Pseudouroctonus peccatum

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Tate A, Riddle R, Soleglad M, Graham M (2013) Pseudouroctonus peccatum, a new scorpion from the Spring Mountains near “Sin City,” Nevada (Scorpiones, Vaejovidae). ZooKeys 364 : 29–45, doi. Versioned wiki page: 2013-12-18, version 40621, , contributors (alphabetical order): Pensoft Publishers.

Citation formats to copy and paste


author = {Tate, Amanda E. AND Riddle, Rebecca R. AND Soleglad, Michael E. AND Graham, Matthew R.},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {Pseudouroctonus peccatum, a new scorpion from the Spring Mountains near “Sin City,” Nevada (Scorpiones, Vaejovidae)},
year = {2013},
volume = {364},
issue = {},
pages = {29--45},
doi = {10.3897/zookeys.364.6288},
url = {},
note = {Versioned wiki page: 2013-12-18, version 40621, , contributors (alphabetical order): Pensoft Publishers.}


RIS/ Endnote:

T1 - Pseudouroctonus peccatum, a new scorpion from the Spring Mountains near “Sin City,” Nevada (Scorpiones, Vaejovidae)
A1 - Tate A
A1 - Riddle R
A1 - Soleglad M
A1 - Graham M
Y1 - 2013
JF - ZooKeys
JA -
VL - 364
IS -
UR -
SP - 29
EP - 45
PB - Pensoft Publishers
M1 - Versioned wiki page: 2013-12-18, version 40621, , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/zookeys.364.6288

Wikipedia/ Citizendium:

<ref name="Tate2013ZooKeys364">{{Citation
| author = Tate A, Riddle R, Soleglad M, Graham M
| title = Pseudouroctonus peccatum, a new scorpion from the Spring Mountains near “Sin City,” Nevada (Scorpiones, Vaejovidae)
| journal = ZooKeys
| year = 2013
| volume = 364
| issue =
| pages = 29--45
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.364.6288
| url =
| pmc =
| accessdate = 2020-04-10

}} Versioned wiki page: 2013-12-18, version 40621, , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Ordo: Scorpiones
Familia: Vaejovidae
Genus: Pseudouroctonus


Pseudouroctonus peccatum Tate & Riddle & Soleglad & Graham, 2013 sp. n.Wikispecies linkZooBank linkPensoft Profile

Type material

United States: Nevada: female holotype, Kyle Canyon Road, Spring Mountains, 36.2666°N, 115.5988°W, 10 May 2013, A.E. Tate, R.R. Riddle, and M.R. Graham leg. (AMNH, MRG1251); paratype, mature male, same location as holotype, A.E. Tate and R.R. Riddle leg. (AMNH, MRG1252); paratype, juvenile female, collected with holotype (AMNH, MRG1222); paratype, juvenile female, same location as holotype, 23 April 2013, A.E. Tate, R.R. Riddle, and D.R. Tate leg. (AMNH, MRG1221); juvenile female, same location as holotype, 2 July 2013, R.R. Riddle leg. (AMNH, MRG1253).


The specific epithet is Latin for “sin” in reference to the proximity of the type locality to Las Vegas, which is known informally as “Sin City.”


Medium to large sized species for the genus, females reaching 50 mm; pectinal tooth counts 13–14 for females and 15 for males. Hemispermatophore lacking secondary lamellar hook; lamellar hook protrudes somewhat from the lamina base creating a modest basal constriction; lamina terminus with a distinct distal crest. Cheliceral movable finger ventral edge with low-profiled pigmented crenulated teeth; fixed finger ventral edge with several small flat pigmented protuberances; movable finger dorsal edge with two subdistal (sd) denticles. Chelal movable finger with seven inner denticles (ID). Metasomal segment V somewhat stout in adults, length compared to width 1.908 in females and 1.953 in males.

