View source for Madagascarchaea\according to Wood et al 2017


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{{Publication to wiki notice
 | author = Wood, Hannah M. AND Scharff, Nikolaj
 | author_abbreviated = Wood H AND Scharff N
 | year = 2017
 | title = A review of the Madagascan pelican spiders of the genera ''Eriauchenius'' O. Pickard-Cambridge, 1881 and ''Madagascarchaea'' gen. n. (Araneae, Archaeidae)
 | journal = ZooKeys
 | volume = 
 | issue = 727
 | pages = 1--96
 | doi = 10.3897/zookeys.727.20222
 | citationurl = https://zookeys.pensoft.net/articles.php?id=20222&element_type=9 
 | url = https://zookeys.pensoft.net/articles.php?id=20222
 | publisher = Pensoft Publishers
 | publisherurl = http://www.pensoft.net/
}}

{{Treatment start
 | Ordo = Araneae
 | Familia = Archaeidae
 | Genus = Madagascarchaea
 | Specific name = 
 | Infraspecific name = 
 | Taxon rank = 	
 | Taxon authority = Wood & Scharff, 2017
 | Taxon status = gen. n.
 | Wikispecies page name = Madagascarchaea
 | ZooBank ID = 041CF907-A633-440F-A0DE-EA875E1265DF
 | Pensoft Profile = Madagascarchaea
}}

==Type species==
''{{Taxon name|Archaea gracilicollis}}'' Millot, 1948

==Etymology==
The name refers to the Madagascan distribution and is feminine in gender.

==Diagnosis==
Distinguished from all other archaeids by the presence of six protrusions, each with a small spine, on the crown of the cephalon, and by the presence of a retrolateral apophysis on the male pedipalpal patella.

==Description==
See Wood (2008)<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref > for a complete description.

==Included species==
14 described species ''{{Taxon name|M. ambre}}'' (Wood, 2008), ''{{Taxon name|M. anabohazo}}'' (Wood, 2008), ''{{Taxon name|M. borimontsina}}'' (Wood, 2008), ''{{Taxon name|M. gracilicollis}}'' (Millot, 1948), ''{{Taxon name|M. griswoldi}}'' (Wood, 2008), ''{{Taxon name|M. halambohitra}}'' (Wood, 2008), ''{{Taxon name|M. jeanneli}}'' (Millot, 1948), ''{{Taxon name|M. lavatenda}}'' (Wood, 2008), ''{{Taxon name|M. legendrei}}'' (Platnick, 1991), ''{{Taxon name|M. namoroka}}'' (Wood, 2008), ''{{Taxon name|M. spiceri}}'' (Wood, 2008), ''{{Taxon name|M. tsingyensis}}'' (Lotz, 2003), ''{{Taxon name|M. vadoni}}'' (Millot, 1948), ''{{Taxon name|M. voronakely}}'' (Wood, 2008), and 4 new species described here: ''{{Taxon name|Madagascarchaea fohy}}'' sp. n., ''{{Taxon name|Madagascarchaea lotzi}}'' sp. n., ''{{Taxon name|Madagascarchaea moramora}}'' sp. n., ''{{Taxon name|Madagascarchaea rabesahala}}'' sp. n

