Difference between revisions of "Mystrium oberthueri"

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{{Publication to wiki notice
 
{{Publication to wiki notice
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  | author = Forel, A.
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  | author = Yoshimura, Masashi AND Fisher, Brian L.
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  | author_abbreviated = Forel, A.
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  | author_abbreviated = Yoshimura M AND Fisher B
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  | year = 1897
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  | year = 2014
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  | title = Ameisen aus Nossi-Be, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar. Gesammelt von Herrn Dr. A. Voeltzkow aus Berlin.
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  | title = A revision of the ant genus ''Mystrium'' in the Malagasy region with description of six new species and remarks on ''Amblyopone'' and ''Stigmatomma'' (Hymenoptera, Formicidae, Amblyoponinae)
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  | journal = Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft
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  | journal = ZooKeys
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  | volume = 21
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  | volume = 394
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  | pages = 192 -- 193
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  | pages = 1--99
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  | doi = TODO
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  | doi = 10.3897/zookeys.394.6446
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  | citationurl = http://antbase.org/ants/publications/3965/3965.pdf
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  | citationurl = http://www.pensoft.net/journals/zookeys/article/6446/citation/  
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  | url = http://antbase.org/ants/publications/3965/3965.pdf
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  | url = http://www.pensoft.net/journals/zookeys/article/6446/abstract
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  | publisher =  
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  | publisher = Pensoft Publishers
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  | publisherurl =  
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  | publisherurl = http://www.pensoft.net/
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}}
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}}
  
 
{{Treatment start
 
{{Treatment start
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  | Ordo =  
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  | Ordo = Hymenoptera
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  | Familia =  
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  | Familia = Formicidae
 
  | Genus = Mystrium
 
  | Genus = Mystrium
 
  | Specific name = oberthueri
 
  | Specific name = oberthueri
 
  | Infraspecific name =  
 
  | Infraspecific name =  
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  | Taxon rank = species
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  | Taxon rank =
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  | Taxon authority = Forel, A., 1897
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  | Taxon authority = Forel, 1897
 
  | Taxon status =  
 
  | Taxon status =  
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  | Nomenclature citation = {{Nomenclature citation| ''{{Taxon name|Mystrium oberthueri}}'' Forel, A., 1897, Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft 21: 192-193.}}  
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  | Nomenclature citation = {{Nomenclature citation |''{{Taxon name|Mystrium oberthueri}}'' Forel, 1897. MADAGASCAR. Syntypes: one worker (as major) and two ergatoid queens (as minor) <nowiki>[</nowiki>lectotype is designated below<nowiki>]</nowiki>.}} {{Nomenclature citation |''{{Taxon name|Mystrium oberthueri}}'': Menozzi 1929<ref name="B27">{{aut|Menozzi C}} (1929) Revisione delle formiche del genere ''Mystrium'' Roger. Zoologischer Anzeiger  82: 518-536.</ref > (in part?: uncertain male identification)}} {{Nomenclature citation |''{{Taxon name|Mystrium oberthueri}}'' (Not) (male): Emery 1899<ref name="B9">{{aut|Emery C}} (1899) Formiche di Madagascar raccolte dal Sig. A. Mocquerys nei pressi della Baia di Antongil (1897–1898). Bullettino della Societa Entomologica Italiana  31: 263-290.</ref >.}} {{Nomenclature citation |''{{Taxon name|Mystrium rogeri}}'' (male): Forel 1899<ref name="B18">{{aut|Forel A}} (1899) Trois notices myrmécologiques. Annales de la Société Entomologique de Belgique  43: 303-310.</ref >.}}
 
  | Wikispecies page name = Mystrium_oberthueri
 
  | Wikispecies page name = Mystrium_oberthueri
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| ZooBank ID =
 
 
  | Pensoft Profile = Mystrium_oberthueri
 
  | Pensoft Profile = Mystrium_oberthueri
 
}}
 
}}
  
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{{Section|id=description|heading = Description|content=&#x263F; minor L. 6,8 bis 7 mm ohne, 8,3 bis 8,5 mm mit Oberkiefer. Oberkiefer kuerzer als bei ''[[Mystrium_voeltzkowi|voeltzkowi]]'', ungefaehr wie bei ''[[Mystrium_mysticum|mysticum]]'', aber fast gerade, gleichmaessig schwach, nach aussen konvex und nach innen konkav, an der Spitze kaum breiter, stumpf abgestutzt, mit zwei Reihen von je ca. 10 kleinen Zaehnen, die etwa so stark sind wie bei ''[[Mystrium_mysticum|mysticum]]''. Der Oberkiefer an der Basis mindestens so dick (sogar etwas dicker) als an der Spitze, sehr fein genetzt und matt.
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==Lectotype==
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'''<nowiki>[</nowiki>here designated<nowiki>]</nowiki>.''' Worker: [http://data.antweb.org/specimen/CASENT0101993 CASENT0101993], MADAGASCAR, Insel Ste. Marie (Island of Sainte Marie), Perrot <nowiki>[</nowiki>MHNG: examined<nowiki>]</nowiki>.
  
