Bolboceras inaequale

From Species-ID
Jump to: navigation, search
Notice: This page is derived from the original publication listed below, whose author(s) should always be credited. Further contributors may edit and improve the content of this page and, consequently, need to be credited as well (see page history). Any assessment of factual correctness requires a careful review of the original article as well as of subsequent contributions.

If you are uncertain whether your planned contribution is correct or not, we suggest that you use the associated discussion page instead of editing the page directly.

This page should be cited as follows (rationale):
Krikken J (2013) Distinguishing Bolboceras inaequale Westwood, 1848 and two new relatives from India (Coleoptera, Geotrupidae, Bolboceratinae). ZooKeys 276 : 55–66, doi. Versioned wiki page: 2013-03-08, version 31870, https://species-id.net/w/index.php?title=Bolboceras_inaequale&oldid=31870 , contributors (alphabetical order): Pensoft Publishers.

Citation formats to copy and paste

BibTeX:

@article{Krikken2013ZooKeys276,
author = {Krikken, Jan},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {Distinguishing Bolboceras inaequale Westwood, 1848 and two new relatives from India (Coleoptera, Geotrupidae, Bolboceratinae)},
year = {2013},
volume = {276},
issue = {},
pages = {55--66},
doi = {10.3897/zookeys.276.4786},
url = {http://www.pensoft.net/journals/zookeys/article/4786/abstract},
note = {Versioned wiki page: 2013-03-08, version 31870, https://species-id.net/w/index.php?title=Bolboceras_inaequale&oldid=31870 , contributors (alphabetical order): Pensoft Publishers.}

}

RIS/ Endnote:

TY - JOUR
T1 - Distinguishing Bolboceras inaequale Westwood, 1848 and two new relatives from India (Coleoptera, Geotrupidae, Bolboceratinae)
A1 - Krikken J
Y1 - 2013
JF - ZooKeys
JA -
VL - 276
IS -
UR - http://dx.doi.org/10.3897/zookeys.276.4786
SP - 55
EP - 66
PB - Pensoft Publishers
M1 - Versioned wiki page: 2013-03-08, version 31870, https://species-id.net/w/index.php?title=Bolboceras_inaequale&oldid=31870 , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/zookeys.276.4786

Wikipedia/ Citizendium:

<ref name="Krikken2013ZooKeys276">{{Citation
| author = Krikken J
| title = Distinguishing Bolboceras inaequale Westwood, 1848 and two new relatives from India (Coleoptera, Geotrupidae, Bolboceratinae)
| journal = ZooKeys
| year = 2013
| volume = 276
| issue =
| pages = 55--66
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.276.4786
| url = http://www.pensoft.net/journals/zookeys/article/4786/abstract
| pmc =
| accessdate = 2020-03-06

}} Versioned wiki page: 2013-03-08, version 31870, https://species-id.net/w/index.php?title=Bolboceras_inaequale&oldid=31870 , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Taxonavigation

Ordo: Coleoptera
Familia: Geotrupidae
Genus: Bolboceras

Name

Bolboceras inaequale Westwood, 1848Wikispecies linkPensoft Profile

  • Bolboceras inaequalis Westwood, 1848: 386 (description); 1952: 24 (description), pl. IV: figs 14, 14a.
  • Bolboceras inaequale: Paulian 1941[1]: 11, 33 (records from Sudan, Senegal).
  • Indobolbus inaequalis: Krikken 1984[2]: 34 (recombination in checklist).
  • Bolboceras inaequale: Krikken 2011[3]: 239, 240 (transfer to original combination).

Material examined

Syntype, unsexed, from “Centr Ind \ [J.B.] Hearsey” (handwritten), and two more specimens from Westwood’s collection(OUMNH, see comments below).
Non-type material (specimens without data excluded). Bangladesh: Dhaka: Dacca [Dhaka], ex Pascoe coll., 1 spm., in BMNH. Dinajpur: Dhanjuri, v/1963, Mapelli, 2 spm., in MHNG. vi/1963, Mapelli, 2 spm., in MHNG. Rajshahi: Andharkota, vii/1963, Mapelli, 1 spm., in MHNG.
India: Bihar: Buxar, ex Felsche coll., 1 spm., in SMTD. Chapra, Mackenzie, 6 spm., in BMNH. Pusa, 03/viii/1926, Pillai, at light, 1 spm., in BMNH. Pusa, 1913, Fletcher TB, 1 spm., in BPBM. 19/vi/1911, TMH, 1 spm., in BMNH. 30/vii/1915, Bahadur U, at light, 1 spm., in BMNH. 28/vii/1928, Sarkar SC, 1 spm., in BMNH. 20/vii/1908, TNS, at light, 1 spm., in BMNH. 24/vii/1919, Austin GD, verandah 1-10 PM, 1 spm., in BMNH. 08/vii/1915, at light, 1 spm., in BMNH. 12/vii/1915, at light, 1 spm., in BMNH. 05/vii/1924, Mukerjee, 1 spm., in BMNH. 31/vii/1918, Ghosh, underground, 1 spm., in BMNH. 15/viii/1908, RDD, at light, 1 spm., in BMNH. 20/viii/1919, Austin GD, 1 spm., in BMNH. 08/ix/1912, NML, 1 spm., in BMNH. 27/ix/1915, Bahadur U, at light, 1 spm., in BMNH. 04/ix/1915, Bahadur U, at light, 1 spm., in BMNH. 04/x/1915, Bahadur U, at light, 1 spm., in BMNH. 18/xi/1920, Sarkar SC, 1 spm., in BMNH. Tamil Nadu: Madras [Chennai; to be confirmed], ex Felsche coll., 2 spm., in SMTD. Uttar Pradesh: Allahabad, ex Bowring coll., 1 spm., in BMNH. Sitapur, HGC [Champion HG], 4 spm., in BMNH. West Bengal: Calcutta [Kolkata], 1 spm., in BMNH. Kalkutta [Kolkata], ex Felsche coll., 5 spm., in SMTD.
Bengal: unspecified, Parish HM, 1 spm., in BMNH. unspecified, ex Gillet coll., 1 spm., in IRSNB.
India: “Inde”, unspecified, ex Boucomont coll., 1 spm., in MNHN. unspecified, 1 spm., in USNM.
Total 47 males and females, 33 collection records, and Westwood’s material.

