Oryctophileurus

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Taxonavigation

Ordo: Coleoptera
Familia: Scarabaeoidea

Name

Oryctophileurus Kolbe, 1878 – Wikispecies link – Pensoft Profile

Type species

Phileurus nasicornis Burmeister, 1847 (original combination) (Fig. 3).
Species of Oryctophileurus are distinguished from other Phileurini by a combination of the following apomorphic characters: a single cephalic horn with lateral carina, pronotal cavity with ocellate punctures and two teeth or tubercles close to the anterior pronotal margin, teeth vertically positioned at about the same level as the outer eye margins.

Taxonomy and discussion

The following characters were cited by Endrödi (1985) for Oryctophileurus, Amblyodus, Microphileurus Kolbe, Metaphileurus Kolbe, Trioplus Burmeister, and Goniophileurus Kolbe and are here not considered as generic: outer side of mandible tridentate; antenna 10-jointed; elytra with punctate rows; proleg sexually monomorphic, protarsus not thickened in male. Oryctophileurus and Amblyodus are distinguished from the other mentioned genera by large horns, a distinctly developed and posteriorly carinate pronotal cavity and four external teeth on the protibia (Endrödi 1985; Ide 1998). However, in Amblyodus the two cephalic horns lack the lateral carina, the punctures in the pronotal cavity are predominately fused to short, transverse ridges, and the teeth close to the anterior pronotal margin are absent. Except for the horn number, these generic differences between Oryctophileurus and Amblyodus have not been recognized by previous workers.
Based on the combination of apomorphic characters of Oryctophileurus that lacks in Amblyodus we suggest retaining the genus Oryctophileurus.

Key to species of Oryctophileurus

Key to species of Oryctophileurus — R. Perger , P. Grossi

1		Horn on frons not projecting above anterior edges of pronotal protuberance (Figs 3B, D). Distance between inner teeth of dorsal pronotal protuberance (Figs 3A, C) narrower than distance between eyes. Development of horn and pronotal protuberance equal in both sexes. Color dark brown    Oryctophileurus nasicornis (Burmeister)
–		Horn on frons projecting above anterior edges of pronotal protuberance in both sexes. Horn longer in males than in females of the same size. Color black    2
2		Development of pronotal protuberance subequal in both sexes. Elytral striae and interstitial punctures arranged in irregular lines, punctures enlarged (Figs 4A, C)    Oryctophileurus varicosus Prell
–		Pronotal protuberance more strongly developed in males than females of the similar size. Elytral striae and interstitial punctures arranged in regular lines    3
3		Distance between inner teeth of dorsal pronotal protuberance in male wider than width between eyes (Fig. 7A); in females separated by a concavity with ocellate punctures (Fig. 6C). Pronotal concavity accounts for 40–50 % of the dorsal pronotal surface in males. Ocellate punctures above posterior-lateral pronotal margin continuous (Fig. 7C)    Oryctophileurus armicollis Prell
–		Distance between inner teeth of dorsal pronotal protuberance in male as wide as width between eyes (Fig. 7B), in females separated by a small fissure only (Fig. 8C). Anterior pronotal concavity accounts for about 30% of the dorsal pronotal surface in males. Area of ocellate punctures above posteriolateral pronotal margin reduced laterally (Fig. 7D), punctures small, shallow and sparse    Oryctophileurus guerrai Perger & Grossi, sp. n.

Taxon Treatment

• Perger, R; Grossi, P; 2013: Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini) ZooKeys, 346: 1-16. doi

Images

Figure 3. Dorsal and lateral habitus of Oryctophileurus nasicornis (Burmeister, 1847) A and B male C and D female, scale bar 5 mm.  Figure 4. Dorsal and lateral habitus of Oryctophileurus varicosus Prell, 1934, A and B male C and D female, scale bar 5 mm.  Figure 6. Dorsal and lateral habitus of Oryctophileurusarmicollis Prell, 1911, A and B holotype male, body length 18.5 mm C and D female, body length 18 mm, scale bar 5 mm.  Figure 7. Pronotum of males with body length 20 mm, dorsal, higher areas light, inner teeth of transversal pronotal carina indicated by white arrow: A Oryctophileurus armicollis Prell B Oryctophileurus guerrai; ocellate punctures at posteriolateral pronotal surface, character state does not differ between sexes C Oryctophileurus armicollis D Oryctophileurus guerrai.  Figure 8. Dorsal and lateral habitus of Oryctophileurus guerrai sp. n., A and B holotype male, body length 20 mm C and D allotype female, body length 19 mm, scale bar 5 mm.