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Kopf wie bei ''[[Mystrium_voeltzkowi|voeltzkowi]]'', viel breiter als lang, sogar noch breiter im Verhaeltnis zur Laenge, hinten weniger verengt als bei genannter Art, vorn seitlich mit zwei sehr dicken und viel laengeren Zaehnen als bei ''[[Mystrium_voeltzkowi|voeltzkowi]]'' &#x263F; minor. Augen ziemlich konvex, etwas hinter der Mitte. Clypeusrand schaerfer gezaehnt als bei ''[[Mystrium_mysticum|mysticum]]'', aber kuerzer als bei ''[[Mystrium_voeltzkowi|voeltzkowi]]''. Der Fuehlerschaft erreicht gerade den Hinterhauptrand. Geisselglieder etwas laenger als bei ''[[Mystrium_mysticum|mysticum]]'', cylindrischer und kuerzer als bei ''[[Mystrium_voeltzkowi|voeltzkowi]]'', mit Ausnahme des langen zweiten Gliedes. Thorax, Stielchen und Hinterleib so geformt wie bei ''[[Mystrium_mysticum|mysticum]]'', aber etwas schmaeler und laenglicher, Thorax staerker eingeschnuert, Stielchen etwas laenger im Verhaeltnis zur Breite (etwas mehr als doppelt so breit als lang).
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==Worker==
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'''Description.''' Measurements: lectotype. HL 2.40, HW 2.61, SL 1.80, ML 2.92, HD 1.67, WL 2.79, PnW 1.39, PpW 1.21, PtW 1.18, PtL 0.81, CI 108.7, SI 69.0, MI 111.8, PpI 87.4, PtI 144.9.<br />
 +
HL 2.00–2.58, HW 2.12–2.76, SL 1.55–1.96, ML 2.39–3.17, HD 1.33–1.69, WL 2.43–2.96, PnW 1.17–1.38, PpW 1.02–1.29, PtW 1.00–1.32, PtL 0.67–0.81, CI 104.4–108.5, SI 68.8–74.9, MI 106.9–122.9, PpI 87.2–93.8, PtI 139.5–163.2 (10 specimens measured).<br />
 +
Posterolateral corner of head moderately expanding posteriorly. Posterior face of vertex forming almost right angle with dorsal face on median line of head, so that declivity of vertex on lateral part as steep as on median part. Whole region of vertex finely striated. Eye moderately small. Anterior margin of clypeus straight or weakly convex with long conical setae. Genal tooth of head relatively long, as long as lateral lobe of clypeus. Masticatory surface of mandible almost invisible in full-face view, and width of dorsal surface of mandible almost identical from mandibular shaft to distal portion. Second maxillary palpomere longer than third. First flagellomere (third antennal segment) about 1.2–1.5× length of pedicel (second antennal segment). Strong, deep longitudinal striae regularly impressed on whole central part of pronotal dorsum. Strong, deep longitudinal striae impressed on lateral surface of pronotum. Mesonotum differentiated from propodeum in dorsal view, length slightly shorter than that of propodeum. Metanotal groove shallowly and gently impressed in lateral view. Metapleural gland bulla moderately developed, and propodeal declivity in lateral view almost straight. Petiole widened on posterior 1/3 and gently narrowing anteriorly in dorsal view, anterior margin straight to gently rounded and often edged by thick striae.<br />
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Body color black.
  
