Cymatodera pueblae

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Burke A, Zolnerowich G (2014) Four new species of Cymatodera Gray from Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 387 : 33–49, doi. Versioned wiki page: 2014-03-11, version 42550, , contributors (alphabetical order): Pensoft Publishers.

Citation formats to copy and paste


author = {Burke, Alan F. AND Zolnerowich, Gregory},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {Four new species of Cymatodera Gray from Mexico (Coleoptera, Cleridae, Tillinae)},
year = {2014},
volume = {387},
issue = {},
pages = {33--49},
doi = {10.3897/zookeys.387.6827},
url = {},
note = {Versioned wiki page: 2014-03-11, version 42550, , contributors (alphabetical order): Pensoft Publishers.}


RIS/ Endnote:

T1 - Four new species of Cymatodera Gray from Mexico (Coleoptera, Cleridae, Tillinae)
A1 - Burke A
A1 - Zolnerowich G
Y1 - 2014
JF - ZooKeys
JA -
VL - 387
IS -
UR -
SP - 33
EP - 49
PB - Pensoft Publishers
M1 - Versioned wiki page: 2014-03-11, version 42550, , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/zookeys.387.6827

Wikipedia/ Citizendium:

<ref name="Burke2014ZooKeys387">{{Citation
| author = Burke A, Zolnerowich G
| title = Four new species of Cymatodera Gray from Mexico (Coleoptera, Cleridae, Tillinae)
| journal = ZooKeys
| year = 2014
| volume = 387
| issue =
| pages = 33--49
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.387.6827
| url =
| pmc =
| accessdate = 2022-08-18

}} Versioned wiki page: 2014-03-11, version 42550, , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Ordo: Coleoptera
Familia: Cleridae
Genus: Cymatodera


Cymatodera pueblae Burke sp. n.Wikispecies linkZooBank linkPensoft Profile

Type material

Holotype: male, Mexico, Puebla, Highway 18, 6 miles S Esperanza, 14-V-1983, 8100 ft, L. O’Brien and G. B. Marshall, printed red label, holotype deposited in CSCA. Paratypes: 2 males and 5 females. 1 male: México, Puebla, San Esteban Necoxcalco, 15-X-1992, C. Mayorga (CNIN); 1 male: Mexico, Mexico, km 41 highway Texcoco – Calpulalpan, 2685 m, 19 30 43 N 98 52 40 W, 3-VII-2001, beating oak, R. L. Westcott (JNRC); 1 female: México, Puebla, km 50 Plan de San Miguel, carretera Huajuapan de León - Oaxaca, 14-VIII-1992, C. Mayorga and E. Barrera (CNIN); 1 female: México, Puebla, Nuevo Vicencio, km 50 Carretera El Seco – Amozoc, 20-V-1995, G. Ortega and E. Barrera (CNIN); 3 females: Mexico, Pachuca, H90, 8000 ft, 6, 9-VII-1937, M. A. Embury (WFBM).

Differential diagnosis

Cymatodera pueblae is readily distinguished from similar species and other congeners by its small size, shape, sinuate midelytral fascia (Fig. 2), male terminalia (Figs 18–19), genitalia (Fig. 12), and geographic distribution (Fig. 34). No other species has this combination of characters.


