Telestes dabar
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Ordo: Cypriniformes
Familia: Cyprinidae
Genus: Telestes
Name
Telestes dabar Bogutskaya & Zupančič & Bogut & Naseka, 2012 sp. n. – Wikispecies link – Pensoft Profile
Holotype
NMW 95295, 79.1 mm SL, BOSNIA & HERZEGOVINA: Dabarsko Polje, Opačica River at Potkom, 43°5.9'N, 18°7.6'E, 15 Sept. 2006, coll. Zupančič.
Paratypes
NMW 95300, 6, 55.2–71.1 mm SL, same data as holotype; PZC 525, 72, 32.5–51.3 mm SL, same data as holotype; PZC 526, 13, 48.0–73.8 mm SL, same data as holotype; PZC 565, 21, 35.5–56.7 mm SL, same locality, 8 July 2011; ZISP 54995, 15, 38.1–58.9 mm SL, same locality, 8 July 2011; SMNH 444, 35.5–60.8 mm SL, same locality, 8 July 2011; PZC 279, 12, 44.2–71.9 mm SL, BOSNIA & HERZEGOVINA: Dabarsko Polje, Vrijeka River, 24 May 2001; PZC 521, 13, 54.0–62.6 mm SL, same locality as 279, 15 Sept. 2004; PZC 575, 18, 40.5–69.8 mm SL, same locality as 279, 15 Sept. 2006.
Diagnosis
Telestes dabar is distinguished from Telestes metohiensis and Telestes miloradi by having the following combination of characters: slightly curved dark stripe (obvious in live and preserved specimens) present from just behind operculum to vertical just anterior to origin of anal fin, this stripe narrow and separated from dark pigmented area on back along its entire length; scales on most of body not overlapping but situated close to one another; scales overlapping behind pectoral girdle along lateral line and usually on caudal peduncle; snout with fleshy tip projecting over upper lip; mouth subterminal with tip of mouth cleft at or below level of ventral margin of eye; lateral line usually interrupted, with 24–69 total lateral-line scales; branched dorsal-fin rays usually 8½; branched anal-fin rays usually 8½; gill rakers 9 or 10; total vertebrae 39–41, mode 40; abdominal vertebrae 22–24, mode 22; caudal vertebrae 16–18, mode 17; head width 43–52% HL; and lower jaw long, length 10–12% SL.
Description
Morphometric data are summarised in Table 1a, selected counts in Tables 2–4. General appearance can be seen in Figs 1a and 2a. Body compressed, elongate. Caudal-peduncle depth only slightly less than half maximum body depth; head length greater than maximum body depth. Eye small, its diameter smaller than snout length. Snout fleshy, slightly to markedly projecting beyond upper lip (similar to a feature Kottelat and Freyhof [2007: fig. 39] called the “rostral cap,” which covers all or part of upper lip); snout terminating laterally in prominent crease along anterior edge of first infraorbital. Mouth subterminal, tip of mouth cleft at level of ventral margin of eye or, more frequently, below it. Lower jaw-quadrate junction at vertical through anterior half of eye. Length of lower jaw 10–12% SL or 36–41% HL, or 102–132% depth of operculum.
| Telestes dabar | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| Holotype | females, n=26 | males, n=13 | |||||||
| min | max | mean | SD | min | max | M | SD | ||
| SL, mm | 79.1 | 60.3 | 81.8 | 67.8 | 55.5 | 69.0 | 61.1 | ||
