Acomoptera

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Kerr P (2011) Six new species of Acomoptera from North America (Diptera, Mycetophilidae). ZooKeys 137 : 41–76, doi. Versioned wiki page: 2011-10-14, version 17510, https://species-id.net/w/index.php?title=Acomoptera&oldid=17510 , contributors (alphabetical order): Pensoft Publishers.

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BibTeX:

@article{Kerr2011ZooKeys137,
author = {Kerr, Peter H.},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {Six new species of Acomoptera from North America (Diptera, Mycetophilidae)},
year = {2011},
volume = {137},
issue = {},
pages = {41--76},
doi = {10.3897/zookeys.137.1764},
url = {http://www.pensoft.net/journals/zookeys/article/1764/abstract},
note = {Versioned wiki page: 2011-10-14, version 17510, https://species-id.net/w/index.php?title=Acomoptera&oldid=17510 , contributors (alphabetical order): Pensoft Publishers.}

}

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TY - JOUR
T1 - Six new species of Acomoptera from North America (Diptera, Mycetophilidae)
A1 - Kerr P
Y1 - 2011
JF - ZooKeys
JA -
VL - 137
IS -
UR - http://dx.doi.org/10.3897/zookeys.137.1764
SP - 41
EP - 76
PB - Pensoft Publishers
M1 - Versioned wiki page: 2011-10-14, version 17510, https://species-id.net/w/index.php?title=Acomoptera&oldid=17510 , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/zookeys.137.1764

Wikipedia/ Citizendium:

<ref name="Kerr2011ZooKeys137">{{Citation
| author = Kerr P
| title = Six new species of Acomoptera from North America (Diptera, Mycetophilidae)
| journal = ZooKeys
| year = 2011
| volume = 137
| issue =
| pages = 41--76
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.137.1764
| url = http://www.pensoft.net/journals/zookeys/article/1764/abstract
| pmc =
| accessdate = 2024-12-13

}} Versioned wiki page: 2011-10-14, version 17510, https://species-id.net/w/index.php?title=Acomoptera&oldid=17510 , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Taxonavigation

Ordo: Diptera
Familia: Mycetophilidae

Name

Acomoptera Vockeroth – Wikispecies link – Pensoft Profile

  • Acomoptera Vockeroth, 1980: 534

Type species

Eudicrana plexipus Garrett, 1925: 4, by subsequent designation.

Diagnosis

This genus may be distinguished from most mycetophilid genera by the following combination of wing characters: Sc ending in C, sc-r present near middle of Sc, and R4 present and displaced from Rs by more than 2.5× its own length. Acomoptera may be distinguished from Paratinia by having wing membrane bare and from Drepanocercus by having the cubital fork near the level or distad of sc-r. Acomoptera is distinguished from Phoenikiella Chandler by having a setose Sc, with sc-r positioned near the middle of this vein.