Holotype. Small, rather robust, posterior wings present, brachypterous, TL = 8.67 mm. Color: Head fuscous; pronotum, prosternum, mesosternum, mestasternum, legs and elytra brown; abdomen testaceous mesally, becoming brown toward sides; mouthparts pale testaceous. Each elytron bearing a pale testaceous, median fascia that extends from first stria to epipleuron (Fig. 2).
Head: HL = 1.15 mm, HW = 1.88 mm. Measured across eyes conspicuously wider than pronotum; rugose; frons bi-impressed; strongly, coarsely punctate; clothed with short, semirecumbent setae intermixed with long, semierect and erect setae; eyes moderately large, subsinuate, longer than wide, moderately emarginate in front, feebly bulging laterally, separated by approximately 3.5 eye-widths. Antennae slender; loosely composed; extending slightly beyond posterior margin of elytra; antennomeres 2–3 subequal in length; fourth antennomere slightly longer than third antennomere; antennomeres 4–5 subequal in length; antennomeres 6–10 subequal in length, each slightly shorter than fifth antennomere; antennomeres 5–10 weakly serrate; last antennomere flattened apically, as long as tenth antennomere (Fig. 7).
Thorax: PL = 2.35 mm, PW = 1.67 mm. Pronotum elongate; widest at middle; middle slightly broader than anterior margin; sides constricted subapically; somewhat more constricted behind middle; disc flat; very feebly impressed in front of middle; subbasal tumescence pronounced; surface rugose; moderately coarsely punctate; somewhat vested with short, recumbent setae, intermingled with less numerous, long erect setae. Prosternum wider than long, rugulose, puncticulate. Mesosternum coarsely, deeply punctate. Metasternum convex; reduced in length, rugulose, moderately shallowly punctate.
Legs: Femora clothed with short, recumbent setae interspersed with few erect and semierect setae; tibiae vested with short and long erect and semierect setae; femorae and tibiae transversely, moderately rugose.
Elytra: EL = 5.17 mm, EW = 2.8 mm. Anterior margin arcuately emarginate; narrower than widest portion of pronotum; humeri very feebly indicated; sides subovoid; widest behind middle; disc convex; apex rounded, broadly dehiscent; surface smooth, clothed with intermixed setae of three sizes; sixth tergite exposed dorsally; sculpturing consisting of regular, rather coarse and deep striae that gradually reduce in size after first third of elytral length, striae not reaching posterior 1/4 of elytral length; interstices about 1.5× the width of punctation at elytral base.
Abdomen: Ventrites 1–5 rugulose, shallowly, moderately punctate; each segment with a pair of large, shallow impressions near sides; clothed with short, recumbent setae interspersed with less numerous, long, semi-erect setae. Fifth visible ventrite convex; sides oblique; posterior margin broadly, very deeply emarginate, emargination extends beyond first third of ventrite length (Fig. 18); sixth visible ventrite subquadrate; rugulose; surface strongly concave, excavated; moderately coarsely punctate; lateral margins subparallel; posterior margin broadly, very deeply emarginate, emargination extends to near base of segment; posterolateral angles arcuate, recurved ventrally. Fifth tergite moderately convex; rugulose; lateral margins subparallel; posterior margin broadly, shallowly, triangularly emarginate (Fig. 19). Sixth tergite subtriangular; surface strongly convex, smooth, shiny, shallowly punctate; lateral margin oblique; posterior margin subtruncate, narrowly, very shallowly, triangularly emarginate. Sixth tergite extending beyond sixth visible ventrite, fully covering it from dorsal view. Aedeagus: 1.4 mm long; ratio of length of parameres to whole tegmen 0.68: 1; tegmen robust, fully covering phallus; parameres robust, pointed at apex, lateral margins moderately oblique, procurved, posterior portion feebly curved ventrally; phallobase broad; phallus with copulatory piece somewhat acuminate distally; phallic plate armed with a row of long, posteriorly pointed denticles along dorsal margin; phallobasic apodeme short, robust, dilated distally; endophallic struts slender throughout their length (Fig. 12).
Females differ from males by having the posterior margin of the fifth visible ventrite longitudinally truncate; the sixth visible ventrite is rather convex, rugose and broader than long, the lateral margins are strongly oblique, giving the appearance of a broadly rounded margin (Fig. 27); posterior margin of fifth tergite broadly, very shallowly, triangularly emarginate; sixth tergite subtriangular, rugulose, surface moderately convex, broader than long, lateral and posterior margins strongly oblique, forming a semicircular perimeter.


Length of males 7.9–9.65 mm, length of females 8.85–10.2 mm; n = 5. Length to width ratio of head: males average 0.71, of females 0.83. Length to width ratio of thorax: males average 1.42, females average 1.6. Length to width ratio of elytra: males average 1.82, females average 1.94. One male and one female are darker than the rest of the type series; the mid-elytral fascia of one female is paler and the legs display a paler, yellowish coloration.


The type series of this species was collected at different localities in south-central Mexico, a high-altitude region characterized by various ranges surrounded by semiarid plateaus. Two males and two females were collected in the central and south region of the state of Puebla, Mexico; three females were collected in the vicinity of Pachuca, Hidalgo, Mexico; and one female was collected in km 41, highway Texcoco - Calpulalpan in the State of Mexico, a region that borders the central-western portion of the state of Puebla. The vegetation type in all collecting localities is predominantly a low to mid-altitude mixture of Quercus-Juniperus-Cupressus spp. in association with thorny species (Fig. 34).


The specific epithet refers to Puebla, the Mexican state where the holotype was collected.

Original Description

  • Burke, A; Zolnerowich, G; 2014: Four new species of Cymatodera Gray from Mexico (Coleoptera, Cleridae, Tillinae) ZooKeys, 387: 33-49. doi