Eotrichocera longensis
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Ordo: Diptera
Familia: Trichoceridae
Genus: Eotrichocera
Name
Eotrichocera longensis Dong, Shih & Ren, 2014 sp. n. – Wikispecies link – ZooBank link – Pensoft Profile
Etymology
“longensis” is a Latin word, referring to the long leg of this specimen.
Diagnosis
Sc rather short about 0.65 times as long as the wing and ending at anterior margin proximad of R2; Rs forking proximad of 2/3 (at about 0.55) times of wing length; the d-cell narrow and long (about 2.5 times as long as wide); A2 short and bending sharply toward anterior margin (angle about 128°).
Holotype
An almost complete female specimen with well-preserved body, wings and head. Specimen number CNU-DIP-NN2013133. Wing length 9.0 mm, width 3.8 mm (Figs 1A, 2, 3A).
Paratype
A female specimen with body and wings, specimen number CNU-DIP-NN2013131. Wing length 7.7 mm, width 3 mm (Figs 1B, 3B).
Locality and horizon
Jiulongshan Formation, Late Middle Jurassic, Daohugou Village, Ningcheng County, Inner Mongolia Autonomous Region, China.
Description
Based on Holotype, different characters of the paratype CNU-DIP-NN2013131 in brackets. Medium-sized winter crane flies, body length (including head) 13 mm with well preserved wings, body and head, [paratype body length (excluding head) 10.5 mm].
Head: Antenna very long, about 3.5 times as long as the head length, palpi about two times as long as the head length, compound eyes preserved.
Thorax: Much higher, in lateral view, than that of the abdomen, subcircular in shape, with robust and well-developed mesonotum. The halters spoon-type and the length of halters as long as thorax.
Wings: Wing is shorter than abdomen, not covering the end of the abdomen. Length 9.0 mm [Paratype with wing length of 7.7 mm], narrow and long (about 2.5 times as long as wide); venation clear, Sc rather short about 0.68 times as long as the wing [Paratype Sc rather short, about 0.65 times as long as the wing] and ending at anterior margin proximad of R2; crossvein sc-r locating at 2/3 of Rs; Rs arising about one-fifth from the base of the wing; R2+3 about 0.8 times as long as R2+3+4; R2 about one-tenth of length of R3; R3 almost three times as long as the R2+3; dM1+2 0.6 times as long as mM1+2, while M1 2.5 times of the dM1+2; a well developed m-m crossvein about three-fourth length of bM3, closing the d-cell and nearly 0.3 length of d-cell; bM1+2 nearly 1.0 times as long as the length of the r-m and the latter at one-fifth of the d-cell; d-cell narrow and long (about 2.5 times as long as wide) and almost one-fifth length of wing; both crossveins m-m and m-cu intersecting with M4 at the same point; Cu long, curved (angle about 135°) and reaching the wing posterior margin at 0.6 from the base of the wing; the stem of A divided into A1 and A2; A1 long, slightly curving and reaching the wing posterior margin; A2 short, 0.15 (right wing) [Paratype 0.14] times as long as wing and almost 0.3 times as long as length of A1, bending sharply (angle about 128°) and reaching the wing posterior margin.
Legs: Legs slender and long; the hind leg nearly 1.2 times as long as the abdomen and 1.3 times as long as the wing. Tarsus with five segments; the first segment of tarsus (t1) is 1.2 times as long as t2 in mid leg.
Abdomen: Abdomen relatively long and thin, with ten segments. Female genital discernible.
Remarks
Eotrichocera (Archaeotrichocera) longensis sp. n. is assigned to Thrichocerinae based on the following characters: d-cell medium, m-cu present; A2 short, antennae long, flagellomeres thin, much longer than two times of the head length. It belongs to Eotrichocera (Archaeotrichocera) because of wing length from 7.7 to 9.0 mm and d-cell almost one-fifth of wing length. In addition, it differs from all other known Thrichocerinae by its A2 rather short and bending sharply toward anterior margin (angle about 128°), R2 relatively long, Sc forking proximad of 2/3 (at about 0.55) times of wing length, and d-cell narrow and long. To compare the key characters among the new species and other species of Eotrichocera (Archaeotrichocera), we set up the Table 2.
| Key characters | Eotrichocera (Archaeotrichocera) ephemera Zhang, 2006 | Eotrichocera (Archaeotrichocera) conica Krzemińska, Krzemiński & Ren, 2009 | Eotrichocera (Archaeotrichocera) rara Krzemińska, Krzemiński & Ren, 2009 | Eotrichocera (Archaeotrichocera) spatiosa Liu, Shih & Ren, 2012 | Eotrichocera (Archaeotrichocera) longensis sp. n. | Eotrichocera (Archaeotrichocera) amabilis sp. n. |
|---|---|---|---|---|---|---|
| Wing length, in mm and (L/W) | 7.1 (L/W=2.8) | 10.0 (L/W=3) | 5.5 | 12.0 (L/W=2.7) | 9.0 (L/W=2.3) | 5.2 (L/W=2.2) |
| Sc length and ending at anterior margin | 0.77 times of wing length, ending proximad of R2 | 0.71 times of wing length, ending proximad of R2 | 0.77 times of wing length, ending distad of R2 | 0.84 times of wing length, and ending distad of R2 | about 0.65 times of wing length, ending proximad of R2 | about 0.71 times of wing length, ending proximad of R2 |
| sc-r position | at 2/3 of length of Rs | at 1/3 of length of Rs | at 1/2 of length of Rs | at 2/3 of length of Rs | at 2/3 of length of Rs | at 1/3 of length of Rs |
| Position of Rs forking | 0.77 times of wing length | 0.64 times of wing length | 0.57 times of wing length | 0.58 times of wing length | 0.53 times of wing length | 0.55 times of wing length |
| d-cell W/L (length) | W/L=0.53 (1/6 of wing length) | W/L=0.43 (0.2 times of wing length) | W/L=0.39 (0.21 times of wing length) | W/L=0.47 (0.19 times of wing length) | W/L= 0.4 (almost 1/5 of wing length) | W/L=0.58 (almost 0.17 of wing length) |
| A2 length | long (about 1/4 of wing length), curved evenly to posterior margin | medium (0.22 times of wing length), not reaching posterior margin | rather short (1/5 of wing length) and not reaching posterior margin | short (about 0.21 times of wing length) and curving to posterior margin | short (0.14) times of wing length) and curved to posterior margin | short (0.13) times of wing length) and curved to posterior margin |
| r-m length | ....... | 1/5 of length of d-cell | ....... | about 1/3 of length of d-cell | 1/5 of length of the d-cell | 0.24 or 0.15 of length of the d-cell |
Original Description
- Dong, F; Shih, C; Ren, D; 2014: Two new species of Trichoceridae from the Middle Jurassic Jiulongshan Formation of Inner Mongolia, China ZooKeys, 411: 145-160. doi
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