Orthomorpha elevata
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Ordo: Diplopoda
Familia: Paradoxosomatidae
Genus: Orthomorpha
Name
Orthomorpha elevata Likhitrakarn & Golovatch & Panha, 2011 sp. n. – Wikispecies link – ZooBank link – Pensoft Profile
Holotype
♂ (CUMZ), Malaysia, Perak State, Bondig, Air Banum, ca 500 m, 5°38'13"N, 101°42'41"E, 27.05.2011, leg. S. Panha.
Paratypes
2 ♀ (CUMZ), same data, together with holotype. 1 ♀ (CUMZ), Malaysia, Johor State, Sungai Bantang, ca 100 m, 2°19'50"N, 103°09'45"E, 21.05.2011, leg. R. Chanabun. 1 ♀ (CUMZ), same State, Sungai Bantang, ca 100 m, 2°19'50"N, 103°09'45"E, 21.05.2011, leg. R. Chanabun. 2 ♀ (CUMZ), Kelantan State, near Gua Pulai, ca 120 m, 4°74'36"N, 101°56'32"E, 26.05.2011, leg. R. Chanabun. 2 ♀ (CUMZ), Kelantan State, near Gua Matu Madu, ca. 130 m, 4°50'13"N, 101°56'56"E, 26.05.2011, leg. R. Chanabun.
Name
To emphasize the mostly strongly elevated paraterga.
Diagnosis
Differs by mostly strongly elevated paraterga, coupled with two small sternal cones between ♂ coxae 4 (see also Key below).
Description
Length 34 (holotype) or 31–38 mm (♀), width of midbody pro- and metazona 2.8 and 4.3 mm (holotype), 2.9–3.5 and 4.3–5.0 mm (♀), respectively.
Coloration of live material (Fig. 100A) blackish to blackish-brown with contrasting pinkish paraterga legs, and tip of epiproct; coloration of alcohol material after long-term preservation blackish-brown with contrasting light yellowish calluses; two paramedian spots divided by a broad, blackish, axial stripe on prozona; lateral parts of paraterga near calluses, tip of epiproct, legs and venter light yellow-brown, legs slightly infuscate distally; antennae brown, distal part of antennomere 5, distal 2/3 of antennomere 6 and entire antennomere 7 blackish-brown (Fig. 100B-H).
Clypeolabral region densely setose, vertex bare, epicranial suture distinct. Antennae short, slightly clavate (antennomere 6 broadest), extending behind body segment 3 (♂) or 2 (♀) dorsally. Head in width < collum < segments 3 and 4 < 2 < 5–17 (♂, ♀); thereafter body gently and gradually tapering. Collum with three transverse rows of medium-sized setae, pattern untraceable; paraterga slightly declivous, subtriangular, with a faint lateral incision near midway, evidently discontinuing dorsum’s convexity, caudal corner pointed, about level to rear tergal margin (Fig. 100B & C). Tegument rather poorly shining, metaterga, prozona and most of metazona below paraterga finely shagreened; metaterga dull, leathery, roughly shagreened, their caudal margin evidently ribbed, with 3+3 or 4+4 short wrinkles; surface below paraterga rugulose, on segments 2–4(5) microgranulate. Postcollum metaterga with fully abraded setae, setation pattern untraceable even as insertion points. Axial line visible on collum and both on following pro- and metazona. Paraterga very strongly developed (Fig. 100A-H), especially well so in ♂, set at about 1/4 body height, postcollum ones upturned in lateral view, moderately enlarged on pore-bearing segments, thinner on poreless ones; shoulders broadly rounded and narrowly bordered, fused to callus; paraterga 2 and 19 (♂) or 19 (♀) lying below dorsum, paraterga 3 (♂) or 2–4 and 18 (♀) level to dorsum, remaining postcollum paraterga clearly above dorsum; caudal corner of postcollum paraterga pointed or narrowly rounded, always extending beyond rear tergal margin, best developed on segments 17 and 18 (Fig. 100F & G). Calluses delimited by a sulcus both dorsally and ventrally, with an evident front denticle only on paraterga 2, lateral margin virtually unincised on following paraterga, except for a small sinuosity in front of ozopore on pore-bearing segments. Posterior edge of paraterga strongly concave, especially strongly so on segments 18 and 19. Ozopores evident, lateral, lying in an ovoid groove at about 1/4 in front of caudal corner. Transverse sulcus complete on metaterga 4–18, narrow, superficial, not reaching bases of paraterga. Stricture between pro- and metazona narrow, rather deep, beaded at bottom down to base of paraterga (Fig. 100B-F & H). Pleurosternal carinae complete crests with a caudal tooth on segments 2–4, thereafter increasingly strongly reduced and remaining visible only as a front bulge and a caudal tooth until segment 17 (♂) or 13 (♀). Epiproct (Fig. 100F-H) conical, flattened dorsoventrally, with two small teeth directed ventrocaudally, emarginate or subtruncate at tip; pre-apical papillae small denticles lying very close to tip. Hypoproct (Fig. 100G) semi-circular, caudal margin with two strong and well-separated setiferous knobs.
Sterna sparsely setose, shining, without modifications; cross-impressions shallow; a paramedian pair of rather small, but evident tubercles in front of gonopod aperture; a small central lobe with a paramedian pair of evident, setose, apical cones between ♂ coxae 4 (Fig. 100I & J). Legs long and slender, slightly incrassate in ♂, midbody ones ca 1.3–1.4 (♂) or 1.1–1.2 times (♀) as long as body height, prefemora without modifications; ♂ tarsal brushes visible on legs 1–3, thereafter gradually thinning out.
Gonopods (Figs 101 & 102) simple. Coxa long and slender, with several strong setae distodorsally. Prefemur densely setose, more than 2 times shorter than femorite + “postfemoral” part. Femorite slightly curved, slender, nearly not enlarged distad, with a “postfemoral” part demarcated by an oblique lateral sulcus. Solenophore tip distinctly trifid, terminal prong being sharp and longest, middle denticle nearly as long as subterminal lobule; solenomere long and flagelliform.
Remark
This new species is rather widespread in northern Malaysia (Map 2).
Original Description
- Likhitrakarn, N; Golovatch, S; Panha, S; 2011: Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae) ZooKeys, 131: 1-161. doi
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