| Maximum body depth (% SL) | 21.6 | 21.6 | 26.0 | 23.6 | 1.3 | 21.7 | 25.0 | 22.6 | 0.9 |
| Depth of caudal peduncle (% SL) | 9.8 | 9.8 | 12.4 | 11.0 | 0.7 | 10.0 | 12.4 | 11.2 | 0.7 |
| Depth of caudal peduncle (% length of caudal peduncle) | 47.9 | 47.7 | 59.6 | 53.6 | 4.3 | 44.6 | 62.2 | 52.9 | 5.2 |
| Maximum body width (% SL) | 13.4 | 11.4 | 16.0 | 13.7 | 1.2 | 12.0 | 22.0 | 14.2 | 2.6 |
| Predorsal length (% SL) | 55.3 | 53.5 | 58.7 | 56.9 | 1.1 | 53.8 | 57.3 | 55.5 | 1.2 |
| Postdorsal length (% SL) | 36.3 | 32.8 | 36.6 | 35.2 | 1.0 | 34.4 | 37.2 | 35.8 | 1.0 |
| Prepelvic length (% SL) | 51.6 | 50.4 | 55.5 | 52.7 | 1.4 | 50.6 | 54.4 | 52.2 | 1.0 |
| Preanal length (% SL) | 71.2 | 68.3 | 73.2 | 70.8 | 1.3 | 68.6 | 71.9 | 69.9 | 1.0 |
| Pectoral – pelvic-fin origin length (% SL) | 24.5 | 24.5 | 27.7 | 25.6 | 0.9 | 22.3 | 26.5 | 24.1 | 1.2 |
| Pelvic – anal-fin origin length (% SL) | 20.2 | 17.2 | 20.2 | 19.0 | 0.6 | 16.6 | 20.3 | 18.4 | 1.3 |
| Length of caudal peduncle (% SL) | 20.6 | 18.5 | 21.9 | 20.5 | 0.8 | 19.2 | 23.5 | 21.2 | 1.1 |
| Dorsal-fin base length (% SL) | 10.3 | 9.3 | 12.3 | 10.9 | 0.8 | 9.9 | 12.2 | 11.2 | 0.7 |
| Dorsal fin depth (% SL) | 18.9 | 16.6 | 20.2 | 18.6 | 1.4 | 18.5 | 21.8 | 19.7 | 0.9 |
| Anal-fin base length (% SL) | 12.1 | 9.2 | 12.1 | 10.6 | 0.8 | 10.3 | 12.4 | 11.2 | 0.5 |
| Anal fin depth (% SL) | 12.3 | 11.9 | 14.8 | 13.2 | 0.8 | 13.4 | 17.2 | 15.9 | 1.1 |
| Pectoral fin length (% SL) | 19.9 | 18.1 | 21.2 | 19.9 | 0.8 | 22.5 | 26.7 | 24.2 | 1.3 |
| Pelvic fin length (% SL) | 14.8 | 12.9 | 16.3 | 14.9 | 0.9 | 16.3 | 17.7 | 16.9 | 0.5 |
| Head length (% SL) | 27.1 | 25.6 | 28.7 | 27.1 | 0.9 | 26.6 | 29.6 | 28.1 | 1.0 |
| Head length (% body depth) | 125.1 | 104.0 | 125.1 | 115.3 | 5.9 | 110.8 | 135.1 | 124.3 | 7.6 |
| Head depth at nape (% SL) | 17.4 | 17.1 | 19.3 | 18.1 | 0.7 | 17.0 | 18.8 | 18.2 | 0.5 |
| Head depth at nape (% HL) | 64.2 | 62.4 | 70.7 | 66.8 | 2.3 | 60.6 | 69.3 | 65.0 | 2.7 |
| Maximum head width (% SL) | 13.5 | 11.6 | 14.6 | 13.6 | 0.7 | 12.4 | 14.5 | 13.6 | 0.6 |
| Maximum head width (% HL) | 50.0 | 42.8 | 52.3 | 50.1 | 2.4 | 44.7 | 52.1 | 48.3 | 2.4 |
| Maximum cranial width (% cranium roof length) | 68.9 | 61.9 | 72.6 | 65.8 | 3.1 | 59.7 | 68.8 | 64.4 | 2.5 |
| Snout length (% SL) | 8.5 | 7.4 | 9.2 | 8.1 | 0.5 | 7.8 | 9.0 | 8.5 | 0.4 |
| Snout length (% HL) | 31.4 | 27.7 | 32.8 | 29.7 | 1.3 | 28.5 | 31.7 | 30.3 | 1.1 |
| Eye horizontal diameter (% SL) | 6.6 | 5.9 | 7.2 | 6.6 | 0.4 | 6.3 | 7.8 | 6.8 | 0.4 |
| Eye horizontal diameter (% HL) | 24.4 | 22.1 | 26.8 | 24.5 | 1.5 | 22.1 | 26.8 | 24.3 | 1.4 |
| Eye horizontal diameter (% interorbital width) | 74.5 | 66.3 | 81.9 | 73.3 | 4.7 | 69.5 | 82.0 | 74.4 | 4.7 |
| Postorbital distance (% HL) | 53.0 | 48.8 | 54.0 | 52.2 | 1.4 | 47.1 | 53.0 | 50.2 | 1.9 |
| Interorbital width (% SL) | 8.9 | 8.2 | 9.6 | 9.1 | 0.4 | 8.5 | 9.9 | 9.2 | 0.4 |
| Interorbital width (% HL) | 32.8 | 31.3 | 36.0 | 33.4 | 1.2 | 29.0 | 35.6 | 32.7 | 1.9 |
| Length of upper jaw (% HL) | 28.6 | 24.5 | 30.5 | 27.8 | 1.9 | 26.2 | 29.5 | 27.7 | 1.1 |
