Psammoecus fasciatus

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Yoshida T, Hirowatari T (2014) A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae). ZooKeys 403 : 15–45, doi. Versioned wiki page: 2014-04-17, version 49499, , contributors (alphabetical order): Pensoft Publishers.

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author = {Yoshida, Takahiro AND Hirowatari, Toshiya},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)},
year = {2014},
volume = {403},
issue = {},
pages = {15--45},
doi = {10.3897/zookeys.403.7145},
url = {},
note = {Versioned wiki page: 2014-04-17, version 49499, , contributors (alphabetical order): Pensoft Publishers.}


RIS/ Endnote:

T1 - A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)
A1 - Yoshida T
A1 - Hirowatari T
Y1 - 2014
JF - ZooKeys
JA -
VL - 403
IS -
UR -
SP - 15
EP - 45
PB - Pensoft Publishers
M1 - Versioned wiki page: 2014-04-17, version 49499, , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/zookeys.403.7145

Wikipedia/ Citizendium:

<ref name="Yoshida2014ZooKeys403">{{Citation
| author = Yoshida T, Hirowatari T
| title = A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)
| journal = ZooKeys
| year = 2014
| volume = 403
| issue =
| pages = 15--45
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.403.7145
| url =
| pmc =
| accessdate = 2022-11-28

}} Versioned wiki page: 2014-04-17, version 49499, , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Ordo: Coleoptera
Familia: Silvanidae
Genus: Psammoecus


Psammoecus fasciatus Reitter, 1874Wikispecies linkPensoft Profile

  • Psamoecus [sic.] fasciatus Reitter, 1874: 525. Type locality: Japan; Type deposition: the Natural History Museum, London; Type examined. (misspelling)
  • Psammoecus fasciatus: Hetschko 1930[1]: 82. (catalogue) – Nakane 1963[2]: 196, fig. 16 in pl. 98. – Hisamatsu 1977[3]: 21. – Sasaji 1985[4]: 204, fig. 34 in pl. 32. – Sato 1989[5]: 377. – Halstead et al. 2007[6] – Hirano 2009[7]: 64. – Hirano 2010[8]: 12, 13. – Yoshida and Hirowatari 2013[9]: 5, 11, 16, 17.


This species is closely similar to Psammoecus hiranoi. It can be distinguished by the longer parameres, the shallower incision of the phallobase on the anterior 1/4 and the shorter teeth on the lateral margins of the pronotum.


Body length. 2.75–3.50 mm (n=49).
Coloration (Fig. 2D). Head and pronotum yellowish-brown to reddish-brown. Elytra with a variable horizontal dark band at middle, this band in lighter color specimens thinner towards elytral suture, lateral margins, apical portion and posterior elytral suture dark. Antennae brown or reddish-brown basally, 7th and 8th antennomeres darker, 9th and 10th almost black, 11th (apex) lighter than basal ones. Antennae of some specimens almost unicolorous.
Head (Fig. 11A, B, C). Broad, HW/HL 1.40–2.08; IE/HL 0.94–1.38. Temples enlarged behind eyes, narrowed gradually toward posterior margin. Eyes large, slightly prominent. Ventral surface with rough irregular punctuation, sparser punctuation on dorsal surface. Antennae 1.38–1.57 mm, 2nd antennomere short; covered with thick medium length pubescence and some long erect pubescence on each antennomere; approximate ratio of lectotype as follows: 2.5: 1.0: 1.3: 1.3: 1.3: 1.3: 1.3: 1.3: 1.3: 1.2: 1.8 (Fig. 11A).
Pronotum (Fig. 11B, C). Broad, PW/PL 0.69–1.52. Punctuation on dorsal surface very dense, but no punctures around posterior margin, punctuation on ventral surface sparser than dorsal surface, some of them in rows especially on posterior half. Pubescence composed of frequent short setae on dorsal surface and a long seta on each tooth on lateral margins and at anterior and posterior angles. Each anterior angle with a distinct group of a few very small teeth, each lateral margin with four teeth; tooth I comparatively small, teeth II, III and IV almost the same moderate size, each posterior angle with a comparatively small tooth.
Elytra (Fig. 11E). Elongate-oval, EW/BL 0.41–0.47. Punctures wider than interstices. Pubescence composed of semi-erect medium length setae and some long erect setae in a row around lateral margins, longer toward anterior portion.
9th abdominal sternite (Fig. 14M). Strut cut at around half, diverging widely around the posterior base, ends of each branch curved inwards. Lateral sclerites triangular, flat.
Aedeagus (Figs 14N and O). Parameres simple stick-shaped, elongate, slightly curved inwards, punctuated sparsely except base, a few semi-long setae around apical portion. Phallobase diverging gradually around anterior 1/4, a little bulging along inner margins of each branch, lateral margins broadening. Penis also elongate, punctuated on posterior 1/6, more densely on ventral portion, especially densely on apical portion.

