foveae small, deep, finely punctate, sparsely pubescent. Elytra with intervals convex, intervals 1–7 glabrous medially, with a row of pubescence laterally near striae (Figs 24, 84), intervals 8 and 9 with decumbent pubescence throughout; striae deep, punctate; humeral angle obtuse (Fig. 70). Abdominal sterna IV-VI nearly glabrous medially, densely pubescent laterally; sternum VII broadly rounded at apex in both sexes (Figs 101–102). Aedeagus with lamella subtruncate at apex (Fig. 125), thick and bent laterally (Fig. 126).

Geographical distribution

Fig. 158. Known only from Taiwan.

Remarks

Mensural data cited in the description were based on measurements obtained from 5 males and 5 females selected for maximum variation.
This species was first described as a subspecies of Chlaenius noguchii. Later, Morita (1993) upgraded it to a distinct species. However, the name Chlaenius formosanus was preoccupied by another Chlaenius species of Jedlička, and therefore Lorenz (1998) renamed it Chlaenius formosensis.
Kirschenhofer (2005: 491) considered Chlaenius formosensis Lorenz to be a junior synonym of Chlaenius formosanus Jedlička (= Chlaenius rambouseki Lutshnik, see below) with no comparison. Based on Habu"s original description (type unavailable according to Morita), named specimens of Chlaenius formosensis in Morita's collection (corresponding author and checked by HBL in 2009), and specimens in Chen Chanchin's collection, we treat Chlaenius formosensis as a distinct species. Its members differ from those of Chlaenius rambousekiin having the vertex of the head and pronotal disk glabrous (both sparsely punctate and pubescent in Chlaenius rambouseki), intervals 1–7 glabrous medially (wholly pubescent in Chlaenius rambouseki), and the male aedeagus stout (slender in Chlaenius rambouseki).
Chlaenius formosensis adults are similar to those of Chlaenius noguchii in elytra pubescence, but differ from the latter in having antennomere 1 cylindrical (coniform in Chlaenius noguchii), pronotal disk smooth (finely punctate in Chlaenius noguchii), and male aedeagus stout (slender in Chlaenius noguchii).

Taxon Treatment

• Liu, Y; Kavanaugh, D; Shi, H; Liang, H; 2011: A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species ZooKeys, 128: 15-52. doi