| Length of upper jaw (% SL) | 7.7 | 6.6 | 8.5 | 7.5 | 0.6 | 7.0 | 8.6 | 7.8 | 0.5 |
| Length of lower jaw (% SL) | 10.6 | 9.5 | 11.2 | 10.2 | 0.5 | 9.7 | 11.5 | 10.9 | 0.5 |
| Length of lower jaw (% HL) | 39.2 | 35.8 | 40.7 | 37.6 | 1.3 | 35.6 | 41.1 | 38.6 | 1.6 |
| Length of lower jaw (% interorbital width) | 119.7 | 106.4 | 123.4 | 112.6 | 4.4 | 110.2 | 129.8 | 120.2 | 6.4 |
| Length of lower jaw (% depth of operculum) | 110.2 | 102.4 | 120.0 | 108.2 | 4.5 | 105.7 | 131.7 | 116.6 | 8.2 |
| Telestes miloradi | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| Holotype | females, n=9 | males, n=3 | |||||||
| min | max | mean | SD | min | max | Mean | SD | ||
| SL, mm | 66.7 | 34.1 | 66.7 | 57.9 | 61.8 | ||||
| Maximum body depth (% SL) | 22.4 | 21.7 | 25.8 | 23.5 | 1.5 | 19.6 | 24.6 | 23.0 | 2.9 |
| Depth of caudal peduncle (% SL) | 11.4 | 10.4 | 11.6 | 11.2 | 0.4 | 9.6 | 11.3 | 10.8 | 1.0 |
| Depth of caudal peduncle (% length of caudal peduncle) | 53.8 | 49.8 | 54.8 | 52.4 | 2.1 | 41.8 | 52.8 | 49.1 | 6.4 |
| Maximum body width (% SL) | 14.3 | 12.9 | 15.3 | 14.6 | 0.7 | 11.4 | 16.8 | 15.0 | 3.1 |
| Predorsal length (% SL) | 55.9 | 55.9 | 57.7 | 57.0 | 0.7 | 54.3 | 54.9 | 54.5 | 0.3 |
| Postdorsal length (% SL) | 35.2 | 32.8 | 35.3 | 34.3 | 1.0 | 35.4 | 35.6 | 35.5 | 0.1 |
| Prepelvic length (% SL) | 51.6 | 51.5 | 53.3 | 52.3 | 0.9 | 50.0 | 50.2 | 50.1 | 0.1 |
| Preanal length (% SL) | 72.8 | 68.2 | 72.8 | 70.3 | 1.9 | 68.2 | 68.4 | 68.3 | 0.1 |
| Pectoral – pelvic-fin origin length (% SL) | 25.1 | 25.1 | 27.6 | 26.2 | 1.1 | 23.3 | 25.0 | 24.4 | 1.0 |
| Pelvic – anal-fin origin length (% SL) | 20.8 | 16.6 | 20.8 | 18.3 | 1.7 | 17.6 | 18.4 | 18.2 | 0.5 |
| Length of caudal peduncle (% SL) | 21.2 | 20.5 | 22.1 | 21.3 | 0.7 | 21.4 | 23.0 | 22.0 | 0.9 |
| Dorsal-fin base length (% SL) | 11.6 | 9.9 | 11.6 | 10.9 | 0.7 | 11.2 | 12.6 | 12.2 | 0.8 |
| Dorsal fin depth (% SL) | 19.6 | 19.2 | 20.9 | 19.8 | 0.8 | 20.9 | 22.1 | 21.4 | 0.6 |
| Anal-fin base length (% SL) | 12.4 | 10.7 | 12.4 | 11.9 | 0.6 | 11.8 | 11.9 | 11.8 | 0.1 |
| Anal fin depth (% SL) | 13.2 | 0.0 | 15.0 | 12.5 | 4.7 | 13.7 | 15.5 | 14.3 | 1.0 |
| Pectoral fin length (% SL) | 21.7 | 18.5 | 21.7 | 20.8 | 1.0 | 23.5 | 25.4 | 24.8 | 1.1 |
| Pelvic fin length (% SL) | 15.6 | 13.6 | 15.6 | 14.6 | 0.8 | 16.9 | 17.6 | 17.2 | 0.4 |
| Head length (% SL) | 26.8 | 25.4 | 28.0 | 27.0 | 0.8 | 26.5 | 29.0 | 28.2 | 1.4 |
| Head length (% body depth) | 119.9 | 105.0 | 119.9 | 115.0 | 6.0 | 117.6 | 135.4 | 123.5 | 10.3 |
| Head depth at nape (% SL) | 18.3 | 16.9 | 18.3 | 17.6 | 0.5 | 16.7 | 19.0 | 18.2 | 1.4 |
| Head depth at nape (% HL) | 68.1 | 62.5 | 68.1 | 65.1 | 2.3 | 62.9 | 65.7 | 64.7 | 1.6 |
| Maximum head width (% SL) | 14.1 | 12.3 | 14.5 | 14.0 | 0.7 | 12.6 | 16.9 | 15.4 | 2.5 |
| Maximum head width (% HL) | 52.5 | 48.7 | 53.6 | 52.0 | 1.5 | 47.5 | 58.2 | 54.6 | 6.2 |
| Maximum cranial width (% cranium roof length) | 62.8 | 59.6 | 73.9 | 66.0 | 5.0 | 58.3 | 65.2 | 62.9 | 4.0 |
