Difference between revisions of "Madagascarchaea jeanneli"
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==Diagnosis== | ==Diagnosis== | ||
| − | Distinguished from all ''{{Taxon name|Madagascarchaea}}'', except other ‘pointy head’ species ''{{Taxon name|M. ambre}}'', ''{{Taxon name|M. lotzi}}'', and ''{{Taxon name|M. | + | Distinguished from all ''{{Taxon name|Madagascarchaea}}'', except other ‘pointy head’ species ''{{Taxon name|M. ambre}}'', ''{{Taxon name|M. lotzi}}'', and ''{{Taxon name|M. moramora}}'' sp. n., by having a conductor that is a concave triangular shape (Fig. 26D–E, J–H, J–K), and by having an abdomen that is invaginated across the back (Fig. 26A), rather than rounded (as in Fig. 23A). Typically ''{{Taxon name|M. jeanneli}}'' can be distinguished from ''{{Taxon name|M. ambre}}'' and ''{{Taxon name|M. moramora}}'' sp. n., by having an abdomen that is invaginated in the posterior (Fig. 26A), rather than straight (Fig. 28A). ''{{Taxon name|M. jeanneli}}'' is further distinguished from ''{{Taxon name|M. moramora}}'' by having the anterior portion of the embolus taper off to a point (Fig. 26G–I, “a”) rather than being broad and blunt (Fig. 28D–F, “a”), and from ''{{Taxon name|M. ambre}}'' by having the anterior piece of the embolus broad and curved (Fig. 26G–I, “a”) and not straight, narrow, and jutting out past the conductor in the retrolateral direction (see fig. 21 in Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >). ''{{Taxon name|M. jeanneli}}'' is distinguished from ''{{Taxon name|M. lotzi}}'' sp. n. by having an embolus with a very deep bifurcation (Fig. 26I, L), and with the posterior portion of the embolus having an additional bifurcation at the tip (Fig. 26F, I, “p”). |
==Description== | ==Description== | ||
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==Nomenclature remarks== | ==Nomenclature remarks== | ||
| − | Previous work (Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >) lumped into ''{{Taxon name|M. jeanneli}}'' what we are here calling several different species. The ''{{Taxon name|M. jeanneli}}'' syntypes include two females, and females of the “{{Taxon name|jeanneli}} complex “(''{{Taxon name|M. ambre}}'', ''{{Taxon name|M. jeanneli}}'', ''{{Taxon name|M. lotzi}}'' sp. n., and ''{{Taxon name|M. | + | Previous work (Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref >) lumped into ''{{Taxon name|M. jeanneli}}'' what we are here calling several different species. The ''{{Taxon name|M. jeanneli}}'' syntypes include two females, and females of the “{{Taxon name|jeanneli}} complex “(''{{Taxon name|M. ambre}}'', ''{{Taxon name|M. jeanneli}}'', ''{{Taxon name|M. lotzi}}'' sp. n., and ''{{Taxon name|M. moramora}}'' sp. n.) cannot be confidently distinguished. However, the type locality of “La Mandraka” is an area about 40 kilometers east of Antananarivo, and so we propose that the female syntypes are conspecific to the males found in this area. The male and female specimens that were described and illustrated as ''{{Taxon name|M. jeanneli}}'' in Wood 2008<ref name="B25"> {{aut|Wood H}} (2008) A revision of the assassin spiders of the ''{{Taxon name|Eriauchenius gracilicollis}}'' group, a clade of spiders endemic to Madagascar ({{Taxon name|Araneae}} : {{Taxon name|Archaeidae}}). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x</ref > are here being called ''{{Taxon name|M. lotzi}}'' n. sp. |
==Taxon Treatment== | ==Taxon Treatment== | ||
Revision as of 11:15, 9 January 2018
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Ordo: Araneae
Familia: Archaeidae
Genus: Madagascarchaea
Name
Madagascarchaea jeanneli (Millot, 1948) comb. n. – Wikispecies link – Pensoft Profile
- Archaea jeanneli Millot, 1948: 12, figs 1A,2B,3F.