Images

Figures 22–27. Habitus ofChlaenius (Lithochlaenius) spp22 Chlaenius agilis Chaudoir, holotype, dorsal view23 Chlaenius agilis Chaudoir, holotype, ventral view24 Chlaenius formosensis Lorenz, male, in CCCC, dorsal view25 Chlaenius formosensis Lorenz, ventral view26 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, dorsal view27 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, ventral view. Scale line = 5.0 mm.  Figures 52–65. Antennomere 1 of Chlaenius (Lithochlaenius) species52 Chlaenius chuanqianensis sp. n., holotype53 Chlaenius linwensini sp. n., holotype54 Chlaenius propeagilis sp. n., holotype55 Chlaenius agilis Chaudoir, holotype56 Chlaenius formosensis Lorenz, CCCC57 Chlaenius leishanensis Kirschenhofer, IZCAS58 Chlaenius noguchii Bates, IZCAS59 Chlaenius wrasei Kirschenhofer, IZCAS60 Chlaenius agiloides Jedlička, IZCAS61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS62 Chlaenius anchomenoides Bates, paratype63 Chlaenius nuristanus Jedlička, paratype64 Chlaenius nuristanus a. rubridipes Jedlička, paratype65 Chlaenius rambouseki Lutshnik from Taiwan,CCCC. Scale line = 1.0 mm.  Figures 66–79. Humeral region of elytra of Chlaenius (Lithochlaenius) spp66 Chlaenius chuanqianensis sp. n., holotype67 Chlaenius linwensini sp. n., holotype68 Chlaenius propeagilis sp. n., holotype69 Chlaenius agilis Chaudoir, holotype70 Chlaenius formosensis Lorenz, CCCC71 Chlaenius leishanensis Kirschenhofer, IZCAS72 Chlaenius noguchii Bates, IZCAS73 Chlaenius wrasei Kirschenhofer, IZCAS74 Chlaenius agiloides Jedlička, IZCAS75 Chlaenius rambouseki Lutshnik from Ussuri region, ZCAS76 Chlaenius anchomenoides Bates, paratype77 Chlaenius nuristanus Jedlička, paratype78 Chlaenius nuristanus Jedlička, paratype79 Chlaenius rambousekiLutshnik from Taiwan, CCCC. Scale line = 1.0 mm.  Figures 80–93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype81 Chlaenius linwensini sp. n., holotype82 Chlaenius propeagilis sp. n., holotype83 Chlaenius agilis Chaudoir, holotype84 Chlaenius formosensis Lorenz, CCCC85 Chlaenius leishanensis, IZCAS86 Chlaenius noguchii Bates, IZCAS87 Chlaenius wrasei Kirschenhofer, IZCAS88 Chlaenius agiloides Jedlička, IZCAS89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS90 Chlaenius anchomenoides Bates, paratype91 Chlaenius nuristanusJedlička, paratype92 Chlaenius nuristanus Jedlička, paratype93 Chlaenius rambouseki Lutshnik from Taiwan,CCCC. Scale line = 1.0 mm.  Figures 94–116. Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species94 Chlaenius chuanqianensis sp. n., male, holotype95 Chlaenius chuanqianensis sp. n., female, paratype96 Chlaenius linwensini sp. n., male, holotype97 Chlaenius linwensini sp. n., female, paratype98 Chlaenius propeagilis sp. n., male, holotype99 Chlaenius propeagilis sp. n., female, paratype100 Chlaenius agilis Chaudoir, male, holotype101 Chlaenius formosensis Lorenz, male102 Chlaenius formosensis Lorenz, female103 Chlaenius leishanensis Kirschenhofer, male104 Chlaenius leishanensis Kirschenhofer, female105 Chlaenius noguchii Bates, male106 Chlaenius noguchii Bates, female107 Chlaenius wrasei Kirschenhofer, male108 Chlaenius wrasei Kirschenhofer, female109 Chlaenius agiloides Jedlička, male110 Chlaenius agiloides Jedlička, female111 Chlaenius rambouseki Lutshnik, from Ussuri region, male112 Chlaenius rambouseki Lutshnik from Ussuri region, female113 Chlaenius anchomenoides Bates, male, paratype114 Chlaenius nuristanus Jedlička, male, paratype115 Chlaenius nuristanus Jedlička, female, paratype116 Chlaenius formosanus Jedlička from Taiwan, male. Scale line = 1.0 mm.  Figures 117–130. Aedeagi of Chlaenius (Lithochlaenius) species117 Chlaenius chuanqianensis sp. n., holotype (dorsal view)118 Chlaenius chuanqianensis sp. n., holotype (left lateral view)119 Chlaenius linwensini sp. n., holotype (dorsal view)120 Chlaenius linwensini sp. n., holotype (left lateral view)121 Chlaenius propeagilis sp. n., holotype (dorsal view)122 Chlaenius propeagilis sp. n., holotype (left lateral view)123 Chlaenius agilis Chaudoir, holotype (dorsal view)124 Chlaenius agilis Chaudoir, holotype (left lateral view)125 Chlaenius formosensis Lorenz, holotype (dorsal view)126 Chlaenius formosensis Lorenz, holotype (left lateral view)127 Chlaenius leishanensis Kirschenhofer (dorsal view)128 Chlaenius leishanensis Kirschenhofer (left lateral view)129 Chlaenius noguchiiBates (dorsal view)130 Chlaenius noguchiiBates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130).  Figures 158. Map showing the known geographical distributions of Chlaenius (Lithochlaenius)spp.