| Snout length (% SL) | 9.0 | 8.0 | 9.7 | 8.9 | 0.5 | 8.5 | 8.6 | 8.6 | 0.0 |
| Snout length (% HL) | 33.7 | 31.4 | 34.7 | 33.0 | 1.2 | 29.5 | 32.4 | 30.5 | 1.7 |
| Eye horizontal diameter (% SL) | 6.5 | 6.3 | 6.9 | 6.5 | 0.2 | 6.6 | 6.7 | 6.7 | 0.0 |
| Eye horizontal diameter (% HL) | 24.4 | 22.5 | 26.4 | 24.2 | 1.4 | 22.9 | 25.2 | 23.7 | 1.3 |
| Eye horizontal diameter (% interorbital width) | 70.2 | 64.7 | 83.0 | 71.3 | 5.9 | 69.4 | 83.4 | 74.1 | 8.1 |
| Postorbital distance (% HL) | 49.1 | 48.9 | 51.6 | 50.2 | 1.2 | 50.2 | 51.0 | 50.5 | 0.5 |
| Interorbital width (% SL) | 9.3 | 8.1 | 9.7 | 9.2 | 0.5 | 8.0 | 9.6 | 9.1 | 0.9 |
| Interorbital width (% HL) | 34.8 | 31.9 | 34.8 | 34.1 | 0.9 | 30.2 | 33.1 | 32.1 | 1.7 |
| Length of upper jaw (% HL) | 27.9 | 26.0 | 28.1 | 27.2 | 0.9 | 23.5 | 25.1 | 24.0 | 0.9 |
| Length of upper jaw (% SL) | 7.5 | 6.6 | 7.9 | 7.3 | 0.4 | 6.7 | 6.8 | 6.8 | 0.1 |
| Length of lower jaw (% SL) | 9.3 | 8.4 | 10.4 | 9.5 | 0.6 | 8.7 | 10.4 | 9.8 | 1.0 |
| Length of lower jaw (% HL) | 34.7 | 33.1 | 38.4 | 35.3 | 1.8 | 32.9 | 35.9 | 34.9 | 1.7 |
| Length of lower jaw (% interorbital width) | 99.8 | 99.8 | 114.3 | 103.7 | 6.1 | 108.5 | 109.1 | 108.7 | 0.4 |
| Length of lower jaw (% depth of operculum) | 104.0 | 95.6 | 104.0 | 101.1 | 2.9 | 95.6 | 107.1 | 103.3 | 6.7 |
| Telestes metohiensis, Gatačko and Cerničko poljes | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| lectotype | females, n=25 | males, n=10 | |||||||
| min | max | mean | SD | min | max | mean | SD | ||
| SL, mm | 87.9 | 60.26 | 102.1 | 74.9 | 54.4 | 75.7 | 63.1 | ||
| Maximum body depth (% SL) | 23.9 | 19.9 | 25.1 | 22.5 | 1.4 | 20.7 | 24.1 | 22.5 | 1.0 |
| Depth of caudal peduncle (% SL) | 11.3 | 9.7 | 11.9 | 10.7 | 0.5 | 10.4 | 11.8 | 11.2 | 0.5 |
| Depth of caudal peduncle (% length of caudal peduncle) | 61.5 | 46.7 | 61.5 | 53.0 | 4.1 | 46.7 | 55.7 | 51.0 | 3.1 |
| Maximum body width (% SL) | 16.0 | 11.1 | 16.4 | 14.3 | 1.7 | 14.1 | 15.6 | 14.8 | 0.6 |
| Predorsal length (% SL) | 57.6 | 49.4 | 60.1 | 56.5 | 2.2 | 53.7 | 57.7 | 55.7 | 1.3 |
| Postdorsal length (% SL) | 33.4 | 31.0 | 36.5 | 33.7 | 1.6 | 32.5 | 38.0 | 35.2 | 1.6 |
| Prepelvic length (% SL) | 52.0 | 46.0 | 55.9 | 52.7 | 1.9 | 50.4 | 53.8 | 51.9 | 1.3 |
| Preanal length (% SL) | 69.1 | 65.6 | 73.6 | 70.2 | 1.9 | 67.1 | 69.7 | 68.5 | 0.9 |
| Pectoral – pelvic-fin origin length (% SL) | 27.5 | 24.7 | 29.7 | 27.0 | 1.3 | 24.1 | 27.2 | 25.5 | 1.1 |
| Pelvic – anal-fin origin length (% SL) | 19.8 | 16.4 | 20.1 | 18.5 | 1.1 | 15.6 | 18.2 | 17.0 | 0.8 |
| Length of caudal peduncle (% SL) | 18.3 | 19.1 | 22.3 | 20.3 | 1.2 | 20.9 | 22.8 | 22.0 | 0.6 |
| Dorsal-fin base length (% SL) | 11.1 | 9.5 | 11.5 | 10.8 | 0.6 | 10.8 | 13.6 | 12.0 | 1.0 |
| Dorsal fin depth (% SL) | 17.6 | 14.9 | 18.3 | 17.2 | 0.8 | 16.3 | 19.6 | 18.5 | 1.1 |
| Anal-fin base length (% SL) | 11.3 | 9.1 | 12.0 | 10.9 | 0.7 | 10.0 | 13.2 | 11.9 | 1.1 |
| Anal fin depth (% SL) | 12.3 | 12.4 | 15.3 | 13.7 | 0.8 | 12.8 | 15.4 | 14.3 | 1.0 |
| Pectoral fin length (% SL) | 18.4 | 17.0 | 19.8 | 18.4 | 0.8 | 22.7 | 25.3 | 23.6 | 0.8 |