==Distribution==
Madagascar

==Discussion==
We elevate the “{{Taxon name|gracilicollis}} group” (Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >) to genus level based on phylogenetic analysis that strongly supports two monophyletic clades, ''{{Taxon name|Eriauchenius}}'' and ''{{Taxon name|Madagascarchaea}}'' gen. n., that have diversified on Madagascar (Fig. 1) (Wood et al. 2015<ref name="B27"> {{aut|Wood H}}, {{aut|Gillespie R}}, {{aut|Griswold C}}, {{aut|Wainwright P}} (2015) Why is Madagascar special? The extraordinarily slow evolution of pelican spiders ({{Taxon name|Araneae}}, {{Taxon name|Archaeidae}}). Evolution 69: 462–481. https://doi.org/10.1111/evo.12578</ref >; Wood et al. 2012<ref name="B28"> {{aut|Wood H}}, {{aut|Griswold C}}, {{aut|Gillespie R}} (2012) Phylogenetic placement of pelican spiders ({{Taxon name|Archaeidae}}, {{Taxon name|Araneae}}), with insight into evolution of the “neck” and predatory behaviours of the superfamily {{Taxon name|Palpimanoidea}}. Cladistics 28: 598–626. https://doi.org/10.1111/j.1096-0031.2012.00411.x</ref >). These two genera are not sister clades, but instead ''{{Taxon name|Afrarchaea}}'' is sister to ''{{Taxon name|Eriauchenius}}'' in Wood et al (2015)<ref name="B27"> {{aut|Wood H}}, {{aut|Gillespie R}}, {{aut|Griswold C}}, {{aut|Wainwright P}} (2015) Why is Madagascar special? The extraordinarily slow evolution of pelican spiders ({{Taxon name|Araneae}}, {{Taxon name|Archaeidae}}). Evolution 69: 462–481. https://doi.org/10.1111/evo.12578</ref >, and ''{{Taxon name|Afrarchaea}}'' is sister to ''{{Taxon name|Madagascarchaea}}'' gen. n. in Wood et al (2012)<ref name="B28"> {{aut|Wood H}}, {{aut|Griswold C}}, {{aut|Gillespie R}} (2012) Phylogenetic placement of pelican spiders ({{Taxon name|Archaeidae}}, {{Taxon name|Araneae}}), with insight into evolution of the “neck” and predatory behaviours of the superfamily {{Taxon name|Palpimanoidea}}. Cladistics 28: 598–626. https://doi.org/10.1111/j.1096-0031.2012.00411.x</ref >. ''{{Taxon name|Eriauchenius}}'' and ''{{Taxon name|Afrarchaea}}'' are never recovered as sister clades.<br />
''{{Taxon name|Madagascarchaea}}'' gen. n. contains two main clades, the “{{Taxon name|vadoni}} group” and the “{{Taxon name|jeanneli}} group” (Fig. 1). The “{{Taxon name|vadoni}} group” contains ''{{Taxon name|M. borimontsina}}'', ''{{Taxon name|M. legendrei}}'', ''{{Taxon name|M. vadoni}}'', ''{{Taxon name|M. fohy}}'' sp. n., and ''{{Taxon name|M. rabesahala}}'' sp. n., and these species share the following traits: a retrolateral apophysis on the male pedipalpal femur (compare fig. 12A with fig. 12B in Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >) and a highly reduced FSGP that lacks “wings” (Figs 23B, 24B, 25B). The female genitalia of “{{Taxon name|vadoni}} group” species do not have interspecific variation. Instead, somatic traits are more useful for identifying “{{Taxon name|vadoni}} group” species. We split what was previously considered ''{{Taxon name|M. legendrei}}'' into two species: ''{{Taxon name|M. legendrei}}'' and ''{{Taxon name|M. rabesahala}}'' sp. n. For clarity, we describe both ''{{Taxon name|M. legendrei}}'' and ''{{Taxon name|M. rabesahala}}'' sp. n. here. In addition, there is a single female specimen ([http://data.antweb.org/specimen/CASENT9046596 CASENT9046596]), denoted as “''{{Taxon name|Madagascarchaea}}'' sp. 1” in Fig. 1 that may be a new species, however, since this is the only known specimen and females are difficult to diagnose, we chose not to describe it (collection information: Madagascar, Fianarantsoa, Réserve Forestiére d’Agnalazaha, Mahabo, 42.9 km 215° SW Farafangana, [http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=23%C2%B011%2738%22S%2C+47%C2%B043%2723%22E&coordinates%5B0%5D=23%C2%B011%2738%22S%2C+47%C2%B043%2723%22E 23°11'38"S, 47°43'23"E], 20 m, 19 Apr 2006, littoral rainforest, maxi winkler litter extraction, B.L. Fisher et al.).<br />
''{{Taxon name|M. jeanneli}}'', ''{{Taxon name|M. ambre}}'', ''{{Taxon name|M. lotzi}}'' sp. n., and ''{{Taxon name|M. moramora}}'' sp. n. belong to the “{{Taxon name|jeanneli}} group” and are part of a species complex distinguished from other ''{{Taxon name|Madagascarchaea}}'' by having an elongated and pointy head (Figs 26A, 27A, 28A), a triangular abdomen that is either straight across the posterior (Fig. 28A) or invaginated (Figs 26A, 27A), and a conductor that is a large triangular piece in the prolateral view (Figs 26D, 27D). The current study splits the ''{{Taxon name|M. jeanneli}}'' of Wood (2008)<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref > into several different species: ''{{Taxon name|M. jeanneli}}'', ''{{Taxon name|M. lotzi}}'' sp. n., and ''{{Taxon name|M. moramora}}'' sp. n. For clarity, we also redescribe ''{{Taxon name|M. jeanneli}}'' here. ''{{Taxon name|M. ambre}}'', is considered a part of the “{{Taxon name|jeanneli}}-complex” but is not redescribed here, see Wood (2008)<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref > for a complete description. These four species form a monophyletic group (Fig. 1), although ''{{Taxon name|M. jeanneli}}'' is not included in the phylogeny. Females cannot be distinguished. For the males, embolus shape is the main morphological difference among species.

