Trichosiopsis supralata

Taxonavigation

Ordo: Diptera
Familia: Sciaridae
Genus: Trichosiopsis

Name

Trichosiopsis supralata  unpublished

Type material

Holotype: ♂, 29.9.–7.10.2014, Malaise trap, leg. Jostein Kjærandsen, TSZD-JKJ-104097 in TSZD

Type locality

Norway, Finnmark, Alta, Gargia

Barcoded material

Stadium	Country	Province	Locality	Habitat	Method	Date	Collector	Collection Number	Collection	Latitude	Longitude	Elevation (m)
1 ♂ (holotype)	Norway	Finnmark	Alta, Gargia, along Vinabekken		Malaise trap	29.9.–7.10.2014	Jostein Kjærandsen	TSZD-JKJ-104097	TSZD	69.84	23.429
1 ♂	Norway	Finnmark	Alta, Gargia, along Vinabekken		Malaise trap	29.9.–7.10.2014	Jostein Kjærandsen	TSZD-JKJ-104096	TSZD	69.84	23.429
1 ♂	Finland	Lapland	Kemijärvi, Huttoja	spruce mire	Malaise trap	11.6.–11.7.2014	Jukka Salmela	ZFMK-TIS-2547878	ZFMK	66.9983	27.0261	193
1 ♂	Finland	Lapland	Savukoski, Tyroja	spruce forest	Malaise trap	1.7.–5.8.2014	Jukka Salmela	ZFMK-TIS-2541417	ZFMK	68.1383	28.5722	200
1 ♂	Finland	Lapland	Savukoski, Ainijaervi 4	swampy meadow near river	Malaise trap	2.6.–17.6.2015	Jukka Salmela	JS-COI-2016-0209	LMM	67.7616	29.4406

Other material studied

Finland: 2 ♂, Salla, Varriö, Malaise trap, Salmela, 4.-29.06.2013, MZH 9415/9418; 1 ♂, Ks, Taivalkoski, Pulkkaoja, by a brook, Malaise trap, Salmela, 03.07.2006, MZH 9258; 1 ♂, Li, Utsjoki, Tundra, Malaise trap, Fatouros, 17.-26.06.2000, PKHH 3243; Norway: 4 ♂, Finnmark, Nilsrud, moist birch forest, Malaise trap, Jaschhof, 20.6.-10.07.2003, SDEI 2977-2980; Sweden: 1 ♂, Hälsingland, Stensjön-Lomtjärn, marsh pine wood close to bog, Malaise trap, Swedish Malaise Trap Project, 15.-29.06.2005, PKHH 7934; 3 ♂, Härjedalen, Nyvallen, Jämtlands län, Nyvallens fäbod: alpine birch and spruce wood, Malaise trap, Swedish Malaise Trap Project, 4.7.-04.08.2004, NHRS 3277/3278/4827; 1 ♂, Pite lappmark, Arjeplog, Lake Sädvajaure (northern end); subalpine birch forest, exhaustor, Jaschhof & Jaschhof, 07.07.2005, SDEI 2461.

Description (male)

Head. Eye bridge 4–5 rows of facets. Antenna unicolour. LW-index of 4^th flagellomere 1.7–2.1; neck 0.2–0.35 × segment width; transition of basal part to neck pronounced. Colour of neck unicolour. Antennal setae shorter than segment width; of normal strength; dense. Palpus darkened, or dark; of normal length; palpomeres 3. First palpomere of normal shape; with 4–6 setae; with only sparse sensilla. Second palpomere short and oval. Third palpomere as long as first. Thorax. Colour dark brown, or brown. Notum unicolour. Thoracic setae normal; black. Posterior pronotum bare. Mesothoracic sclerites bare. Legs. Colour brown. Hind coxa of same colour as femora, or darkened. Setae on front coxa black, or darkened. Front tibial organ as patch of setae; pale; front tibial organ not bordered. Tibial setae on hind legs normal, shorter than tibial width. Tibial spurs of equal length. Claws untoothed. Wing. Wing slightly darkened; of normal shape. Wing membrane without macrotrichia. Wing venation weak, with faint stM. M-fork of normal shape. R₁ ending at or slightly before base of m-fork; posterior veins with macrotrichia; stM mostly with macrotrichia, or with a few macrotrichia; CuA₁ and CuA₂ mostly with macrotrichia; bM bare; r-m with few setae, or mostly setose; bM:r-M 0.4–0.6; st-Cu:bM 0.4–0.65; R₁:R 1.25–1.5; c:w 0.55–0.65. Halter darkened; of normal length. Abdomen. Abdominal setae strong; on tergites black; on sternites black. Hypopygium concolour with abdomen; LW-index 0.5–0.7. Base of gonocoxites with weak setae; gonocoxites forming an inverse V; inner margin of gonocoxites narrowly U-shaped; inner membrane of hypopygium scarcely setose; ventral margin of gonocoxite with short setae. Gonostylus elongate; LW-index 1.8–2.2; Inner margin concave; apex nearly rectangular. Apical tooth present; without internal structure; strong; LW-Index 2.3–2.7. Awl-like setae normal; beneath apical tooth absent. Megasetae absent. Whiplash-hair absent. Tegmen 0.7–0.8 × longer than broad; rectangular with rounded edges; without special features; central process absent. Length of ejaculatory apodeme/hypopygium 25–40 %; base of ejaculatory apodeme absent. Field with aedeagal teeth inconspicuous. Measurements. Body size 2.7–3 mm. Hind tibia 1.27–1.45 mm. Wing length 2.5–2.9 mm.

Diagnosis

Tr. supralata is is easily to recognize by the extremely distended apical part of the gonostylus. This and the inverse-V structure at the hypopygial base are typical characters for the subgenus Leptospina, but as the prolonged dorsal awl-like spines are missing and the posterior wing veins are setose, the new species is assigned to Trichosiopsis The nearly rectangular shape of the gonostylus resembles Trichosiopsis scutellata, from which it can be separated otherwise by the not modified setae on the hypopygial base and the shorter setae at the antennae. Unusual for Trichosiopsis is the pale setosity of the tibial organ. In most other known species except for Trichosiopsis claviforceps and the Trichosiopsis trochanterata complex it consists of dark setae. The species is most similar to Trichosiopsis wiebke. For differences see under the latter.

DNA Barcoding

The COI sequence is assigned to BIN BOLD:ACU4367 (average distance 0.13%, max. 0.32%, n=5, K2P: 9.63%).

Etymology

lat. supralatus = very broad; making reference to apically enlarged gonostylus.

Ecology

Trichosiopsis supralata has a strictly boreal distribution. It was found to date only in Fennoscandia and only north of 59° N.

Distribution

Finland, Norway, Sweden.

Images

Figure 1. Trichosiopsis supralata male, habitus, scale 1.0 mm  Figure 2. Trichosiopsis supralata male, hypopygium, scale 0.1 mm  Figure 3. Trichosiopsis supralata male, gonostylus, scale 0.1 mm  Figure 4. Trichosiopsis supralata male, aedeagal complex, scale 0.1 mm  Figure 5. Trichosiopsis supralata male, 4th antennal flagellomere, scale 0.1 mm  Figure 6. Trichosiopsis supralata male, palpus, scale 0.1 mm  Figure 7. Trichosiopsis supralata male, tibial organ, scale 0.1 mm  Figure 8. Trichosiopsis supralata male, wing, scale 1.0 mm