| Pelvic fin length (% SL) | 12.9 | 12.4 | 15.3 | 13.7 | 0.8 | 15.4 | 17.7 | 16.6 | 0.7 |
| Head length (% SL) | 27.0 | 23.8 | 28.2 | 26.6 | 1.1 | 26.1 | 28.2 | 27.3 | 0.6 |
| Head length (% body depth) | 112.9 | 111.0 | 132.5 | 120.4 | 6.6 | 114.4 | 131.8 | 122.8 | 5.2 |
| Head depth at nape (% SL) | 17.4 | 15.5 | 18.7 | 17.3 | 0.8 | 15.9 | 18.1 | 17.3 | 0.7 |
| Head depth at nape (% HL) | 64.6 | 62.9 | 71.8 | 65.1 | 2.2 | 58.5 | 66.1 | 63.5 | 2.7 |
| Maximum head width (% SL) | 14.8 | 12.9 | 15.7 | 14.8 | 0.7 | 14.5 | 15.7 | 14.9 | 0.4 |
| Maximum head width (% HL) | 54.9 | 50.6 | 59.3 | 55.7 | 2.7 | 52.8 | 56.9 | 54.7 | 1.6 |
| Maximum cranial width (% cranium roof length) | 63.5 | 62.9 | 78.6 | 68.6 | 4.4 | 65.4 | 71.7 | 68.2 | 2.7 |
| Snout length (% SL) | 8.1 | 7.3 | 8.8 | 8.2 | 0.4 | 8.5 | 9.4 | 8.9 | 0.4 |
| Snout length (% HL) | 30.0 | 27.9 | 33.1 | 30.8 | 1.6 | 31.2 | 34.2 | 32.5 | 1.1 |
| Eye horizontal diameter (% SL) | 6.0 | 4.6 | 6.9 | 5.9 | 0.5 | 5.3 | 6.8 | 6.0 | 0.4 |
| Eye horizontal diameter (% HL) | 22.4 | 19.4 | 25.2 | 22.1 | 1.5 | 19.5 | 24.8 | 22.1 | 1.8 |
| Eye horizontal diameter (% interorbital width) | 70.7 | 57.0 | 80.2 | 68.2 | 6.9 | 60.2 | 71.8 | 65.1 | 4.3 |
| Postorbital distance (% HL) | 51.9 | 47.9 | 55.6 | 51.9 | 2.4 | 46.2 | 53.1 | 50.8 | 2.3 |
| Interorbital width (% SL) | 8.5 | 7.7 | 9.2 | 8.7 | 0.4 | 8.4 | 9.8 | 9.3 | 0.5 |
| Interorbital width (% HL) | 31.6 | 29.6 | 34.8 | 32.6 | 1.7 | 30.8 | 35.8 | 34.0 | 1.5 |
| Length of upper jaw (% HL) | 27.8 | 27.0 | 31.0 | 29.2 | 1.0 | 27.0 | 30.9 | 29.0 | 1.1 |
| Length of upper jaw (% SL) | 7.5 | 6.4 | 8.3 | 7.8 | 0.4 | 7.5 | 8.4 | 7.9 | 0.3 |
| Length of lower jaw (% SL) | 9.9 | 8.3 | 11.3 | 10.3 | 0.8 | 9.5 | 11.0 | 10.3 | 0.6 |
| Length of lower jaw (% HL) | 36.7 | 35.0 | 42.2 | 38.7 | 2.2 | 34.8 | 41.2 | 37.8 | 2.1 |
| Length of lower jaw (% interorbital width) | 116.0 | 106.5 | 132.5 | 120.6 | 6.4 | 101.6 | 121.9 | 113.7 | 6.9 |
| Length of lower jaw (% depth of operculum) | 107.2 | 106.5 | 117.9 | 112.5 | 3.0 | 109.6 | 119.2 | 112.7 | 3.5 |
| Telestes metohiensis (affinis nomen museale), Nevesinjsko Polje | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| lectotype | females, n=33 | males, n=14 | |||||||
| min | max | mean | SD | min | max | M | sd | ||
| SL, mm | 87.9 | 57.8 | 113.8 | 79.5 | 57.7 | 86.2 | 70.4 | ||
| Maximum body depth (% SL) | 23.9 | 19.3 | 24.1 | 21.8 | 1.1 | 19.8 | 23.8 | 21.6 | 1.4 |
| Depth of caudal peduncle (% SL) | 11.3 | 9.0 | 12.0 | 10.7 | 0.7 | 9.6 | 12.3 | 11.1 | 0.7 |
| Depth of caudal peduncle (% length of caudal peduncle) | 61.5 | 42.7 | 60.0 | 54.2 | 4.5 | 43.6 | 59.9 | 51.8 | 4.2 |
| Maximum body width (% SL) | 16.0 | 11.6 | 17.6 | 14.5 | 1.4 | 11.0 | 16.6 | 14.3 | 1.7 |
| Predorsal length (% SL) | 57.6 | 54.2 | 59.4 | 57.1 | 1.3 | 54.4 | 58.4 | 56.3 | 1.0 |
| Postdorsal length (% SL) | 33.4 | 29.9 | 36.4 | 34.0 | 1.3 | 33.8 | 36.5 | 35.2 | 0.7 |
| Prepelvic length (% SL) | 52.0 | 51.6 | 55.1 | 53.3 | 1.0 | 50.1 | 54.6 | 51.6 | 1.2 |
