Sinogodavaria

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Taxonavigation

Ordo: Diptera
Familia: Phoridae

Name

Sinogodavaria Liu, 2020 gen. nov. – Wikispecies link – ZooBank link – Pensoft Profile

Type species

Sinogodavaria multiformis sp. nov.

Diagnosis

Supra-antennal setae absent; flagellomere 1 not elongate; costa of female thickened; wing vein Rs with several fine setulae along upper side; vein Rs deflected slightly at the junction with vein M1; vein R2+3 absent; tip of vein R4+5 not enlarged; hind tibia with an antero-apical seta but without an antero-basal seta; male hypandrium without dense microsetae; aedeagus with a long, curled sclerotized process.

Description

Head. Frons generally broader than long. Median furrow present, vestigial. Supra-antennal setae absent. First and second rows of setae convex. Flagellomere 1 globose; arista sub-apical. Palpus oval, with apical setae and ventral setulae.
Thorax. Propleuron of thorax with three ventral setae, two posterior setae, and scattered setulae. Anepisternum with fine setulae on upper part. Notopleura with four setae, the second being smaller than the others. Scutellum with an anterior pair of small setulae and a posterior pair of setae.
Legs. Fore tibia with a dorsal seta near basal two fifths and several small setulae below it. Mid tibia with the normal basal pair of setae and an antero-apical seta. Hind tibia with two dorsal longitudinal setal palisades, one antero-apical seta, one robust ventral spur and a series of weak posterior and postero-dorsal spurs.
Wing. Costa usually extending to half of wing length. Wing vein Rs with several fine setulae along upper side. Female costa thickened around junction with vein R1 or at the first section of costa. Vein Sc reaching vein R1. Axillary ridge with 4–6 long, black, feathered setae. Haltere yellowish brown, knob black.
Abdomen. Female without tergite VII. Left side of epandrium slender, rounded apically; right side of epandrium large, triangular. Hypandrium with ventrally directed outer lobe covered with rounded spinuli. Aedeagus dark brown, supported by aedeagal apodeme; its left plate with a long, curled, sclerotized process. Anal tube short.

Etymology

The genus name is derived from Sino and Godavaria and refers to the locality and to the relationship with the genus Godavaria Brown.

Distribution

China (Liaoning, Hebei, Shaanxi, Sichuan).

Recognition

In Disney’s (1994) male and female keys to genera, the new genus runs to couplets 9 (males) and 119 (females), respectively, both Borophaga Enderlein, 1924. There is no further division of the genus Borophaga in these keys. In the modified key to the Borophaga group (Bänziger and Disney 2006), the new genus runs to couplet 3 as Peromitra Enderlein, 1924 and Latiborophaga Brown, 1992. Sinogodavaria gen. nov. is distinguished from Peromitra by a subcircular and non-erect anterior ocellus and thickened costa of the female wing. It differs from Latiborophaga by the more-or-less straight vein Rs and the absence of an antero-basal seta on the hind tibia. It differs from the genus Chaetogodavaria Liu, 1996 by the absence of a long seta on the anepisternum (Liu 1996). In consideration of new data, the following modifications of the male and female keys to genera by Disney (1994) are proposed.

Partial modification of the key to genera (males) by Disney (1994)

Partial modification of the key to genera (males) by Disney (1994) — G. Liu

9		Posterior ocelli close to eye margin and ocellar triangle strongly demarcated at front by a sinuous furrow     Stichillus
–		Posterior ocelli well removed from eye margin and ocellar triangle not demarcated in this way at front    9a
9a		Tip of wing vein Rs thickened     Borophaga
–		Tip of wing vein Rs not thickened    9b
9b		Hind tibia with antero-apical seta, but without antero-basal seta    9c
–		Hind tibia with both antero-apical and antero-basal setae    9e
9c		Anepisternum with a strong seta and short setulae     Chaetogodavaria
–		Anepisternum only with short setulae    9d
9d		Wing vein R1+2 present and strongly developed     Godavaria
–		Wing vein R1+2 absent; aedeagus with a long, curled, sclerotized process    Sinogodavaria gen. nov.
9e		Wing vein Rs deflected at mid length; anterior ocellus not elevated; hypandrium with dense microsetae     Latiborophaga
–		Wing vein Rs more or less straight; anterior ocellus elevated; hypandrium without dense microsetae     Peromitra

Partial modification of the key to genera (females) by Disney (1994)

Partial modification of the key to genera (females) by Disney (1994) — G. Liu

114		Anepisternum bare; hind tibia without longitudinal setal palisades    115
–		Anepisternum with setulae; hind tibia with two dorsal longitudinal setal palisades    114a
114a		Wing vein R2+3 very weak; tip of vein R1 thickened    Borophaga (part)
–		Wing vein R2+3 strongly developed; tip of vein R1 not thickened     Godavaria
119		Posterior ocelli clearly closer to eyes than to anterior ocellus, and ocellar region usually clearly demarcated in front by a sinuous furrow     Stichillus
–		Posterior ocelli clearly closer to anterior ocellus than to eyes, and ocellar region not clearly demarcated in this way in front    119a
119a		Tip of vein R1 thickened    Borophaga (part)
–		Tip of vein R1 not thickened    119b
119b		Anterior ocellus elevated, broader than high; costa not thickened     Peromitra
–		Anterior ocellus not elevated, subcircular; costa thickened    119c
119c		Hind tibia with antero-basal seta; vein Rs deflected at mid length     Latiborophaga
–		Hind tibia without antero-basal seta; vein Rs deflected at the junction with vein M1    Sinogodavaria gen. nov.

Key to the species of Sinogodavaria gen. nov.

Key to the species of Sinogodavaria gen. nov — G. Liu

1		Female    2
–		Male    4
2		Abdominal tergites light brown with some dark area; tergite III wide-mouth-urn shaped (Fig. 7)    S. multiformis sp. nov.
–		Abdominal tergites uniformly dark brown; tergite III trapezoid (Fig. 20)    3
3		Tergite VI triangular (Fig. 28)    S. bathmis (Liu)
–		Tergite VI trapezoid (Fig. 20)    S. tenebrosa sp. nov.
4		Tergites with light rear band; hypandrium with a short pointed fronto-ventral process (Figs 11, 12, 13)    S. multiformis sp. nov.
–		Tergites without light rear band; hypandrium with a long hockey-stick-shaped fronto-ventral process (Figs 24–26)    S. tenebrosa sp. nov.

Original Description

• Liu, G; 2020: A new genus of the Hypocera group (Diptera, Phoridae), with descriptions of two new species from China ZooKeys, 932: 113-127. doi

Images

Figures 1–7. S. multiformis sp. nov. 1–3 male 1 lateral view of body 2 head 3 wing 4–7 female 4 lateral view of body 5 head 6 wing 7 abdominal tergites. Scale bars: 0.5 mm.  Figures 8–13. Male terminalia of S. multiformis sp. nov. 8, 12 left view 9, 13 right view 10 front view 11 ventral view. Scale bars: 0.1 mm.  Figures 14–20. S. tenebrosa sp. nov. 14–16 Male 14 lateral view of body 15 head 16 wing 17–20 female 17 lateral view of body 18 head 19 wing 20 abdominal tergites. Scale bars: 0.5 mm.  Figures 21–26. Male terminalia of S. tenebrosa sp. nov. 21–25 left view 22–26 right view 23 front view 24 ventral view. Scale bars: 0.1 mm.  Figures 27–29. S. bathmis (Liu) 27 head 28 abdominal tergites V–VI 29 lateral view of body. Scale bars: 0.1 mm (27–28); 0.5 mm (29)