Type series

Lectotype: male, Mt. Maya-san, Kobe City, Hyogo Prefecture, Japan, 1871, G. Lewis leg. (BMNH). Paralectotype: 1 female, same locality, 1871, G. Lewis leg. (BMNH). (here designated)

Specimens examined

Russia: [Primorsky Krai] 1 ex., Chuguevka Village, 27–VI–1999, Y. Notsu leg. (EUMJ). JAPAN: [Niigata Pref.] 1 male & 1 ex., Renge-Onsen, Itoigawa City, 14–VIII–2003, I. Tanaka leg. (ITC). [Tokyo Pref.] 7 exs., Mikura Island, Mikurajima Village, 11–VI–1977, T. Nakane leg. (HUSE). [Nagano Pref.] 1 ex., Togakushi-Kôgen, Nagano City, 5–6–VII–2008, K. Mizuno leg. (KMC); 2 ex., Ôbora, Ueda City, 24–VII–2013, T. Yoshida leg. (ELKU). [Kyoto Pref.] 1 ex., Kifune, Kyoto City, 6–IV–1988, I. Tanaka leg. (ITC). [Hyogo Pref.] 1 ex., Itoi, Asago City, 26–IX–2004, I. Tanaka leg. (ITC); 5 exs., Sakanotani, Shisô City, 19–VII–2009, K. Itô leg. (KMC); 1 ex., Onzui, Shisô City, 19–VII–2009, K. Utô leg. (KMC); 1 ex., Mt. Yuzuruha-san, Nantan Town, Awaji Island, 18–X–2003, I. Tanaka leg. (ITC). [Nara Pref.] 1 ex., Mt. Kasuga-yama, Nara City, 5–V–1985, I. Tanaka leg. (ITC); 2 exs., same locality, 5–V–2012, K. Ikeuchi leg. (ELKU); 1 ex., same locality, 31–VII–2012, K. Matsuda leg. (ELKU). [Tokushima Pref.] 1 ex., Mt. Shibakoya-yama, Kamiyama Town, 31–VII–1975, M. Yoshida leg. (EUMJ). [Ehime Pref.] 1 ex., Nishidani, Yanadani Village, 15–16–VII–1994, K. Aita leg. (EUMJ). [Kochi Pref.] 1 ex., Muroto Cape, Muroto City, 7–VI–1959, S. Hisamatsu leg. (EUMJ). [Nagasaki Pref.] 1 ex., Mt. Tatera-san, Tsushima Island 16–IX–1995, N. Narukawa leg. (ELKU); 3 exs., Midake, 12–X–1977, Y. Notsu leg. (EUMJ). [Kagoshima Pref.] 1 male & 4 exs., Kirishima, 29–VII–1971, T. Nakane leg. (HUSE); 2 exs., Shiratani-unsuikyô, Yakushima Island, 15–VIII–2005, J. Ogawa leg. (EUMJ); 1 male & 1 ex., Onoaida Hodô, Yakushima Island, 14–15–IX–2002, N. Ohbayashi leg. (EUMJ); 2 exs., Kosugidani, Yakushima Island, 2–V–1984, K. Mizuno leg. (OMNH); 4 exs., Anbô, Yakushima Island, 16–VII–1989, N. Narukawa leg. (ELKU); 5 exs., Nogi, Nishino-omote, Tanegashima Island, 10–VII–1974, S. Hisamatsu leg. (EUMJ).


JAPAN: Hokkaido, Honshu, Shikoku, Kyushu, Mikura (Izu Islands), Awaji, Yakushima and Tanegashima Islands; Burma; Korea; Russia.

Biological notes

The first author (Yoshida) collected this species from various kinds of dead leaves from evergreen trees, deciduous trees and bamboo.


The syntypes of this species consist of three specimens: two of them belong to this species, but the remaining one represents Psammoecus trimaculatus or Psammoecus triguttatus. We designate a male specimen as lectotype, and a female specimen as paralectotype. The type specimens were collected from Mt. Maya-san, Kobe City, Hyogo.

Taxon Treatment

  • Yoshida, T; Hirowatari, T; 2014: A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae) ZooKeys, 403: 15-45. doi

Other References

  1. Hetschko A (1930) Fam. Cucujidae. In: Coleopterorum Catalogus Pars 109. W. Junk, Berlin, 1–90.
  2. Nakane T (1963) Silvanidae. In: Nakane T Ohbayashi K Nomura S Kurosawa Y (Eds) Iconographia Insectorum Japonicorum Colore naturali edita. Vol. 2 (Coleoptera), Hokuryukan, Tokyo, 195–196 (incl. pl. 98). [In Japanese]
  3. Hisamatsu S (1977) Some notes of clavicorn beetles from Hiroshima Prefecture (Hiroshima-ken san kyuukakugun kochu note). Hiroshima Mushi-no-kai Kaiho 16: 207-209. [In Japanese]
  4. Sasaji H (1985) Silvanidae. In: Kurosawa Y Hisamatsu S Sasaji H (Eds) The Coleoptera of Japan in Color Vol. III. Hoikusha Publishing, Osaka, 202–205 (incl. pl. 32). [In Japanese, with English title]
  5. Sato M (1989) Coleoptera. In: Hirashima Y (Ed) A Check List of Japanese Insects I. Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka, 197-538. [In Japanese]
  6. Halstead D, Löbl I, Jelínek J (2007) Silvanidae. In: Löbl I Smetana A (Eds) Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea – Derodontoidea – Bostrichoidea – Lymexyloidea – Cleroidea – Cucujoidea. Apollo Books, Stenstrup, 496-501.
  7. Hirano Y (2009) Notes on Japanese Silvanidae (Nihonsan hosohiratamushi-ka ni tsuite). Kanagawa-chûhô 168: 57-83. [In Japanese]
  8. Hirano Y (2010) Cucujoidea of Japan Vol.2 Silvanidae, Byturidae, Biphyllidae. Roppon-Ashi Entomological Books, Tokyo, 61 pp. [In Japanese, with English title]
  9. Yoshida T, Hirowatari T (2013) A New Species of the Genus Psammoecus (Coleoptera, Silvanidae) from the Nansei Islands, Japan. Japanese Journal of Systematic Entomology 19: 85-90.