| Preanal length (% SL) | 69.1 | 67.8 | 73.4 | 70.9 | 1.3 | 65.9 | 70.3 | 68.8 | 1.3 |
| Pectoral – pelvic-fin origin length (% SL) | 27.5 | 24.2 | 30.0 | 25.9 | 1.2 | 21.7 | 25.4 | 23.8 | 1.0 |
| Pelvic – anal-fin origin length (% SL) | 19.8 | 16.7 | 44.1 | 19.4 | 4.6 | 15.6 | 19.9 | 17.5 | 1.3 |
| Length of caudal peduncle (% SL) | 18.3 | 18.0 | 21.3 | 19.7 | 0.9 | 20.0 | 23.2 | 21.4 | 0.8 |
| Dorsal-fin base length (% SL) | 11.1 | 9.4 | 12.9 | 11.1 | 0.8 | 8.1 | 12.1 | 10.4 | 1.2 |
| Dorsal fin depth (% SL) | 17.6 | 15.8 | 20.4 | 17.9 | 1.4 | 15.9 | 21.8 | 18.0 | 1.5 |
| Anal-fin base length (% SL) | 11.3 | 9.4 | 12.0 | 10.9 | 0.6 | 10.5 | 13.2 | 11.4 | 0.7 |
| Anal fin depth (% SL) | 12.3 | 10.6 | 15.0 | 13.1 | 1.1 | 12.0 | 16.2 | 14.0 | 1.4 |
| Pectoral fin length (% SL) | 18.4 | 18.0 | 21.9 | 20.2 | 1.0 | 21.8 | 26.4 | 23.8 | 1.4 |
| Pelvic fin length (% SL) | 12.9 | 12.5 | 15.6 | 14.1 | 0.8 | 14.2 | 17.5 | 15.7 | 0.9 |
| Head length (% SL) | 27.0 | 26.7 | 29.4 | 27.8 | 0.8 | 26.3 | 30.1 | 28.0 | 1.1 |
| Head length (% body depth) | 112.9 | 114.5 | 142.6 | 127.4 | 6.6 | 116.5 | 142.9 | 130.0 | 8.5 |
| Head depth at nape (% SL) | 17.4 | 16.5 | 18.7 | 17.6 | 0.6 | 15.7 | 18.8 | 17.3 | 0.9 |
| Head depth at nape (% HL) | 64.6 | 60.0 | 67.9 | 63.4 | 1.8 | 57.1 | 65.9 | 62.0 | 2.9 |
| Maximum head width (% SL) | 14.8 | 13.7 | 16.6 | 14.9 | 0.6 | 13.3 | 16.2 | 14.7 | 0.8 |
| Maximum head width (% HL) | 54.9 | 50.5 | 61.2 | 53.7 | 2.3 | 51.0 | 57.4 | 53.0 | 2.4 |
| Maximum cranial width (% cranium roof length) | 63.5 | 64.9 | 79.6 | 72.5 | 4.3 | 66.3 | 75.8 | 69.1 | 2.8 |
| Snout length (% SL) | 8.1 | 7.9 | 9.8 | 8.7 | 0.5 | 7.7 | 9.3 | 8.6 | 0.4 |
| Snout length (% HL) | 30.0 | 29.4 | 33.4 | 31.3 | 1.1 | 27.2 | 33.0 | 30.8 | 1.6 |
| Eye horizontal diameter (% SL) | 6.0 | 4.7 | 7.7 | 6.1 | 0.7 | 5.4 | 8.0 | 6.3 | 0.7 |
| Eye horizontal diameter (% HL) | 22.4 | 17.3 | 27.3 | 21.9 | 2.5 | 20.6 | 26.4 | 22.5 | 2.0 |
| Eye horizontal diameter (% interorbital width) | 70.7 | 52.0 | 86.7 | 67.5 | 8.3 | 59.8 | 92.5 | 73.1 | 11.9 |
| Postorbital distance (% HL) | 51.9 | 49.7 | 57.2 | 53.3 | 1.9 | 48.1 | 54.4 | 51.2 | 1.8 |
| Interorbital width (% SL) | 8.5 | 8.0 | 10.0 | 9.0 | 0.6 | 7.5 | 9.9 | 8.7 | 0.8 |
| Interorbital width (% HL) | 31.6 | 29.7 | 35.9 | 32.5 | 1.9 | 26.2 | 34.8 | 31.2 | 2.7 |
| Length of upper jaw (% HL) | 27.8 | 26.7 | 30.5 | 28.7 | 0.9 | 27.4 | 29.8 | 28.5 | 0.8 |
| Length of upper jaw (% SL) | 7.5 | 7.2 | 8.7 | 8.0 | 0.4 | 7.3 | 8.9 | 8.0 | 0.5 |
| Length of lower jaw (% SL) | 9.9 | 10.0 | 12.2 | 10.7 | 0.5 | 10.0 | 12.5 | 11.0 | 0.9 |
| Length of lower jaw (% HL) | 36.7 | 36.3 | 43.8 | 38.5 | 1.5 | 36.3 | 44.0 | 38.9 | 2.5 |
| Length of lower jaw (% interorbital width) | 116.0 | 110.0 | 133.8 | 121.2 | 6.0 | 109.1 | 138.7 | 123.0 | 9.8 |
| Length of lower jaw (% depth of operculum) | 116.5 | 110.8 | 133.8 | 116.1 | 4.9 | 111.5 | 132.3 | 118.5 | 7.0 |
Total gill rakers (Table 2) 9 (20) or 10 (20), 10 in holotype. Pharyngeal teeth 5–4, hooked, slightly serrated (examined in 5 specimens).