Description of holotype

Color: Carapace, trochanter, femur, patella, tergites, and metasoma have a yellow-orange base color with dark brown to black markings along the carinae of the pedipalp and metasoma. Legs are light yellow. Pedipalp chelae are dark brown in color with darker reddish-brown coloration at the anterior portion of the palm where the fixed finger and movable finger meet. Chelicerae are light yellow on proximal half with dark reddish-brown fingers. Vesicle portion of the telson is yellow-orange with a dark reddish-brown to black aculeus. Pectines and genital operculum are light yellow.
Morphology: Carapace: trapezoidal with noticeably emarginated anterior margin; surface finely granular with scattered small granules, with larger granules symmetrically flanking the lateral and median eyes; median furrow is slight and traverses length of carapace; ratio of location of median eyes location (from anterior edge)/carapace length = 0.338. Tergites: surface with small granules on distal 1/3–1/2 of tergites I–VI; tergite VII with two pairs of granular lateral carinae, and a slight median hump. Sternites: III–VI smooth to very finely granular and without carinae; VII with granular ventral lateral carinae on posterior 4/5, median carinal pair essentially obsolete. Spiracles: ellipsoid and with median side rotated 40° away from posterior sternite margin. Genital Operculum: sclerites separated on posterior 1/5. Pectines: tooth count 13/13; middle lamellae 8/8; sensorial areas present on all pectine teeth. Metasoma: ratio of segment I length/width 0.91; segment II length/width 1.00; segment III length/width 1.06; segment IV length/width 1.36; segment V length/width 1.91. Segments I–IV: dorsal carinae are moderately denticulate on segments I–IV and have slightly enlarged distal denticles; dorsolateral carinae are moderately crenulate on segment I and moderately denticulate on segments II–IV with enlarged posterior denticles; posterior denticles are largest and most pronounced on dorsolateral carinae IV; ventrolateral carinae are moderately crenulate on segments I–IV; segments II and III have sparse and moderately crenulate intermediary carinae forming an approximately 30° angle with the dorsolateral carinae on the posterior 2/3 on segment II and posterior 1/3 segment III; ventromedian carinae are smooth to obsolete on segment I, and crenulate on segments II–IV; ventrolateral setae 2/2:2/2:2/2:2/2; ventral submedian setae 3/3:3/3:3/3:3/3. Segment V: dorsolateral carinae subtly denticulate; lateral carinae crenulate and obsolete on posterior 1/3; ventrolateral carinae crenulate; ventromedian carinae crenulate; intercarinal spaces finely granular; dorsolateral setation 3/3; lateral setation 3/3; ventrolateral setation 4/4; ventromedian setation 4/4. Telson: smooth to slightly granular with no subaculear tubercule and lacking LAS denticles (Fet et al. 2006). Chelicerae: dorsal edge of fixed finger with four teeth, one distal, one subdistal, one median, and one basal, the latter two denticles formed as a bicuspid; ventral edge with seven low-profile pigmented protuberances, the distal two smaller in size; dorsal edge of movable finger has five teeth total comprised of one distal, two subdistal, one median tooth, and one basal tooth; ventral edge of movable finger has seven small pigmented crenulated teeth; serrula with approximately 23 tines. Pedipalps:trichobothrial pattern type C, orthobothriotaxic: trichobothria ib-it positioned on extreme base of fixed finger, Dt located considerably basal of palm midpoint, Db ventral of digital carina, patellar trichobothria v3 adjacent to et3; ratio of chela length/width 3.57; femur length/width 2.60; patella length/width 2.79; fixed finger length/carapace length 0.78. Chela: median denticles (MD) of fixed finger aligned and divided into six subrows by five outer denticles (OD); flanked by six inner denticles (ID); movable finger with six subrows of MD, five OD and seven ID. Chela carinae: Digital carina rounded, rough, and somewhat flattened; subdigital essentially obsolete, formed by two granules; dorsosecondary flat, rounded, and roughly textured; dorsomarginal very rounded, with large scattered granules; dorsointernal rounded, with scattered granules; ventroexternal rounded, roughly textured, terminating at external condyle of movable finger; ventromedian flat, essentially obsolete; ventrointernal rounded, with small granules scattered distally; external carina very rounded, roughly textured. Femur: dorsointernal, and ventrointernal carinae denticulate and black in color, ventroexternal and dorsoexternal carinae denticulate with small granules throughout, internal surface has 6 prominent dentate granules. Patella:dorsointernal carinae are denticulate, dorsoexternal carinae are mildly crenate, ventral internal carinae are dentate, ventroexternal carinae are moderately denticulate, internal surface has 11 prominent granules. Legs: Ventral surface of tarsus with single median row of 7–17 spinules terminating distally with one to two pairs of spinules. The terminal spinules are slightly stouter than the those comprising the median row. Flanking setal pairs are essentially absent in legs I–II, with 2–3 irregularly positioned and sized pairs on legs III–IV. Basitarsus spinule rows are limited, 2 rows on leg I, 1 row on leg II, 1 weak row on III, and absent on IV. Basitarsus populated with large irregularly placed darkly pigmented setae. Hemispermatophore: Left hemispermatophore is 5.7 mm in length; lamina length 3.71, lamellar hook length 1.03, and trough different (vertical distance between ventral and dorsal troughs) 0.55. Lamellar edges sub-parallel, except for slight expansion on mid-area of the internal edge; terminus blunted with a distinct distal crest on the dorsal side. Lamellar hook extends somewhat from lamina base, is distinctly bifurcated, and is formed entirely from the dorsal trough. A modest basal constriction and a deep truncal flexure are present. Sclerotized mating plugs with smooth barbs were extracted from the ventrointernal aspect of both hemispermatophore median areas.