==''{{Taxon name|Madagascarchaea}}'' gen. n.: to supplement “Gracilicollis group” key==
When using the identification key from Wood (2008)<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >, if a specimen is identified as either ''{{Taxon name|M. jeanneli}}'' or ''{{Taxon name|M. ambre}}'', then use the following key to separate ''{{Taxon name|M. jeanneli}}'', ''{{Taxon name|M. ambre}}'', ''{{Taxon name|M. lotzi}}'' sp. n., and ''{{Taxon name|M. moramora}}'' sp. n.; females of these four species are indistinguishable: 

{{Key Start
| title = ''{{Taxon name|Madagascarchaea}}'' gen. n.: to supplement “Gracilicollis group” key
| description =
| flags = jkey-hidekeymetadata 
| taxon name = 
| common names =
| taxon rank = 
| parent taxon =
| parent key =
| parent key text =
| geoscope = 
| category =
| source =
| audience =
| status =
| creators = H. Wood & N. Scharff
}}		
{{Lead | colspan="1" rowspan="1" | 1 | colspan="1" rowspan="1" | Embolus bifurcation shallow, both parts of bifurcation roughly equal thickness (Fig. 27E–F, G–H, “a” and “p”); posterior of abdomen usually invaginated in both males and females (Fig. 27H, arrow)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. lotzi}}'' '''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Embolus with a deeper bifurcation, both parts of bifurcation different thickness (Figs 26F, I, L, 28E–F, compare shape of “a” with shape of “p”); abdomen invaginated or straight across the posterior in both sexes (Fig. 28A, arrow)
	| colspan="1" rowspan="1" | result text = '''2'''}}
{{Lead | colspan="1" rowspan="1" | 2 | colspan="1" rowspan="1" | Anterior portion of embolus bifurcation thinner than the posterior part, and jutting out past the conductor in the ventral view (see fig. 21 in Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >); usually abdomen straight across the posterior in both sexes (as in Fig. 28A, arrow; also see fig. 1C in Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. ambre}}'' '''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Anterior portion of embolus bifurcation broad, posterior portion not extending past conductor, and posterior portion with a bifurcation (Fig. 26E–F,H–I)
	| colspan="1" rowspan="1" | result text = '''3'''}}
{{Lead | colspan="1" rowspan="1" | 3 | colspan="1" rowspan="1" | Anterior portion of embolus bifurcation thick and blunt at tip (Fig. 28D–F, “a”), posterior portion with a bifurcation at tip with each piece of unequal width (Fig. 28D–F, “p”); posterior part of abdomen usually straight in both sexes (Fig. 28A, arrow)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. moramora}}'' '''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Anterior portion of embolus bifurcation tapering off to a point (Fig. 26D–L, “a”), posterior portion with a bifurcation at the tip (Fig. 26E–F, H–I); posterior part of abdomen usually invaginated in both sexes (Fig. 26A)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. jeanneli}}'' '''}}
{{Lead | colspan="1" rowspan="1" | 1 | colspan="1" rowspan="1" | In males and females, cephalon crown rounded (Figs 23A, 24A)
	| colspan="1" rowspan="1" | result text = '''2'''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Cephalon crown not as round in both sexes (Fig. 25A)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. rabesahala}}'' '''}}
{{Lead | colspan="1" rowspan="1" | 2 | colspan="1" rowspan="1" | In males, pedipalpal bulb, with a sclerotized rod-shaped piece on embolus (Fig. 23H, J, arrow); presence of a cymbium process that has setae on the retrolateral side of the cymbium (Fig. 23C); females of ''{{Taxon name|M. fohy}}'' and ''{{Taxon name|M. legendrei}}'' are indistinguishable
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. fohy}}'' '''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Embolus lacking heavily sclerotized rod-shaped piece (Figs 24G–I, 25G–I); retrolateral process on cymbium present but lacking setae (Fig. 24C)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. legendrei}}'' '''}}
{{Key End}} When using the identification key from Wood (2008)<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >, if a specimen is identified as ''{{Taxon name|M. legendrei}}'', then use the following key to separate species of ''{{Taxon name|M. legendrei}}'', ''{{Taxon name|M. fohy}}'' sp. n., and ''{{Taxon name|M. rabesahala}}'' sp. n.: 