- Eriauchenius jeanneli (Millot, 1948): Wood 2008[1]: 286, figs 2E,12B,18A,20A–D (new combination)
Type material
2 females, 1 juv: as Archaea jeanneli Millot, 1948: Madagascar, La Mandraka (deposited in MNHN, examined; MNHN 13/1970)
Other material examined
MADAGASCAR: 2M,1F, Toamasina, Res. Analamazaotra, Parc National Andasibe, 23 road km E Moramanga, 18°56'38.2"S, 48°25'03.2"E, 960 m, 16–18 Jan 2003, rainforest, general collecting night, C. Griswold, D. Silva, and D. Andriamalala (CASENT9005233, CASENT9018315); 7M,4F,1juv, Antananarivo, 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe, 18°28'24"S, 47°57'36"E, 1300 m, 5–13 Dec 2000, montane rainforest, general collecting, Fisher, Griswold et al. (CASENT9004085); 3M,4F,1juv, same data as previous except beating and sweeping (CASENT9004009); 6F, Toamasina, Parc National Perinet, nr Andasibe, 18°56’S, 48°24’E, 1000 m, 4–5 Nov 1993, J. Coddington, S. Larcher, C. Griswold, R. Andriamasimanana, and N. Scharff (CASENT9012329, CASENT9046592, CASENT9046601); 1M, Toamasina, Parc National Perinet, 18°55’S, 48°25’E, 1-3 Aug 1992, V. & B. Roth (CASENT9012003); 2F, Toamasina, Station Forestier Analamazaotra, administered by Mitsinjo, 0.75 km N Andasibe, 18°55.783’S, 48°24.696’ E, 964m, 31 Jan – 3 Feb 2009, primary montane rainforest, hand collected at night in vegetation, C. Griswold, A. Saucedo and H. Wood (USNMENT01377201, USNMENT01377202); 1F, Antananarivo, Réserve Spéciale d’Ambohitantely, Forêt d’Ambohitantely, 20.9 km 72° NE Ankazobe, 18°13'31"S, 47°17'13"E, 1410 m, 17–22 Apr 2001, montane rainforest, EB17 beating low vegetation, Fisher, Griswold et al. (CASENT9012336); 2M,2F,1juv, Antananarivo, 7 km SE Andasibe Parc National (=Perinet) 18°58’S, 48°27’E, 5 Sep 2001, montane forest, beating foliage, D. Ubick (CASENT9001265).
Diagnosis
Distinguished from all Madagascarchaea, except other ‘pointy head’ species M. ambre, M. lotzi, and M. moramora sp. n., by having a conductor that is a concave triangular shape (Fig. 26D–E, J–H, J–K), and by having an abdomen that is invaginated across the back (Fig. 26A), rather than rounded (as in Fig. 23A). Typically M. jeanneli can be distinguished from M. ambre and M. moramora sp. n., by having an abdomen that is invaginated in the posterior (Fig. 26A), rather than straight (Fig. 28A). M. jeanneli is further distinguished from M. moramora by having the anterior portion of the embolus taper off to a point (Fig. 26G–I, “a”) rather than being broad and blunt (Fig. 28D–F, “a”), and from M. ambre by having the anterior piece of the embolus broad and curved (Fig. 26G–I, “a”) and not straight, narrow, and jutting out past the conductor in the retrolateral direction (see fig. 21 in Wood 2008[1]). M. jeanneli is distinguished from M. lotzi sp. n. by having an embolus with a very deep bifurcation (Fig. 26I, L), and with the posterior portion of the embolus having an additional bifurcation at the tip (Fig. 26F, I, “p”).