Table 1. Measurements (in millimeters) of all known specimens of Pseudouroctonus peccatum.
Scorpion ID Holotype MRG1251 Female MRG1221 Female MRG1222 Male MRG1252 Female MRG1253
Total Length 52.45 mm 32.37 mm 27.71 mm 36.12 mm 39.83 mm
Carapace L 5.59 mm 4.37 mm 3.66 mm 4.53 mm 4.90 mm
Mesosoma L 19.89 mm 8.30 mm 8.67 mm 9.79 mm 13.94 mm
Metasoma L 20.8 mm 15.00 mm 11.70 mm 17.03 mm 16.36 mm
MET I L 3.15 mm 2.43 mm 1.98 mm 2.66 mm 2.53 mm
MET I W 3.45 mm 2.55 mm 2.12 mm 2.63 mm 2.81 mm
MET I D 2.65 mm 1.98 mm 1.62 mm 1.99 mm 2.20 mm
MET II L 3.43 mm 2.40 mm 1.79 mm 2.84 mm 2.69 mm
MET II W 3.42 mm 2.55 mm 2.12 mm 2.65 mm 2.79 mm
MET II D 2.91 mm 1.98 mm 1.55 mm 2.03 mm 2.31 mm
MET III L 3.59 mm 2.52 mm 1.90 mm 3.00 mm 2.82 mm
MET III W 3.39 mm 2.55 mm 2.06 mm 2.69 mm 2.69 mm
MET III D 2.62 mm 1.95 mm 1.62 mm 2.07 mm 2.22 mm
MET IV L 4.41 mm 3.06 mm 2.29 mm 3.61 mm 3.20 mm
MET IV W 3.25 mm 2.48 mm 1.98 mm 2.51 mm 2.63 mm
MET IV D 2.57 mm 1.95 mm 1.59 mm 2.15 mm 2.52 mm
MET V L 6.22 mm 4.59 mm 3.74 mm 4.98 mm 5.12 mm
MET V W 3.26 mm 2.48 mm 2.01 mm 2.55 mm 2.59 mm
MET V D 2.47 mm 1.77 mm 1.54 mm 1.95 mm 2.17 mm
Telson L 6.17 mm 4.47 mm 3.68 mm 4.77 mm 4.63 mm
Vesicle L 4.21 mm 3.02 mm 2.30 mm 3.39 mm 3.49 mm
Vesicle W 3.13 mm 2.31 mm 1.70 mm 2.33 mm 2.19 m
Vesicle D 2.41 mm 1.68 mm 1.49 mm 1.67 mm 2.62 mm
Aculeus L 1.96 mm 1.45 mm 1.38 mm 1.38 mm 1.14 mm
Pedipalp L 18.79 mm 15.09 mm 11.80 mm 16.08 mm 16.44 mm
Femur L 4.71 mm 3.74 mm 2.96 mm 3.88 mm 4.30 mm
Femur W 1.81 mm 1.26 mm 1.11 mm 1.39 mm 1.41 mm
Femur D 1.24 mm 0.95 mm 0.82 mm 1.00 mm 0.95 mm
Patella L 5.02 mm 3.80 mm 3.40 mm 4.05 mm 4.41 mm
Patella W 1.80 mm 1.43 mm 1.19 mm 1.42 mm 1.59 mm
Patella D 1.71 mm 1.38 mm 0.92 mm 1.34 mm 1.38 mm
Chela L 9.06 mm 7.55 mm 5.27 mm 8.15 mm 7.73 mm
Palm W 2.54 mm 1.81 mm 1.26 mm 2.10 mm 1.49 mm
Palm L 4.84 mm 2.85 mm 2.46 mm 3.79 mm 3.38 mm
Chela D 3.31 mm 2.23 mm 1.44 mm 2.84 mm 2.65 mm
MF L 5.55 mm 4.05 mm 3.33 mm 4.36 mm 4.35 mm
FF L 4.37 mm 3.32 mm 2.94 mm 3.25 mm 3.65 mm
Cara to eyes 1.89 mm 1.65 mm 1.20 mm 1.37 mm 1.67 mm
Pectine count 13/13 14/14 13/13 15/15 14/14
Table 2. Matrix showing character set that distinguishes Pseudouroctonus peccatum sp. n. from the other seventeen species in the genus. If a species differs significantly for a given data character, its state is shaded. Note, except for Pseudouroctonus peccatum sp. n., most of the data contained in this table is based on published information (see reference list). ID = inner denticle.