Description

Body length 4.5 to 7.1 mm. Ocelli three, subequal or median ocellus reduced, lateral ocellus separated from eye margin by approximately its own diameter or clearly less. Eye with a broad distinct emargination above antennal base, with microsetae usually scattered and short, but sometimes more numerous and longer. Frons bare between ocelli and antennal bases. Frontal tubercle present. Antenna two to four times as long as thorax, 2/3rds to approximately same length as abdomen; antennal bases nearly touching, separated only by narrow tip of frontal tubercle; all flagellomeres densely setulose, elongate, flagellomere length decreases gradually toward tip. Palpus with five palpomeres; palpomere 1 short, as wide as long, remaining palpomeres longer than wide (except sometimes palpomere 2 as wide as long); palpomere 2 clearly shorter than or subequal to palpomere 3; palpomere 4 three to six times longer than wide; palpomere 5 thinner than others, six to eleven times longer than wide, usually subequal to combined length of palpomeres 3 and 4. Scutum with short appressed acrostichal and dorsocentral setae and longer and more erect sublateral setae, the intervening areas bare. Scutellum with three or four irregular rows of short to long setae, sometimes bare medially. Paratergite present; antepronotum with setae on posterior half, proepisternum bare; mesopleuron, metapleuron, and prosternum bare; metanotum with one to three erect setae laterad of base of halter; mediotergite and laterotergite bare; phragma not well developed. Wing unmarked, with dense microtrichia, without macrotrichia. Costal vein extends beyond R5, between approx. 0.25× and 0.33× distance between R5 and M1; sc-r crossvein present, slightly distad of middle of Sc, proximal of Rs; R4 present (when missing, teratogenic), located approx. 3× its own length from Rs, forming an elongate radial cell; medial fork approx. 3× longer than stem; cubital fork arises near level or distad of sc-r and proximad of medial fork, cubital stem shorter than CuA1 and approx. equal in length to or longer than CuA2; all cross veins bare on upper surface, all longitudinal veins setose on upper surface, except CuP and the first two sections of M bare (as an exception, Acomoptera crispa sp. n. may have a few setae on M1+2); wing veins R1, R5, M1, and sometimes M2 with at least some setae on lower surface. Hind coxa with a single vertical row of setae on at least proximal half (sometimes weak). Tibial bristles short but distinct, the longest between half and full tibial diameter in length. Anteroapical depressed area of fore tibia ovate and well developed. Tibial spur pairs of equal length, hind tibial spurs usually longer than mid tibial spurs, but sometimes mid and hind spurs subequal in length. Tarsal claws each with one or two small ventral teeth. Empodia small. Sternite 1 bare. Sternites 2 to 7 each with a pair of broad, poorly defined, submedian to sub lateral fold-lines. Terminalia not rotated. Epandrium (tergite 9) between three times as wide as long to slightly wider than long. Hypandrium (sternite 9) fused with gonocoxites, the synsclerite with or without a narrow transverse membranous area across ventral surface, with or without a ventral preapical hook-like process directed posteriorly. Gonostylus variously formed, often divided into two or three lobes. Posterodorsal process attached to the median dorsal angles of the gonocoxites variously shaped, sometimes arising as a distinctive bilobed wing-like structure whose posterior margins are darkened and toothed. Cerci broad, flat, broadly rounded apically, with fine setae. Hypoproct broad, semicircular, weakened anteromedially.
Female sternite 8 deeply emarginate posteriorly, the rest of the median area membranous or lightly sclerotized. Sternite 9 with two weakly sclerotized anterolateral areas and two slender, more heavily sclerotized submedian processes projecting posteriorly. Tergite 10 short, setose, fused on either side with sternite 10. Sternite 10 well developed, membranous medially, tapering on posterior half, posterior margin extending to apex of first segment of cercus. Cercus 2-segmented, first segment almost twice as long as wide, second segment oval.

Key to the Acomoptera of the world (males)