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Matt. Kopf sehr grob genetzt, mit Tendenz zur Laengsrunzelung, besonders auf der Stirne, Clypeus mit feiner Skulptur, sowie die abschuessige Flaeche des Metanotum, die Vorderflaeche des Stielchens und die drei letzten Hinterleibssegmente. Thorax, Stielchen und erstes Hinterleibssegment sehr grob und regelmaessig laengsgerunzelt, (Metanotum, Stielchen und erstes Hinterleibssegment fast gestreift). Zweites Hinterleibssegment fein laengsgerunzelt. Ueberall ist die feine Grundskulptur unregelmaessig, sehr fein genetzt-gerunzelt (am Hinterleib punktiert-genetzt).
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==Ergatoid queen==
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'''Description.''' Measurements: HL 1.40–1.93, HW 1.46–1.96, SL 1.07–1.47, ML 1.49–2.00, HD 1.00–1.35, WL 1.92–2.56, PnW 0.83–1.11, PpW 0.85–1.13, PtW 0.81–1.07, PtL 0.53–0.70, CI 101.6–108.6, SI 71.6–76.7, MI 96.3–108.8, PpI 96.0–104.4, PtI 149.1–177.1 (10 specimens measured).<br />
 +
Wings vestigial and reduced to small but distinct appendages. Wing sclerites undeveloped. Posterolateral corner of head moderately expanding posteriorly, expansion not differentiated from that in workers. Posterior face of vertex forming almost a right angle with dorsal face on median line of head, so that declivity of vertex on lateral part as steep as on median part. Ventral half of vertex sculptured. Eye small but distinct. Ocelli absent. Anterior margin of clypeus almost straight with long to moderate conical setae. Anterolateral portion of head with short spine. Masticatory margin of mandible almost invisible in full-face view, and dorsal surface on distal portion as wide as that on mandibular shaft. Spatulate seta present on basal side of each basal denticle on masticatory margin of mandible. First flagellomere (third antennal segment) long, about 1.2-1.5× length of pedicel (second antennal segment). Setae on pronotum distinctly spatulate, widened distally with sharp or blunt apex. Metapleural gland bulla moderately developed, not expanding dorsally to propodeal spiracle, so that propodeal declivity in lateral view weakly convex and rounded posteriorly on its ventral 1/3. Petiole relatively long in dorsal view, about 0.8× length of abdominal segment III.<br />
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Body color blackish to reddish brown.
  
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Behaarung zerstreut, aeusserst kurz, dick, ueberall ziemlich anliegend, aber nirgends deutlich kolbig oder loeffeifoermig, wodurch diese Art von allen anderen abweicht. An den Beinen ist die Behaarung reichlicher und dicker.
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==Male==
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'''Description.''' Measurements: HL 1.44, HW 2.09, SL 0.62, EL 0.95, WL 3.55, MnW 1.94, CI 145.1, SI 29.6, EI 66.2, MnI 92.9 (1 specimen measured).<br />
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Eye moderately large, occupying 0.6× of head length. Ocelli relatively distant from dorsal margin of head in full-face view. Dorsal margin of head in full-face view straight. Both anterior and lateral ocelli small. Lateral ocellus small and distant from eye: distance between these more than 1.5× maximum diameter of lateral ocellus. Posterior face of vertex not clearly differentiated from dorsal face, so that vertex almost continuously rounded. Palpal formula 4,3. First maxillary palpomere flattened and distinctly wider than second segment. Second maxillary palpomere longer than third. Notauli clearly impressed on mesoscutum. Petiole in dorsal view thin, 0.8× as long as abdominal tergite III. Petiolar dorsum covered with shallow, irregular punctures. Abdominal tergum VIII without deep punctures, almost smooth.<br />
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Distal portion of abdominal sternum IX smooth and not punctured. Basal ring moderately long, expanding basoventrally. Telomere distinctly extending distally farther than digitus. Basoventral expansion of aedeagus moderately developed basoventrally, longer than dorsal extension. Ventral margin of aedeagus gently convex in lateral view. Aedeagus moderately narrowing distally and distal portion rounded.<br />
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On forewing, cu-a located far basal from junction of Media (M) and Cubitus (Cu).<br />
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Body color reddish brown to black.
  
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Schwarz. Fuehlerkeule, erstes Geisselglied, Spitze der Mandibeln, Stirnleisten, Tarsen, Gelenke und Segmentraender roetlichgelb oder rotbraun, Fuehler, Beine und Basis der Mandibeln im uebrigen braun.
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==Distribution==
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MADAGASCAR: as in Figure 56D.
  