| 7 | 8 | 9 | 10 | mean | SD | |
|---|---|---|---|---|---|---|
| Telestes dabar | 20 | 20 | 9.5 | 0.51 | ||
| Telestes miloradi | 1 | 11 | 6 | 9.3 | 0.57 | |
| Telestes metohiensis, Gatačko Polje | 2 | 33 | 15 | 1 | 8.3 | 0.58 |
| Telestes metohiensis, Nevesinjsko Polje | 6 | 28 | 12 | 1 | 8.2 | 0.67 |
| Total lateral-line scales | Total lateral-series scales | ||||||||||
| 0–53 | 54–57 | 58–61 | 62–65 | 66–69 | 70–73 | mean | SD | range | mean | SD | |
| Telestes dabar | 5 | 17 | 8 | 7 | 3 | 54.2 | 9.58 | 62–69 | 66.6 | 2.41 | |
| Telestes miloradi | 6 | 5 | 5 | 2 | 60.3 | 3.91 | 58–69 | 62.8 | 2.93 | ||
| Telestes metohiensis, Gatačko Polje | 11 | 10 | 25 | 5 | 61.0 | 3.92 | 56–71 | 63.5 | 3.74 | ||
| Telestes metohiensis, Nevesinjsko Polje | 1 | 9 | 11 | 19 | 5 | 2 | 61.1 | 5.32 | 60–71 | 65.0 | 3.07 |
Total vertebrae (Table 4) 39 (49), 40 (101) or 41 (11), 40 in holotype; abdominal vertebrae 22 (101), 23 (58) or 24 (2), 22 in holotype; caudal vertebrae 16 (9), 17 (82) or 18 (70), 18 in holotype; predorsal vertebrae 13 (24), 14 (126) or 15 (11), 14 in holotype; intermediate vertebrae 3 (123) or 4 (38), 3 in holotype. Most frequent vertebral formulae 22+17 (41), 22+18 (60) and 23+17 (40), 22+18 in holotype.
| 38 | 39 | 40 | 41 | 42 | mean | SD | |
|---|---|---|---|---|---|---|---|
| Telestes dabar | 49 | 101 | 11 | 39.8 | 0.56 | ||
| Telestes miloradi | 1 | 10 | 11 | 40.5 | 0.60 | ||
| Telestes metohiensis, Gatačko Polje | 6 | 33 | 12 | 2 | 39.2 | 0.68 | |
| Telestes metohiensis, Nevesinjsko Polje | 10 | 112 | 35 | 39.2 | 0.52 | ||
| Telestes croaticus | 5 | 21 | 38.8 | 0.40 | |||
| Telestes fontinalis | 1 | 7 | 3 | 39.2 | 0.60 | ||
| Telestes karsticus | 2 | 2 | 3 | 41.1 | 0.89 | ||
| Telestes polylepis | 2 | 9 | 6 | 41.2 | 0.66 | ||
| Telestes turskyi | 2 | 9 | 6 | 1 | 40.3 | 0.77 | |
| Telestes ukliva | 8 | 42.0 | 0 |
| 21+17 | 22+16 | 22+17 | 22+18 | 22+19 | 23+16 | 23+17 | 23+18 | 23+19 | 24+16 | 24+17 | 24+18 | 24+19 | |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Telestes dabar | 41 | 60 | 8 | 40 | 10 | 1 | 1 | ||||||
| Telestes miloradi | 1 | 6 | 1 | 4 | 10 | ||||||||
| Telestes metohiensis, Gatačko Polje | 6 | 28 | 2 | 5 | 10 | 2 | |||||||
| Telestes metohiensis, Nevesinjsko Polje | 7 | 58 | 12 | 33 | 24 | ||||||||
| Telestes croaticus | 2 | 3 | 16 | 5 | |||||||||
| Telestes fontinalis | 1 | 1 | 7 | 3 | |||||||||
| Telestes karsticus | 1 | 1 | 2 | 1 | 2 | ||||||||
| Telestes polylepis | 1 | 1 | 8 | 4 | 1 | 2 | |||||||
| Telestes turskyi | 1 | 4 | 1 | 5 | 6 | 1 | |||||||
| Telestes ukliva | 3 | 5 |
| Abdominal vertebrae | Caudal vertebrae | Predorsal vertebrae | |||||||||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| 21 | 22 | 23 | 24 | Mean | SD | 15 | 16 | 17 | 18 | 19 | mean | SD | 12 | 13 | 14 | 15 | mean | SD | |
| Telestes dabar | 101 | 58 | 2 | 22.4 | 0.51 | 9 | 82 | 70 | 17.4 | 0.59 | 24 | 126 | 11 | 13.9 | 0.46 | ||||
| Telestes miloradi | 8 | 14 | 22.6 | 0.49 | 5 | 16 | 1 | 17.8 | 0.50 | 15 | 7 | 14.3 | 0.5 | ||||||
| Telestes metohiensis, Gatačko Polje | 36 | 17 | 22.3 | 0.47 | 7 | 42 | 4 | 16.9 | 0.46 | 3 | 38 | 12 | 14.2 | 0.50 | |||||
| Telestes metohiensis, Nevesinjsko Polje | 1 | 92 | 64 | 22.4 | 0.51 | 2 | 43 | 105 | 7 | 16.7 | 0.56 | 2 | 124 | 31 | 14.2 | 0.42 | |||
| Telestes croaticus | 2 | 19 | 5 | 22.1 | 0.52 | 8 | 18 | 16.7 | 0.47 | 21 | 5 | 14.2 | 0.40 | ||||||
| Telestes fontinalis | 2 | 9 | 22.8 | 0.40 | 7 | 4 | 16.4 | 0.50 | 2 | 9 | 14.8 | 0.40 | |||||||
| Telestes karsticus | 1 | 4 | 2 | 23.1 | 0.69 | 1 | 5 | 1 | 18.0 | 0.58 | 4 | 3 | 13.4 | 0.53 | |||||
| Telestes polylepis | 1 | 13 | 3 | 23.1 | 0.49 | 2 | 11 | 4 | 18.1 | 0.60 | 1 | 16 | 13.9 | 0.24 | |||||
| Telestes turskyi | 5 | 13 | 22.7 | 0.46 | 1 | 6 | 10 | 1 | 17.6 | 0.62 | 16 | 2 | 13.1 | 0.32 | |||||
| Telestes ukliva | 3 | 5 | 23.6 | 0.52 | 5 | 3 | 18.4 | 0.52 | 2 | 5 | 1 | 12.9 | 0.64 |
Colouration