{{Key Start
| title = ''{{Taxon name|Madagascarchaea}}'' gen. n.: to supplement “Gracilicollis group” key
| description =
| flags = jkey-hidekeymetadata 
| taxon name = 
| common names =
| taxon rank = 
| parent taxon =
| parent key =
| parent key text =
| geoscope = 
| category =
| source =
| audience =
| status =
| creators = H. Wood & N. Scharff
}}		
{{Lead | colspan="1" rowspan="1" | 1 | colspan="1" rowspan="1" | Embolus bifurcation shallow, both parts of bifurcation roughly equal thickness (Fig. 27E–F, G–H, “a” and “p”); posterior of abdomen usually invaginated in both males and females (Fig. 27H, arrow)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. lotzi}}'' '''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Embolus with a deeper bifurcation, both parts of bifurcation different thickness (Figs 26F, I, L, 28E–F, compare shape of “a” with shape of “p”); abdomen invaginated or straight across the posterior in both sexes (Fig. 28A, arrow)
	| colspan="1" rowspan="1" | result text = '''2'''}}
{{Lead | colspan="1" rowspan="1" | 2 | colspan="1" rowspan="1" | Anterior portion of embolus bifurcation thinner than the posterior part, and jutting out past the conductor in the ventral view (see fig. 21 in Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >); usually abdomen straight across the posterior in both sexes (as in Fig. 28A, arrow; also see fig. 1C in Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. ambre}}'' '''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Anterior portion of embolus bifurcation broad, posterior portion not extending past conductor, and posterior portion with a bifurcation (Fig. 26E–F,H–I)
	| colspan="1" rowspan="1" | result text = '''3'''}}
{{Lead | colspan="1" rowspan="1" | 3 | colspan="1" rowspan="1" | Anterior portion of embolus bifurcation thick and blunt at tip (Fig. 28D–F, “a”), posterior portion with a bifurcation at tip with each piece of unequal width (Fig. 28D–F, “p”); posterior part of abdomen usually straight in both sexes (Fig. 28A, arrow)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. moramora}}'' '''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Anterior portion of embolus bifurcation tapering off to a point (Fig. 26D–L, “a”), posterior portion with a bifurcation at the tip (Fig. 26E–F, H–I); posterior part of abdomen usually invaginated in both sexes (Fig. 26A)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. jeanneli}}'' '''}}
{{Lead | colspan="1" rowspan="1" | 1 | colspan="1" rowspan="1" | In males and females, cephalon crown rounded (Figs 23A, 24A)
	| colspan="1" rowspan="1" | result text = '''2'''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Cephalon crown not as round in both sexes (Fig. 25A)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. rabesahala}}'' '''}}
{{Lead | colspan="1" rowspan="1" | 2 | colspan="1" rowspan="1" | In males, pedipalpal bulb, with a sclerotized rod-shaped piece on embolus (Fig. 23H, J, arrow); presence of a cymbium process that has setae on the retrolateral side of the cymbium (Fig. 23C); females of ''{{Taxon name|M. fohy}}'' and ''{{Taxon name|M. legendrei}}'' are indistinguishable
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. fohy}}'' '''}}
{{Lead | colspan="1" rowspan="1" | – | colspan="1" rowspan="1" | Embolus lacking heavily sclerotized rod-shaped piece (Figs 24G–I, 25G–I); retrolateral process on cymbium present but lacking setae (Fig. 24C)
	| colspan="1" rowspan="1" | result text = ''' ''{{Taxon name|M. legendrei}}'' '''}}
{{Key End}}