Hemispermatophore Cheliceral
Chelal Movable Finger ID Pectinal Tooth Counts Metasoma Seg V (Length/Width)
Pseudouroctonus peccatum Distal crest present on lamina terminus; secondary lamellar hook absent va on MF and FF crenulated; two subdistal
Seven ID 15 ♂13–14 ♀ 1.953 ♂1.908 ♀
Pseudouroctonus andreas Distal crest absent smooth Six ID smaller
9–10 ♂ 7–9 ♀
1.933 ♂2.000 ♀
Pseudouroctonus angelenus Distal crest absent; secondary lamellar hook present smooth Seven ID smaller
11 ♂ ? ♀
3.059 ♂? ♀
Pseudouroctonus apacheanus  ? smooth Seven ID smaller
10–11 ♂ 9–10♀
2.412 ♂2.176 ♀
Pseudouroctonus bogerti Distal crest absent; secondary lamellar hook present smooth Seven ID -? ♂ 12 ♀ thinner
3.444 ♂3.091 ♀
Pseudouroctonus cazieri Distal crest absent va on MF and FF toothed Six ID -
13 ♂ 11–12 ♀
2.150 ♂2.217 ♀
Pseudouroctonus chicano  ? smooth Seven ID smaller
 ? ♂ 9♀
 ? ♂2.067 ♀
Pseudouroctonus glimmei Distal crest absent; secondary lamellar hook present smooth Seven ID smaller
10–12 ♂ 9–11♀
2.600 ♂2.609 ♀
Pseudouroctonus iviei Distal crest absent; secondary lamellar hook present smooth Seven ID smaller
11 ♂ 10♀
2.191 ♂2.304 ♀
Pseudouroctonus lindsayi Distal crest absent smooth Six ID -
14 ♂ 12 ♀
2.286 ♂2.286 ♀
Pseudouroctonus minimus castaneus Distal crest absent smooth Seven ID smaller
10–12 ♂ 10♀
1.402 ♂1.364 ♀
Pseudouroctonus minimus minimus Distal crest absent smooth Six ID smaller
10–11 ♂ 9–10♀
1.650 ♂1.565 ♀
Pseudouroctonus minimus thompsoni  ? smooth Seven ID smaller
10–11 ♂ 10♀
1.667 ♂1.538 ♀
Pseudouroctonus reddelli Distal crest absent va on MF and FF toothed Seven ID larger
18–19 ♂ 15–16♀
3.036 ♂2.871 ♀
Pseudouroctonus rufulus Distal crest absent smooth Seven ID smaller
11 ♂ 9♀
2.400 ♂2.176 ♀
Pseudouroctonus saavasi Distal crest absent Smooth, one subdistal (sd) Seven ID smaller
10–11 ♂ 9♀
2.810 ♂2.563 ♀
Pseudouroctonus sprousei Distal crest absent va on MF and FF toothed Seven ID larger
17 ♂ ?♀
4.214 ♂? ♀
Pseudouroctonus williamsi Distal crest absent; secondary lamellar hook present smooth Seven ID -
13 ♂ 11–12 ♀
3.400 ♂3.400 ♀

DNA barcodes

(COI) – MRG1221 (paratype, GenBank no. KF841448):
MRG1222 (paratype, GenBank no. KF841449):

Original Description

  • Tate, A; Riddle, R; Soleglad, M; Graham, M; 2013: Pseudouroctonus peccatum, a new scorpion from the Spring Mountains near “Sin City,” Nevada (Scorpiones, Vaejovidae) ZooKeys, 364: 29-45. doi