Coloborhynchus clavirostris

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Rodrigues T, Kellner A (2013) Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England. ZooKeys 308 : 1–112, doi. Versioned wiki page: 2013-06-12, version 35256, , contributors (alphabetical order): Pensoft Publishers.

Citation formats to copy and paste


author = {Rodrigues, Taissa AND Kellner, Alexander Wilhelm Armin},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England},
year = {2013},
volume = {308},
issue = {},
pages = {1--112},
doi = {10.3897/zookeys.308.5559},
url = {},
note = {Versioned wiki page: 2013-06-12, version 35256, , contributors (alphabetical order): Pensoft Publishers.}


RIS/ Endnote:

T1 - Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England
A1 - Rodrigues T
A1 - Kellner A
Y1 - 2013
JF - ZooKeys
JA -
VL - 308
IS -
UR -
SP - 1
EP - 112
PB - Pensoft Publishers
M1 - Versioned wiki page: 2013-06-12, version 35256, , contributors (alphabetical order): Pensoft Publishers.

M3 - doi:10.3897/zookeys.308.5559

Wikipedia/ Citizendium:

<ref name="Rodrigues2013ZooKeys308">{{Citation
| author = Rodrigues T, Kellner A
| title = Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England
| journal = ZooKeys
| year = 2013
| volume = 308
| issue =
| pages = 1--112
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.308.5559
| url =
| pmc =
| accessdate = 2019-06-20

}} Versioned wiki page: 2013-06-12, version 35256, , contributors (alphabetical order): Pensoft Publishers.</ref>

See also the citation download page at the journal.


Ordo: Pterosauria
Familia: Anhangueridae
Genus: Coloborhynchus


Coloborhynchus clavirostris Owen, 1874Wikispecies linkPensoft Profile

  • Coloborhynchus clavirostris Owen: Owen 1874[1]: p. 6, pl. I, fig. 1–4
  • Criorhynchus simus (Owen): Hooley 1914[2]: p. 537
  • Criorhynchus clavirostris (Owen): Arthaber 1922[3]: p. 18, fig. 7c
  • Criorhynchus simus (Owen): Kuhn 1967[4]: 38
  • Criorhynchus simus (Owen): Wellnhofer 1978[5]: p. 60
  • Coloborhynchus clavirostris Owen: Lee 1994[6]: p. 756, fig. 4
  • Coloborhynchus clavirostris Owen: Unwin 2001[7]: p. 206
  • Coloborhynchus clavirostris Owen: Veldmeijer 2003[8]: 42
  • Coloborhynchus clavirostris Owen: Rodrigues and Kellner 2008[9]: p. 220, fig. 1.1, 2.1, 3.1
  • Coloborhynchus clavirostris Owen: Martill, Sweetman and Witton 2011[10]: p. 380, fig. 25.8


NHMUK PV R 1822, anterior portion of the rostrum (Fig. 7A–D).

Type locality

St.–Leonards–on–Sea, East Sussex, England.

Type horizon

Hastings Group (late Berriasian / Valanginian).


Anhanguerid pterosaur with the following combination of characters that distinguishes it from other members of the clade (autapomorphies are marked with an asterisk): oval depression beneath the first pair of alveoli*; second, third and fourth pairs of alveoli located laterally*; fifth and sixth pairs of alveoli located more medially than the preceding alveoli on the base of the palatal ridge*; anterior part of the palatal ridge bordered by two shallow longitudinally elongated depressions* (from Rodrigues and Kellner 2008[9]).


The holotype of Coloborhynchus clavirostris (NHMUK PV R 1822) is a fragment of the premaxillae and maxillae, and has previously been described in detail (Owen 1874[1]; Veldmeijer 2003[8]; Rodrigues and Kellner 2008[9]). It has a flattened anterior margin, where the first pair of tooth sockets is located. The second, third and fourth pairs of alveoli face laterally, and the fifth and sixth pairs are situated closer to the midline. Coloborhynchus clavirostris also has an anterior expansion and a strong palatal ridge in addition a robust premaxillary crest, which begins at the tip of the rostrum.


Rodrigues and Kellner (2008)[9] reviewed the taxonomic history of the genus Coloborhynchus and the species Coloborhynchus clavirostris, and thus only the main points are repeated here. The genus and species were erected by Owen (1874)[1]. Later authors such as Hooley (1914)[2], Kuhn (1967)[4] and Wellnhofer (1978)[5] regarded it synonymous with Ornithocheirus simus [=Criorhynchus simus]. Lee (1994)[6] revalidated Coloborhynchus, and Unwin (2001)[7] not only considered it a valid genus, but also referred additional species to the genus. Rodrigues and Kellner (2008)[9], noting several unique characters of Coloborhynchus clavirostris, restricted the genus to the type species. This view is followed here.

Taxon Treatment

  • Rodrigues, T; Kellner, A; 2013: Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England ZooKeys, 308: 1-112. doi

Other References

  1. 1.0 1.1 1.2 Owen R (1874) Monograph on the fossil Reptilia of the Mesozoic Formations. Palaeontographical Society, London, 14 pp.
  2. 2.0 2.1 Hooley R (1914) On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529-557. doi: 10.1080/00222931408693521
  3. Arthaber G (1922) Über Entwicklung, Ausbildung und Austerben der Flugsaurier. Palaeontologische Zeitschrift 4: 1-47.
  4. 4.0 4.1 Kuhn O (1967) Die fossile Wirbeltierklasse Pterosauria. Verlag Oeben, Krailling bei München, 52 pp.
  5. 5.0 5.1 Wellnhofer P (1978) Pterosauria. Handbuch der Paläoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp.
  6. 6.0 6.1 Lee Y (1994) The Early Cretaceous pterodactyloid pterosaur Coloborhynchus from North America. Palaeontology 37: 755-763.
  7. 7.0 7.1 Unwin D (2001) An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189-221. doi: 10.1002/mmng.20010040112
  8. 8.0 8.1 Veldmeijer A (2003) Preliminary description of a skull and wing of a Brazilian Cretaceous (Santana Formation; Aptian-Albian) pterosaur (Pterodactyloidea) in the collection of the AMNH. PalArch’s Journal of Vertebrate Palaeontology 0: 1-14.
  9. 9.0 9.1 9.2 9.3 9.4 Rodrigues T, Kellner A (2008) Review of the pterodactyloid pterosaur Coloborhynchus. Zitteliana B 28: 219-228.
  10. Martill D, Sweetman S, Witton M (2011) Pterosaurs. In: Batten DJ (Ed). English Wealden fossils. The Palaeontological Association, London: 370-390.