==Original Description==
*{{aut|Wood, H}}; {{aut|Scharff, N}}; 2017: A review of the Madagascan pelican spiders of the genera ''Eriauchenius'' O. Pickard-Cambridge, 1881 and ''Madagascarchaea'' gen. n. (Araneae, Archaeidae) [https://zookeys.pensoft.net/ ''ZooKeys'',] '''(727)''': 1-96. {{doi|10.3897/zookeys.727.20222}}

==Images==
{{Gallery | lines=5 | width=250
|1= File:zookeys-727-001-g001.jpg|2= '''Figure 1.''' Total evidence phylogeny from Bayesian analysis of molecular and morphological data, from Wood et al. (2015)<ref name="B27"> {{aut|Wood H}}, {{aut|Gillespie R}}, {{aut|Griswold C}}, {{aut|Wainwright P}} (2015) Why is Madagascar special? The extraordinarily slow evolution of pelican spiders ({{Taxon name|Araneae}}, {{Taxon name|Archaeidae}}). Evolution 69: 462–481. https://doi.org/10.1111/evo.12578</ref >. Outgroups not shown and with the morphospecies names changed to the new species names from the current study. Posterior probabilities are as follows, greater than or equal to: (*) 0.90; (**) 0.95; (***) 0.99.
|3= File:zookeys-727-001-g023.jpg|4= '''Figure 23.''' ''{{Taxon name|Madagascarchaea fohy}}'' sp. n. '''A''' male (holotype, [http://data.antweb.org/specimen/CASENT9015766 CASENT9015766]) habitus, lateral view '''B''' female ([http://data.antweb.org/specimen/CASENT9015766 CASENT9015766]) internal genitalia, dorsal view '''C–L''' male pedipalpal bulbs (holotype, [http://data.antweb.org/specimen/CASENT9015766 CASENT9015766]) '''D–F''' right bulb, image reversed '''C, G–L''' left bulb '''C''' cymbium close-up, dorsal view, arrow showing the retrolateral cymbium protrusion that has setae '''D, G, J''' prolateral view, arrow showing rod-like piece on embolus '''E, H, K''' ventral view, arrow showing rod-like piece on embolus '''F, I, L''' retrolateral view. Scale bars: 1 mm ('''A'''); 0.25 mm ('''B, D''').
|5= File:zookeys-727-001-g024.jpg|6= '''Figure 24.''' ''{{Taxon name|Eriauchenius legendrei}}'' (Platnick, 1991). '''A''' male ([http://hol.osu.edu/spmInfo.html?id=USNMENT01377217 USNMENT01377217]) habitus, lateral view '''B''' female ([http://data.antweb.org/specimen/CASENT9012333 CASENT9012333]) internal genitalia, dorsal view. '''C–I''' male pedipalpal bulbs '''D–F''' left bulb ([http://data.antweb.org/specimen/CASENT9012333 CASENT9012333]) '''C''' left bulb ([http://data.antweb.org/specimen/CASENT9010074 CASENT9010074]) '''G–I''' left bulb ([http://hol.osu.edu/spmInfo.html?id=USNMENT01377217 USNMENT01377217]): '''C''' cymbium close-up, dorsal view, arrow showing the small retrolateral cymbium protrusion that lacks setae '''D, G''' prolateral view '''E, H''' ventral view '''F, I''' retrolateral view. Scale bars: 1 mm ('''A'''); 0.25 mm ('''B, D''').