In live specimens, dark back contrasting sharply with pale area below lateral midline, even in small specimens. Black midlateral stripe extending from head to caudal peduncle forming ventral border of darkly pigmented area on back. Another black lateral stripe occurring more ventrally, on otherwise pale ventral portion of trunk; this stripe extending from eye or opercle (or just behind opercle) to at least vertical through point halfway between origins of pelvic and anal fins, sometimes extending as poorly coalesced spots onto caudal peduncle. Dash-like black marking present along internal procurrent rays of caudal-fin dorsal lobe, and elongate black blotch present at bases of 3rd–7th branched rays of dorsal fin. Black pigment also occurring on rays of dorsal and caudal fins, but its intensity varying among individuals. Peritoneum black. This general pattern of pigmentation retained in formaldehyde-fixed and ethanol-preserved specimens. Live specimens collected from May through September, both males and females, exhibiting yellowish-orange pigment at bases of all fins, especially pectoral and anal fins, and yellowish pigment on iris and along anterior, dorsal and posterior margins of operculum. Colouration of specimens in cold season unknown.
Sexual dimorphism
Genital papilla absent in both males and females. Most morphometric characters not significantly different between males and females (Table 1a) with five exceptions. In males, distance between origins of pectoral and pelvic fins longer than in females (P<0.0001), dorsal fin deeper (P<0.02), anal fin deeper (P<0.0001), pectoral fin longer (P<0.0001), pectoral fin often reaching pelvic-fin origin in males, and pelvic fin longer (P<0.0001), pelvic fin often reaching anal-fin origin in males.
In samples collected in May, ripe males with small but prominent conical breeding tubercles. Tubercles regularly covering entire body, including dorsal and ventral surfaces of caudal peduncle, except for ventralmost surface of head. Single tubercle located on each scale. On all fins (except for caudal fin), tubercles present on both sides along all rays and on fin membrane, being particularly dense along marginal rays. Tubercles forming rows along outer margins of operculum and pectoral fin; tubercles in those rows larger than others on body. Degree of tubercle development varying between males with regard to both size of tubercles and their location. Tubercles always present on head, back, and pectoral fin. Males retaining tubercles, though reduced in size and density, until September.
Distribution
The new species is known from two rivers, Vrijeka and Opačica, in the Dabarsko Polje of Eastern Herzegovina in Bosnia and Herzegovina (Fig. 4).
Habitat and biology
From May through September Telestes dabar is found in shallow water of those river sections that are adjacent to and filled from underground springs. There is no current, and the water is clean (Fig. 5a). Females with eggs and just-spent females were caught on 24 May 2001 in Vrijeka River and mature males and just-spent females on 31 May 2000 in Opačica River. The size of the ripe eggs was 1.3–1.7 mm in diameter. In all examined samples females predominate. The smallest spent female was 45.0 mm SL, and the smallest ripe male 43.7 mm SL. No other fishes were caught in Opačica together with Telestes dabar while Delminichthys ghetaldii were collected in Vrijeka.
Etymology
The specific name, dabar, refers to the type locality, Dabarsko, or Dabar Polje; it is a noun in apposition.
Original Description
- Bogutskaya, N; Zupančič, P; Bogut, I; Naseka, A; 2012: Two new freshwater fish species of the genus Telestes (Actinopterygii, Cyprinidae) from karst poljes in Eastern Herzegovina and Dubrovnik littoral (Bosnia and Herzegovina and Croatia) ZooKeys, 180: 53-80. doi
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