Description
Male (based on CASENT9012003, from Parc National Perinet, Madagascar). Total length 1.55, carapace 0.64 long, 0.47 wide. Abdomen 0.79 long, 0.91 high. Carapace tilt angle 50.2°, tilt height (CtH) 1.19, constriction 0.35, head length 0.68, neck length 0.59. CtH divided by carapace length 1.85. Cephalon with AME on large bulge, and with 6 short post-occular spines at the crown, not on protrusions, and 1 short spine between and posterior to the LE and median eyes (see fig. 18B in Wood 2008[1]). Chelicerae 1.18 long, and with a small spine 0.30 from base of chelicerae and projecting downward (Fig. 26A). Femur I 1.96 long. Sternum 0.44 long, 0.25 wide. Carapace, chelicerae, and sternum reddish brown with white setae. Legs light brown with darker annulations throughout. Abdomen dark brown with lighter circular patches throughout, with white and brown setae (Fig. 26A). Posterior edge of abdomen invaginated (Fig. 27A, arrow). Conductor concave and triangular; MA dark, thick, and curves anteriorad; S1 a thin ridge; embolus dark, with deep bifurcation, anterior portion curved and tapering off and posterior portion with additional bifurcation at tip with each side tapering off (Fig. 26D–L).
Female (based on CASENT9012336, from Réserve Spéciale d’Ambohitantely, Madagascar). Total length 2.00, carapace 0.78 long, 0.59 wide. Abdomen 1.18 long, 1.49 high. Carapace tilt angle 60.0°, tilt height (CtH) 1.53, constriction 0.41, head length 0.94, neck length 0.73. CtH divided by carapace length 1.97. Cephalon as in male. Chelicerae 1.48 long, and with a short spine 0.36 from base of chelicerae and projecting downward. Femur I 2.31 long. Sternum 0.51 long, 0.28 wide. Colors as in male. Posterior edge of abdomen invaginated. Genitalic bursa divided down the middle by sclerotized piece on the anterior-ventral side, with a few clusters of poreplates on either side; FSGP with two strong points arising from either side of anterior edge, having ‘wings’, and lacking posterior elongation (Fig. 26B–C).
Variation
Total length 1.54–1.83 (males; n=5), 1.82–2.00 (females; n=5); Carapace length 0.64–0.76 (males; n=5), 0.74–0.78 (females; n=5); Femur I 2.60–3.06 times the length of carapace in males (n=5), 2.88–3.12 in females (n=5); CtH divided by carapace length 1.72–1.85 in males (n=5), 1.75-2.01 in females (n=5); Average femur I length 2.01 in males (n=5), 2.23 in females (n= 5).
Natural history
Specimens were collected in montane rainforest from 960–1300 m in elevation by beating low foliage, low vegetation, by beating and sweeping, by general collecting at night, and by hand collecting at night in vegetation.
Distribution
Known only from central eastern Madagascar (Fig. 34).
Nomenclature remarks
Previous work (Wood 2008[1]) lumped into M. jeanneli what we are here calling several different species. The M. jeanneli syntypes include two females, and females of the “jeanneli complex “(M. ambre, M. jeanneli, M. lotzi sp. n., and M. moramora sp. n.) cannot be confidently distinguished. However, the type locality of “La Mandraka” is an area about 40 kilometers east of Antananarivo, and so we propose that the female syntypes are conspecific to the males found in this area. The male and female specimens that were described and illustrated as M. jeanneli in Wood 2008[1] are here being called M. lotzi n. sp.
Taxon Treatment
- Wood, H; Scharff, N; 2017: A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae) ZooKeys, (727): 1-96. doi
Images
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Other References
- ↑ 1.0 1.1 1.2 1.3 1.4 Wood H (2008) A revision of the assassin spiders of the Eriauchenius gracilicollis group, a clade of spiders endemic to Madagascar (Araneae : Archaeidae). Zoological Journal of the Linnean Society 152: 255–296. https://doi.org/10.1111/j.1096-3642.2007.00359.x
