Calamagrostis heterophylla

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Sylvester S, Soreng R, Bravo-Pedraza W, Cuta-Alarcon L, Giraldo-Cañas D, Aguilar-Cano J, Peterson P (2019) Páramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in páramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia. PhytoKeys 122 : 29–78, doi. Versioned wiki page: 2019-05-28, version 176649, https://species-id.net/w/index.php?title=Calamagrostis_heterophylla&oldid=176649 , contributors (alphabetical order): Pensoft Publishers.

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BibTeX:

@article{Sylvester2019PhytoKeys122,
author = {Sylvester, Steven P. AND Soreng, Robert J. AND Bravo-Pedraza, William J. AND Cuta-Alarcon, Lia E. AND Giraldo-Cañas, Diego AND Aguilar-Cano, Jose AND Peterson, Paul M.},
journal = {PhytoKeys},
publisher = {Pensoft Publishers},
title = {Páramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in páramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia},
year = {2019},
volume = {122},
issue = {},
pages = {29--78},
doi = {10.3897/phytokeys.122.33032},
url = {https://phytokeys.pensoft.net/articles.php?id=33032},
note = {Versioned wiki page: 2019-05-28, version 176649, https://species-id.net/w/index.php?title=Calamagrostis_heterophylla&oldid=176649 , contributors (alphabetical order): Pensoft Publishers.}

}

RIS/ Endnote:

TY - JOUR
T1 - Páramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in páramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia
A1 - Sylvester S
A1 - Soreng R
A1 - Bravo-Pedraza W
A1 - Cuta-Alarcon L
A1 - Giraldo-Cañas D
A1 - Aguilar-Cano J
A1 - Peterson P
Y1 - 2019
JF - PhytoKeys
JA -
VL - 122
IS -
UR - http://dx.doi.org/10.3897/phytokeys.122.33032
SP - 29
EP - 78
PB - Pensoft Publishers
M1 - Versioned wiki page: 2019-05-28, version 176649, https://species-id.net/w/index.php?title=Calamagrostis_heterophylla&oldid=176649 , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/phytokeys.122.33032

Wikipedia/ Citizendium:

<ref name="Sylvester2019PhytoKeys122">{{Citation
| author = Sylvester S, Soreng R, Bravo-Pedraza W, Cuta-Alarcon L, Giraldo-Cañas D, Aguilar-Cano J, Peterson P
| title = Páramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in páramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia
| journal = PhytoKeys
| year = 2019
| volume = 122
| issue =
| pages = 29--78
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/phytokeys.122.33032
| url = https://phytokeys.pensoft.net/articles.php?id=33032
| pmc =
| accessdate = 2024-12-23

}} Versioned wiki page: 2019-05-28, version 176649, https://species-id.net/w/index.php?title=Calamagrostis_heterophylla&oldid=176649 , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Taxonavigation

Ordo: Poales
Familia: Poaceae
Genus: Calamagrostis

Name

Calamagrostis heterophylla (Wedd.) Pilg., Bot. Jahrb. Syst. 42: 64. 1908. Deyeuxia heterophylla Wedd., Bull. Soc. Bot. France 22: 177, 180. 1875. – Wikispecies link – Pensoft Profile

  • [[ | ]] Type. Bolivia. [Potosí], A. D’Orbigny 202 (lectotype, designated by Rúgolo de Agrasar (2012[1]: 201): P (P00729766 [image!]); isolectotypes: BAA (BAA00001845 [image!]), US (US00133531!), W (W18890120011 [image!])).
  • [[ | ]] = Chaetotropisandina Ball, J. Linn. Soc. Bot. 22: 58. 1885. Type: Peru. Ex Saxosis Andium Peruviae juxta Paquim Chicla, 12000–13000’ s.m. [3658–3962 m alt.], 21–23 Apr. 1882, J. Ball s.n. (holotype: P; isotypes: BAA (BAA00001704 [image!]) fragm., K, US (US00344798!) fragm. ex K & ex LE).
  • [[ | ]] = Calamagrostisheterophylla(Wedd.)Pilg.var.robustior Pilg., Bot. Jahrb. Syst.: 64. l908. Type: Peru. Puno: Azangaro, in saxosis calcareis, 4000 m alt., Feb. 1902, A. Weberbauer 474 (holotype: B (not found); isotype: US (US00153711!)).
  • [[ | ]] = Calamagrostisheterophylla(Wedd.)Pilg.var.pubescens Pilg., Bot. Jahrb. Syst.: 64. 1908. Type: Peru. Puno: in provincia Sandia, supra Cuyocuyo, in campis fructisibus nonnullis intermistix, 3700–3800 m alt., May 1902, A. Weberbauer 905 (holotype: B (not found); isotype: US (US00153712!)).
  • [[ | ]] = Calamagrostiscalvescens Pilg., Bot. Jahrb. Syst. 42: 65. 1908. Type: Peru. Ancash: Prov. Cajatambo, 3000-3300 m alt., 13 Apr. 1902, A. Weberbauer 2842 (lectotype, designated by Nicora and Rúgolo de Agrasar (1998[2]: 168): BAA (BAA00000758 [image!]) fragm. ex B; isolectotypes: MOL, US (US00131526!)).
  • [[ | ]] = Calamagrostismulleri Luces, Bol. Soc. Venez., Cienc. Nat. 15 (80):9. 1953. Type: Venezuela. Edo. Mérida: Páramo de Mucuchíes, 4000 m alt., 11 Nov. 1939, A.S. Müller 897 (holotype: VEN (VEN20682 [image!]); isotypes: MO, US (US00149289!)).
  • [[ | ]] = Calamagrostismacbridei Tovar, Mem. Mus. Javier Prado 11:63. 1960. Type: Perú. Pasco, Huarón, northern part Cerro Pasco, northeastern slope, 14000 ft. [4267 m] alt., 12 Jun. 1922, J.F. Macbride & W. Featherstone 998 (holotype: US (US00153707!); isotypes: F (F0040679F [image!]), SI (SI000619 [image!]) fragm.).
  • [[ | ]] = Calamagrostisswallenii Tovar, Mem. Mus. Hist. Nat. “Javier Prado” 11: 66. 1960. Deyeuxiaswallenii (Tovar) Rúgolo, Rev. ~Deyeuxia~ Bolivien 128. 1995. Type: Peru. Huancavelica: Prov. Huancavelica: Tausiri, cerca a Manta, pajonal de Puna, 4500 m alt., 31 Mar. 1953, O. Tovar 1168 (holotype: US (US00133195!); isotypes: GH (GH00023323 [image!]), K (K000308446!), MO (MO115821 [image!]), USM (USM000722 [image!])).