Diagnosis

Ridge (or saddle) separating discomedian and discolateral concavities on pronotum narrow, lowered in relation to surrounding discal surface; discoparamedian tubercles approximated (connected by saddle), their tip subacute; discolateral concavity in major males extending down, onto anterolateral surface. Posterior edge of discomedian concavity usually distinctly W-shaped in major individuals, in minors the pronotal ornamentation may be only just distinct (in oblique view); bottom of concavities more or less sericeous. Interocular ridge low, not reaching paraocular ridge on either side, situated between posterior part of eyes. Anterolateral corners of clypeus usually protuberant (in majors intervening ridge concave in axial view). Scutellum usually closely punctate. Intermesocoxal lobe of metasternum anteriorly with arcuate ridge. Aedeagus (Fig. 13) with acuminate parameres, median apparatus with pair of strongly sclerotized lateral stalks, their tip broad, shortly recurved-hooked. Colour uniformly brown. Body length usually 11-13, some series down to ca 9 mm or less (see next section).

Variation

There are small individuals (minors) from Northeast India and Bangladesh (a male from Dhanjuri in Bangladesh pictured here, ca 9 mm long, Figs 3–4, 10, 18–19), characterized by a minimal set of pronotal protrusions and impressions, but still with distinct traces of the basic pattern – and also with a smaller, but externally quite similar aedeagus. Some intermediates between these minors and the real majors (including the type) were seen. At this moment I find no morphological support for ranking the minors as a separate taxon – informative material from different locations required. Paulian (1941)[1] mentions a length range for Bolboceras inaequale of 7–12 mm.

Sexual dimorphism

Apparently no notable sexual dimorphism.

Comments

Note the recent retransfer of this species from Indobolbus Nikolajev, 1979, to Bolboceras Kirby, 1819, and the ensuing suffix change of the species-group name, following the ICZN (2006)[4] ruling. The specimen of Bolboceras inaequale labelled as the type (OUMNH) cannot be considered the holotype, as there are two more historical specimens having the usual large (pale blue) rhomboid label with “W” (meaning Westwood’s original collection), one of them with this label only. The specimen here qualified as syntype has a curatorial label of the 1960s, indicating its registration as Coleoptera type 516, plus additional post-Westwood labels. The three specimens are considered conspecific, and consequently, at this moment, an explicit lectotype designation appears unnecessary.

Distribution

India (southern occurrences to be confirmed), Bangladesh; northern sub-Saharan Africa (needs confirmation).

Taxon Treatment

  • Krikken, J; 2013: Distinguishing Bolboceras inaequale Westwood, 1848 and two new relatives from India (Coleoptera, Geotrupidae, Bolboceratinae) ZooKeys, 276: 55-66. doi

Other References

  1. 1.0 1.1 Paulian R (1941) Coprophaga Africana: Deuxième note. Revision des Bolboceras africains. Revue de Zoologie et Botanie africaines 35: 1-71.
  2. Krikken J (1984) A generic reclassification of the Afrotropical Bolboceratini (Coleoptera: Geotrupidae). Zoologische Mededelingen Leiden 58: 23-45.
  3. Krikken J (2011) A new Kenyan species of Bolboceras (Coleoptera: Geotrupidae: Bolboceratinae). Koleopterologische Rundschau 81: 237-242.
  4. ICZN (2006) International Commission on Zoological Nomenclature. Opinion 2138 (Case 3097). Bolboceras Kirby, 1819 (July) (Insecta, Coleoptera): not conserved; priority maintained for Odonteus Samouelle, 1819 (June). Bulletin of Zoological Nomenclature 63: 62-64.

Images