Key to the Acomoptera of the world (males) — P. Kerr
1
Hook-like or fork-like hypandrial lobe present (Figs 5B, 5E, 17B, 17E, 23B, 23E) 
  2
–
Hypandrial lobe absent 
  4
2
Hypandrial lobe fork-like (Figs 16D, 17A–C) 
  Acomoptera forculata sp. n.
–
Hypandrial lobe hook-like (Figs 5B, 23B) 
  3
3
Hypandrial lobe long, subtending the gonostyli in part (Fig. 5E); gonostylus with club-like, ventral process that extends well beyond rest of gonostyus posteriorly (Figs 4A–C, 5A–B, 5E); Europe 
  Acomoptera difficilis Dziedzicki
–
Hypandrial lobe short, not reaching beyond posterior margin of gonocoxites (Figs 22B, 23B, 23E); gonostylus without large, posteriorly-directed club-like process; North America 
  Acomoptera plexipus Garrett
4
Gonostylus with three apical lobes of similar size and shape 
  Acomoptera sinicaWu & Yang
–
Gonostylus variously shaped, without three subequal lobes 
  5
5
Gonocoxites with well-developed, paired structure (‘gonocoxal comb') dorsomedially (Figs 2A, 13A, 19A, 25A) 
  6
–
Gonocoxites without such well-developed dorsomedial structure (e.g., Fig. 11A) 
  9
6
Posterior face of gonostyli with prominent cluster of curved setae dorsally (Figs 3A–B, 3D–E, 26A–B, 26D–E) 
  7
–
Posterior face of gonostyli without such curved setae (Figs 14A–B, 14D–E, 20A–B, 20D–E) 
  8
7
Gonocoxites with transverse, denticulate ridge of cuticle ventrally (Fig. 26B) ; outermost dorsal margin of gonostylus swept forward (Fig. 26D–E); ventromedial process of gonostylus usually unforked (Fig. 26E) 
  Acomoptera vockerothi sp. n.
–
Gonocoxites smooth ventrally, without transverse, denticulate ridge (Fig. 3B) ; uppermost lateral margin of gonostylus swept forward (Fig. 3D–E); ventromedial process usually forked (Fig. 3E) 
  Acomoptera crispa sp. n.
8
Inner face of gonostylus toothed dorsally (Fig. 20A) 
  Acomoptera nelsoni sp. n.
–
Inner face of gonostylus smooth dorsally (Fig. 14A) 
  Acomoptera echinosa sp. n.
9
Fork of cubital wing vein distad of base of r-m (Fig. 8); hypandrium entire (Fig. 11B) 
  Acomoptera digitata sp. n.
–
Cubital fork proximal to r-m (Ševčík 2004[1]: fig. 1, Søli, 1993: fig. 1); hypandrium with unsclerotized area near posterior margin (Søli, 1993: fig. 2B) 
  Acomoptera spinistyla (Søli), comb. n.

Taxon Treatment

  • Kerr, P; 2011: Six new species of Acomoptera from North America (Diptera, Mycetophilidae) ZooKeys, 137: 41-76. doi

Other References

  1. ↑ Ševčík J (2004) New data on Sciaroidea (Diptera) from the Czech and Slovak Republics, with descriptions of seven new species of Mycetophilidae. Časopis Slezského zemského muzea. Série A, Vědy přírodní 53: 49-74.