|7= File:zookeys-727-001-g025.jpg|8= '''Figure 25.''' ''{{Taxon name|Madagascarchaea rabesahala}}'' sp. n. '''A''' male (holotype, [http://data.antweb.org/specimen/CASENT9004011 CASENT9004011]) habitus, lateral view, image reversed '''B''' female ([http://data.antweb.org/specimen/CASENT9004086 CASENT9004086]) internal genitalia, dorsal view '''C–I''' male pedipalpal bulbs '''D–F''' right bulb ([http://data.antweb.org/specimen/CASENT9004086 CASENT9004086]), image reversed '''C, G–I''' left bulb ([http://data.antweb.org/specimen/CASENT9004086 CASENT9004086]): '''C''' cymbium close-up, dorsal view, arrow showing the absence of a retrolateral cymbium protrusion '''D, G''' prolateral view '''E, H''' ventral view '''F, I''' retrolateral view. Scale bars: 1 mm ('''A'''); 0.25 mm ('''B, D''').
|9= File:zookeys-727-001-g026.jpg|10= '''Figure 26.''' ''{{Taxon name|Madagascarchaea jeanneli}}'' (Millot, 1948). '''A''' male ([http://data.antweb.org/specimen/CASENT9004085 CASENT9004085]) habitus, lateral view, image reversed '''B–C''' female ([http://data.antweb.org/specimen/CASENT9012336 CASENT9012336]) internal genitalia '''B''' dorsal view '''C''' anterior view. '''D–L''' male pedipalpal bulbs '''D–I''' left bulb ([http://data.antweb.org/specimen/CASENT9004085 CASENT9004085]) '''J–L''' left bulb ([http://data.antweb.org/specimen/CASENT9005233 CASENT9005233]): '''D, G, J''' prolateral view '''E, H, K''' ventral view '''F, I, L''' retrolateral view. Scale bars: 1 mm ('''A'''); 0.125 mm ('''B, D''').
|11= File:zookeys-727-001-g027.jpg|12= '''Figure 27.''' ''{{Taxon name|Madagascarchaea lotzi}}'' sp. n. '''A''' male ([http://data.antweb.org/specimen/CASENT9018921 CASENT9018921]) habitus, lateral view, arrow showing posterior invagination of abdomen '''B–C''' female ([http://data.antweb.org/specimen/CASENT9012008 CASENT9012008]) internal genitalia '''B''' dorsal view '''C''' anterior view '''D–L''' male pedipalpal bulbs '''D–F''' right bulb ([http://data.antweb.org/specimen/CASENT9012004 CASENT9012004]), image reversed, '''G–L''' left bulb ([http://data.antweb.org/specimen/CASENT9012330 CASENT9012330]) '''D, G, J''' prolateral view '''E, H, K''' ventral view '''F, I, L''' retrolateral view. Scale bars: 1 mm ('''A'''); 0.125 mm ('''B, D''').
|13= File:zookeys-727-001-g028.jpg|14= '''Figure 28.''' ''{{Taxon name|Madagascarchaea moramora}}'' sp. n. '''A''' male (holotype, [http://hol.osu.edu/spmInfo.html?id=USNMENT01377197 USNMENT01377197]) habitus, lateral view, image reversed, arrow showing abdomen posterior is straight. '''B–C''' female ([http://hol.osu.edu/spmInfo.html?id=USNMENT01377198 USNMENT01377198]) internal genitalia '''B''' dorsal view '''C''' anterior view '''D–I''' male pedipalpal bulbs (holotype, [http://hol.osu.edu/spmInfo.html?id=USNMENT01377197 USNMENT01377197]): '''D–F''' left bulb, '''G–I''' right bulb, expanded, image reversed '''D, G''' prolateral view '''E, H''' ventral view; '''F, I''' retrolateral view. Scale bars: 1 mm ('''A'''); 0.125 mm ('''B, D''').
}}

==Other References==

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