Comments

Previously known from high Andean regions of Venezuela (Hokche et al. 2008[3]; Bono 2010[4]; Briceño 2010[5]), Peru (Tovar 1993[6]), Bolivia (Villavicencio 1995[7], 1998[8]), northern Chile and northwest Argentina (Rúgolo de Agrasar 2012[1]). The specimens collected exhibit characters of both Calamagrostisheterophylla and Calamagrostisbrevipaleata Swallen, an Ecuadorian endemic, with both species having heteromorphic leaf blades, the cauline glabrous and those of the innovations pilose. The species bear more affinity to C.heterophylla as the lemmas measure less than 4.2 mm (> 5 mm long in C.brevipaleata) and the leaf blades measure < 10 cm long (10–25 cm long in C.brevipaleata). However, the specimens do have characteristics of C.brevipaleata in terms of the lemma surfaces, which are smooth proximally and scabrous distally and no great differentiation in width between the cauline and tiller leaf blades.

Specimen examined

COLOMBIA. Boyacá: Mun. Chiscas, páramo El Penon, borde de bosque de Polylepis creciendo sobre roca, 6°36.0714'N; 72°26.229'W, 3917 m alt., 5 Mar. 2018, S.P. Sylvester, R.J. Soreng, W. Bravo & L.E. Cuta 3158 (COL, FMB, K, US).

Taxon Treatment

  • Sylvester, S; Soreng, R; Bravo-Pedraza, W; Cuta-Alarcon, L; Giraldo-Cañas, D; Aguilar-Cano, J; Peterson, P; 2019: Páramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in páramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia PhytoKeys, 122: 29-78. doi

Images

Figure 5. A general habit and a spikelet of Calamagrostisguamanensis Escalona, J.R.I. Wood 4064 (FMB), Calamagrostisheterophylla (Wedd.) Pilg., S.P. Sylvester 3158 (US), Calamagrostispisinna Swallen, S.P. Sylvester 3107 (US), and Calamagrostisrigida (Kunth) Trin. ex Steud., S.P. Sylvester 3125 (US). Habit scale bar 5 cm; spikelet scale bar 1 mm.
Figure 5. A general habit and a spikelet of Calamagrostisguamanensis Escalona, J.R.I. Wood 4064 (FMB), Calamagrostisheterophylla (Wedd.) Pilg., S.P. Sylvester 3158 (US), Calamagrostispisinna Swallen, S.P. Sylvester 3107 (US), and Calamagrostisrigida (Kunth) Trin. ex Steud., S.P. Sylvester 3125 (US). Habit scale bar 5 cm; spikelet scale bar 1 mm. 

Other References

  1. ↑ 1.0 1.1 Rúgolo de Agrasar Z (2012) Deyeuxia Clarion ex P. Beauv. In: Zuloaga F Rúgolo de Agrasar Z Antón A (Eds) Flora Argentina.Poaceae: Pooideae Volumen 3 Tomo 2. IBODA-IMBIV, CONICET, Buenos Aires, Argentina, 180–218.
  2. ↑ Nicora E, Rúgolo de Agrasar Z (1998) Tipos de Gramineae conservados en el Herbario Gaspar Xuarez, Buenos Aires (BAA) procedentes de Berlin (B).Darwiniana36: 163–199.
  3. ↑ Hokche O, Berry P, Huber O (Eds) (2008) Nuevo Catálogo de la Flora Vascular de Venezuela. Fundación Instituto Botánico de Venezuela, Caracas, 1–859.
  4. ↑ Bono G (2010) Poaceae. In: Bono G (Ed.) Catálogo de la Flora y Vegetación de los Valles de la Vertiente Occidental de los Andes de la Cordillera de Mérida.Universita degle Studi di Firenze, Firenze, 127–154.
  5. ↑ Briceño B (2010) Capitulo XXI Familia Poaceae (R.Br.) Barnhart (Gramineae). In: Morillo G Briceño B Silva J (Eds) Botánica y Ecología de las Monocotiledóneas de los Páramos en Venezuela, vol.2. Centro Editorial Litoramma C. A., Merida, 599–711.
  6. ↑ Tovar O (1993) Las Gramíneas (Poaceae) del Perú.Ruizia13: 1–480.
  7. ↑ Villavicencio L (1995) Revision der Gattung Deyeuxia in Bolivien: eine taxonomisch-anatomische Studie der in Bolivien auftretenden Arten der Gattung Deyeuxia. Ph.D. Thesis, Freien Universität Berlin, Berlin, 1–304.
  8. ↑ Villavicencio L (1998) Deyeuxia Clarion ex P. Beauv. In: Renvoize S (Ed.) , Gramineas de Bolivia.Royal Botanic Gardens Kew, 181–235.
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