Images

Figure 2. Acomoptera crispa sp. n., male genitalia, images: A dorsal view [691235] B ventral view [691236] C lateral view [691237] D posterior view [691237]. Scale line = 0.1 mm. Abbreviations: cerc cercus epandepandrium gcx comb gonocoxal comb goncx gonocoxites gonst gonostylus v m gonst proc ventromedial gonostylus process.
'Figure 2. Acomoptera crispa sp. n., male genitalia, images: A dorsal view [691235] B ventral view [691236] C lateral view [691237] D posterior view [691237]. Scale line = 0.1 mm. Abbreviations: cerc cercus epandepandrium gcx comb gonocoxal comb goncx gonocoxites gonst gonostylus v m gonst proc ventromedial gonostylus process. 
Figure 3. Acomoptera crispa sp. n., male genitalia, illustrated: A gonopods, dorsal view [691241] B gonopods, ventral view [691242] C gonocoxites, with gonostyli removed, posterior view [691244] D gonostyli, posterior view [691244] E gonopods, lateral view [691243] F epandrium, dorsal view [691240]. Scale line = 0.1 mm. Abbreviations: cerc cercus epand epandrium gcx comb gonocoxal comb goncx gonocoxites gonst gonostylus v m gonst proc ventromedial gonostylus process.
Figure 3. Acomoptera crispa sp. n., male genitalia, illustrated: A gonopods, dorsal view [691241] B gonopods, ventral view [691242] C gonocoxites, with gonostyli removed, posterior view [691244] D gonostyli, posterior view [691244] E gonopods, lateral view [691243] F epandrium, dorsal view [691240]. Scale line = 0.1 mm. Abbreviations: cerc cercus epand epandrium gcx comb gonocoxal comb goncx gonocoxites gonst gonostylus v m gonst proc ventromedial gonostylus process. 
Figure 4. Acomoptera difficilis, male genitalia, images: A dorsal view [691256] B ventral view [691257] C lateral view [691258] D posterior view [691259]. Scale line = 0.1 mm.
Figure 4. Acomoptera difficilis, male genitalia, images: A dorsal view [691256] B ventral view [691257] C lateral view [691258] D posterior view [691259]. Scale line = 0.1 mm. 
Figure 5. Acomoptera difficilis, male genitalia, illustrated: A gonopods, dorsal view [691261] B gonopods, ventral view [691262] C gonocoxites, with gonostyli removed, posterior view [691264] D gonostyli, posterior view [691265] E gonopods, lateral view [691263] F epandrium, dorsal view [691263]. Scale line = 0.1 mm. Abbreviations: gonst gonostylus hyp lobe hypandrial lobe.
Figure 5. Acomoptera difficilis, male genitalia, illustrated: A gonopods, dorsal view [691261] B gonopods, ventral view [691262] C gonocoxites, with gonostyli removed, posterior view [691264] D gonostyli, posterior view [691265] E gonopods, lateral view [691263] F epandrium, dorsal view [691263]. Scale line = 0.1 mm. Abbreviations: gonst gonostylus hyp lobe hypandrial lobe. 
' Figure 8.' Acomoptera digitata sp. n., wing, dorsal view [692360]. Scale line = 1 mm.
' Figure 8. Acomoptera digitata sp. n., wing, dorsal view [692360]. Scale line = 1 mm. 
Figure 11. Acomoptera digitata sp. n., male genitalia, illustrated: A gonopods, dorsal view B gonopods, ventral view C gonocoxites, with gonostyli removed, posterior view D gonostyli, posterior view E gonopods, lateral view F epandrium, dorsal view. Scale line = 0.1 mm.
Figure 11. Acomoptera digitata sp. n., male genitalia, illustrated: A gonopods, dorsal view B gonopods, ventral view C gonocoxites, with gonostyli removed, posterior view D gonostyli, posterior view E gonopods, lateral view F epandrium, dorsal view. Scale line = 0.1 mm. 
Figure 13. Acomoptera echinosa sp. n., male genitalia, images: A dorsal view [691278] B ventral view [691279] C lateral view [691280] D posterior view [691281]. Scale line = 0.1 mm.
Figure 13. Acomoptera echinosa sp. n., male genitalia, images: A dorsal view [691278] B ventral view [691279] C lateral view [691280] D posterior view [691281]. Scale line = 0.1 mm. 
Figure 14. Acomoptera echinosa sp. n., male genitalia, illustrated: A gonopods, dorsal view [691284] B gonopods, ventral view [691285] C gonocoxites, with gonostyli removed, posterior view D gonostyli, posterior view [691287] E gonopods, lateral view [691286] F epandrium, dorsal view [691283]. Scale line = 0.1 mm.
Figure 14. Acomoptera echinosa sp. n., male genitalia, illustrated: A gonopods, dorsal view [691284] B gonopods, ventral view [691285] C gonocoxites, with gonostyli removed, posterior view D gonostyli, posterior view [691287] E gonopods, lateral view [691286] F epandrium, dorsal view [691283]. Scale line = 0.1 mm. 
