Trichosiopsis fuscipalpa fuscipalpoides

Taxonavigation

Ordo: Diptera
Familia: Sciaridae
Genus: Trichosiopsis
Species: Trichosiopsis fuscipalpa

Name

Trichosiopsis fuscipalpa fuscipalpoides  unpublished

Type material

Holotype: ♂, 16.5.–2.6.2014, Malaise trap, leg. Rulik, ZFMK-TIS-2564672 in ZFMK

Type locality

Germany, Rhineland-Palatinate, Niederzissen, Bausenberg

Barcoded material

Stadium	Country	Province	Locality	Habitat	Method	Date	Collector	Collection Number	Collection	Latitude	Longitude	Elevation (m)
1 ♂	Germany	Rhineland-Palatinate	Niederzissen, Bausenberg	Wiese	Malaise Trap	16.5.–2.6.2014	Björn Rulik	ZFMK-TIS-2564672	ZFMK	50.465	7.226	292
1	Germany	Rhineland-Palatinate	Bad Neuenahr-Ahrweiler, Landskrone		Malaise Trap	25.07.2013	Björn Rulik	BIOUG16219-G06	ZFMK	50.552	7.17	194
1 ♂	Germany	Hesse	Lorch a. Rhein, oberhalb der Burg Nollig	Trockenhang	Malaise Trap	10.05.2013	Oliver Niehuis	ZFMK-TIS-2527933	ZFMK	50.05	7.798
1 ♂	Germany	Rhineland-Palatinate	Bad Neuenahr-Ahrweiler, Landskrone	Forest	Malaise Trap	02.06.2014	Björn Rulik	BIOUG17094-B05	ZFMK	50.552	7.17	194
1 ♀	Germany	Rhineland-Palatinate	Bad Neuenahr-Ahrweiler, Landskrone		Malaise Trap	02.06.2014	Björn Rulik	BIOUG17094-C01	ZFMK	50.552	7.17	194
1	Germany	Rhineland-Palatinate	Mayen-Koblenz	Forest	Malaise Trap	16.05.2013	Björn Rulik	BIOUG22302-F10	ZFMK	50.32	7.493	243
1	Germany	Rhineland-Palatinate	Mayen-Koblenz		Malaise Trap	29.05.2013	Björn Rulik	BIOUG22303-H02	ZFMK	50.32	7.493	243
1	Germany	Rhineland-Palatinate	Mayen-Koblenz	Forest	Malaise Trap	29.05.2013	Björn Rulik	BIOUG22304-C11	ZFMK	50.32	7.493	243
1 ♂	Germany	Rhineland-Palatinate	Niederzissen, Bausenberg	Wiese	Malaise Trap	16.5.–2.6.2014	Björn Rulik	ZFMK-TIS-2559687	ZFMK	50.465	7.226	292
1 ♀	Germany	Rhineland-Palatinate	Niederzissen, Bausenberg	Trockenrasen	Malaise Trap	24.04.2014	Björn Rulik	ZFMK-TIS-2564781	ZFMK	50.465	7.222	321
1 ♂	Norway	Buskerud	Hurum, Holtnesdalen		Malaise Trap	07.07.2010	L.O. Hansen	bf-sci-01334	NHMO	59.5409	10.4285	110

Description (male)

Head. Eye bridge 4 rows of facets. Antenna unicolour. LW-index of 4^th flagellomere 2.3–2.8; neck 0.3–0.5 × segment width; transition of basal part to neck pronounced. Colour of neck unicolour. Antennal setae shorter than segment width; of normal strength; dense; salient. Palpus darkened; of normal length; palpomeres 3. First palpomere elongate; with 3–5 setae; with only sparse sensilla. Second palpomere short and oval. Third palpomere as long as first. Thorax. Colour brown. Notum unicolour. Thoracic setae normal; brown. Posterior pronotum setose. Postpronotal setae 1–4; fine. Mesothoracic sclerites bare. Legs. Colour yellow. Hind coxa of same colour as femora. Setae on front coxa darkened. Front tibial organ as patch of setae; dark; front tibial organ not bordered. Tibial setae on hind legs weak and inconspicuous. Tibial spurs of equal length. Claws untoothed. Wing. Wing slightly darkened; of normal shape, or with weakly developed anal lobe. Wing membrane without macrotrichia. Wing venation weak, with faint stM. M-fork of normal shape. R₁ ending at or slightly before base of m-fork; posterior veins with macrotrichia; stM mostly with macrotrichia; CuA₁ and CuA₂ mostly with macrotrichia; bM bare; r-m with few setae, or mostly setose; bM:r-M 0.6–0.8; st-Cu:bM 0.4–0.8; R₁:R 1.3–1.65; c:w 0.71–0.81. Halter darkened; of normal length. Abdomen. Abdominal setae strong; dense; on tergites black; on sternites black. Hypopygium concolour with abdomen; LW-index 0.6–0.75. Base of gonocoxites with weak setae; gonocoxites broadly separated; inner margin of gonocoxites typically U-shaped; inner membrane of hypopygium scarcely setose; ventral margin of gonocoxite with short setae. Gonostylus elongate; LW-index 1.75–1.96; Inner margin concave; apex with one obtuse angle. Apical tooth present; without internal structure; of medium strength. Awl-like setae normal; present beneath apical tooth. Megasetae absent. Whiplash-hair absent. Tegmen 0.6–0.82 × longer than broad; rectangular with rounded edges; without special features; central process absent. Length of ejaculatory apodeme/hypopygium 28–36 %; base of ejaculatory apodeme present. Field with aedeagal teeth inconspicuous. Measurements. Body size 2.8–3.6 mm. Hind tibia 1.5–1.7 mm. Wing length 2.5–3 mm.

Diagnosis

This species is based on its unique COI sequence. Tr. fuscipalpa fuscipalpoides is smaller than Trichosiopsis fuscipalpa fuscipalpa. It has a rather tumid gonostylus with a distinct apico-dorsal angle.

DNA Barcoding

The COI sequence is assigned to BIN BOLD:ACC1319 (average distance 0.09%, max. 0.17%, n=11, K2P: 8.21%).

Discussion

Trichosiopsis fuscipalpa fuscipalpoides is morphologically very similar to the other subspecies in the complex, but has the highest K2P of all. This alone might justify its consideration as a distinct species, but as the morphological distinction is not always possible, for practical reasons only subspecific rank is assigned to this taxon.

Etymology

The name refers to the similarity with Tr. fuscipalpa fuscipalpa.

Distribution

Germany, Norway.

Images

Figure 1. Trichosiopsis fuscipalpa fuscipalpoides male, habitus, scale 1.0 mm  Figure 2. Trichosiopsis fuscipalpa fuscipalpoides male, hypopygium, scale 0.1 mm  Figure 3. Trichosiopsis fuscipalpa fuscipalpoides male, gonostylus (ventral view), scale 0.1 mm  Figure 4. Trichosiopsis fuscipalpa fuscipalpoides male, aedeagal complex, scale 0.1 mm  Figure 5. Trichosiopsis fuscipalpa fuscipalpoides male, 4th antennal flagellar segment, scale 0.1 mm  Figure 6. Trichosiopsis fuscipalpa fuscipalpoides male, wing, scale 1.0 mm