Figure 16. Acomoptera forculata sp. n., male genitalia, images: A dorsal view [691288] B ventral view [691289] C lateral view [691290] D posterior view [691291]. Scale line = 0.1 mm. Abbreviations: hyp lobe hypandrial lobe.
Figure 16. Acomoptera forculata sp. n., male genitalia, images: A dorsal view [691288] B ventral view [691289] C lateral view [691290] D posterior view [691291]. Scale line = 0.1 mm. Abbreviations: hyp lobe hypandrial lobe. 
Figure 17. Acomoptera forculata sp. n., male genitalia, illustrated: A gonopods, dorsal view [691294] B gonopods, ventral view [691295] C gonocoxites, with gonostyli removed, posterior view [691297] D gonostyli, posterior view [691298] E gonopods, lateral view [691296] F epandrium, dorsal view [691293]. Scale line = 0.1 mm.
Figure 17. Acomoptera forculata sp. n., male genitalia, illustrated: A gonopods, dorsal view [691294] B gonopods, ventral view [691295] C gonocoxites, with gonostyli removed, posterior view [691297] D gonostyli, posterior view [691298] E gonopods, lateral view [691296] F epandrium, dorsal view [691293]. Scale line = 0.1 mm. 
Figure 19. Acomoptera nelsoni sp. n., male genitalia, images: A dorsal view [691299] B ventral view [691300] C lateral view [691301] D posterior view [691302]. Scale line = 0.1 mm.
Figure 19. Acomoptera nelsoni sp. n., male genitalia, images: A dorsal view [691299] B ventral view [691300] C lateral view [691301] D posterior view [691302]. Scale line = 0.1 mm. 
Figure 20. Acomoptera nelsoni sp. n., male genitalia, illustrated: A gonopods, dorsal view [691305] B gonopods, ventral view [691306] C gonocoxites, with gonostyli removed, posterior view [691308] D gonostyli, posterior view [691309] E gonopods, lateral view [691307] F epandrium, dorsal view [691304]. Scale line = 0.1 mm.
Figure 20. Acomoptera nelsoni sp. n., male genitalia, illustrated: A gonopods, dorsal view [691305] B gonopods, ventral view [691306] C gonocoxites, with gonostyli removed, posterior view [691308] D gonostyli, posterior view [691309] E gonopods, lateral view [691307] F epandrium, dorsal view [691304]. Scale line = 0.1 mm. 
Figure 22. Acomoptera plexipus, male genitalia, images: A dorsal view [691311] B ventral view [691312] C lateral view [691313] D posterior view [691314]. Scale line = 0.1 mm.
Figure 22. Acomoptera plexipus, male genitalia, images: A dorsal view [691311] B ventral view [691312] C lateral view [691313] D posterior view [691314]. Scale line = 0.1 mm. 
Figure 23. Acomoptera plexipus, male genitalia, illustrated: A gonopods, dorsal view [691317] B gonopods, ventral view [691318] C gonocoxites, with gonostyli removed, posterior view [691320] D gonostyli, posterior view [691321] E gonopods, lateral view [691319] F epandrium, dorsal view [691316]. Scale line = 0.1 mm.
Figure 23. Acomoptera plexipus, male genitalia, illustrated: A gonopods, dorsal view [691317] B gonopods, ventral view [691318] C gonocoxites, with gonostyli removed, posterior view [691320] D gonostyli, posterior view [691321] E gonopods, lateral view [691319] F epandrium, dorsal view [691316]. Scale line = 0.1 mm. 
Figure 25. Acomoptera vockerothi sp. n., male genitalia, images: A dorsal view [691246] B ventral view [691247] C lateral view [691248] D posterior view [691249]. Scale line = 0.1 mm.
Figure 25. Acomoptera vockerothi sp. n., male genitalia, images: A dorsal view [691246] B ventral view [691247] C lateral view [691248] D posterior view [691249]. Scale line = 0.1 mm. 
Figure 26. Acomoptera vockerothi sp. n., male genitalia, illustrated: A gonopods, dorsal view [691252] B gonopods, ventral view [691253] C gonocoxites, with gonostyli removed, posterior view D gonostyli, posterior view [691255] E gonopods, lateral view [691254] F epandrium, dorsal view [691251]. Scale line = 0.1 mm.
Figure 26. Acomoptera vockerothi sp. n., male genitalia, illustrated: A gonopods, dorsal view [691252] B gonopods, ventral view [691253] C gonocoxites, with gonostyli removed, posterior view D gonostyli, posterior view [691255] E gonopods, lateral view [691254] F epandrium, dorsal view [691251]. Scale line = 0.1 mm. 
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