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&#x263F; major: L. ohne Mandibeln 9,7 mm, mit den Mandibeln 12,7 mm. Mandibeln wie bei ''[[Mystrium_voeltzkowi|M. voeltzkowi]]'', &#x263F; major, viel laenger als beim &#x263F; minor und mit staerker nach innen konkaver Endhaelfte, mit viel staerkeren Zaehnen (die Zaehne aber stumpfer und kuerzer als beim ''[[Mystrium_voeltzkowi|voeltzkowi]]'' &#x263F; major). Die Vorderecken mit je einem langen, spitzen und gekruemmten Dorn, der dreimal so lang ist als die Breite seiner Basis und doppelt so lang wie bei ''[[Mystrium_voeltzkowi|voeltzkowi]]'' &#x263F; major. Das 3. Abdominalsegment ist noch gestreift; Kopf querrechteckig, sonst wie der &#x263F; minor.
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==Additional material examined==
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In addition to the type material, specimens from the following localities were examined in this study: MADAGASCAR. Antsiranana. Forêt d’ Antsahabe, 11.4 km 275° W Daraina ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-13.21167%C2%B0%2C+49.55667%C2%B0&coordinates%5B0%5D=-13.21167%C2%B0%2C+49.55667%C2%B0 -13.21167°, 49.55667°]), tropical dry forest, 550 m alt.; Makirovana forest ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-14.17066%C2%B0%2C+49.95409%C2%B0&coordinates%5B0%5D=-14.17066%C2%B0%2C+49.95409%C2%B0 -14.17066°, 49.95409°]), rainforest, 415 m alt.; Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-14.435%C2%B0%2C+49.76%C2%B0&coordinates%5B0%5D=-14.435%C2%B0%2C+49.76%C2%B0 -14.435°, 49.76°]), rainforest, 775 m alt.; 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-14.43667%C2%B0%2C+49.775%C2%B0&coordinates%5B0%5D=-14.43667%C2%B0%2C+49.775%C2%B0 -14.43667°, 49.775°]), rainforest, 450 m alt.; Marojejy R.N.I. #12 ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-14.43583%C2%B0%2C+49.76056%C2%B0&coordinates%5B0%5D=-14.43583%C2%B0%2C+49.76056%C2%B0 -14.43583°, 49.76056°]), rainforest, 610 m alt.; Forêt Ambanitaza, 26.1 km 347° Antalaha ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-14.67933%C2%B0%2C+50.18367%C2%B0&coordinates%5B0%5D=-14.67933%C2%B0%2C+50.18367%C2%B0 -14.67933°, 50.18367°]), rainforest, 240 m alt.; 1 km W Andampibe, Cap Masoala ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-15.69361%C2%B0%2C+50.18139%C2%B0&coordinates%5B0%5D=-15.69361%C2%B0%2C+50.18139%C2%B0 -15.69361°, 50.18139°]), lowland rainforest, 125 m alt.; Fotodriana, Cap Masoala ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-15.69694%C2%B0%2C+50.27028%C2%B0&coordinates%5B0%5D=-15.69694%C2%B0%2C+50.27028%C2%B0 -15.69694°, 50.27028°]), lowland rainforest, 25 m alt.; Toamasina. Montagne d’Anjanaharibe, 18.0 km 21° NNE Ambinanitelo ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-15.18833%C2%B0%2C+49.615%C2%B0&coordinates%5B0%5D=-15.18833%C2%B0%2C+49.615%C2%B0 -15.18833°, 49.615°]), rainforest, 470 m alt.; Montagne d’Akirindro 7.6 km 341° NNW Ambinanitelo ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-15.28833%C2%B0%2C+49.54833%C2%B0&coordinates%5B0%5D=-15.28833%C2%B0%2C+49.54833%C2%B0 -15.28833°, 49.54833°]), rainforest, 600 m alt.; 19 km ESE Maroantsetra ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-15.48333%C2%B0%2C+49.9%C2%B0&coordinates%5B0%5D=-15.48333%C2%B0%2C+49.9%C2%B0 -15.48333°, 49.9°]), rainforest, 350 m alt.; 5.3 km SSE Ambanizana, Andranobe ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-15.66667%C2%B0%2C+49.96667%C2%B0&coordinates%5B0%5D=-15.66667%C2%B0%2C+49.96667%C2%B0 -15.66667°, 49.96667°]), rainforest, 425 m alt.; 6.3 km S Ambanizana, Andranobe ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-15.6813%C2%B0%2C+49.958%C2%B0&coordinates%5B0%5D=-15.6813%C2%B0%2C+49.958%C2%B0 -15.6813°, 49.958°]), rainforest, 5 m alt.; 6.3 km S Ambanizana, Andranobe ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-15.68131%C2%B0%2C+49.958%C2%B0&coordinates%5B0%5D=-15.68131%C2%B0%2C+49.958%C2%B0 -15.68131°, 49.958°]), rainforest, 25 m alt.; Parc National Mananara-Nord, 7.1 km 261° Antanambe ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-16.455%C2%B0%2C+49.7875%C2%B0&coordinates%5B0%5D=-16.455%C2%B0%2C+49.7875%C2%B0 -16.455°, 49.7875°]), rainforest, 225 m alt.; Res. Ambodiriana, 4.8 km 306° Manompana, along Manompana river ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-16.67233%C2%B0%2C+49.70117%C2%B0&coordinates%5B0%5D=-16.67233%C2%B0%2C+49.70117%C2%B0 -16.67233°, 49.70117°]), rainforest, 125 m alt.; Réserve Spéciale Ambatovaky, Sandrangato river ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-16.7633%C2%B0%2C+49.26692%C2%B0&coordinates%5B0%5D=-16.7633%C2%B0%2C+49.26692%C2%B0 -16.7633°, 49.26692°]), rainforest, 520 m alt.; ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-16.77274%C2%B0%2C+49.26551%C2%B0&coordinates%5B0%5D=-16.77274%C2%B0%2C+49.26551%C2%B0 -16.77274°, 49.26551°]), rainforest, 450 m alt.; ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-16.7755%C2%B0%2C+49.26427%C2%B0&coordinates%5B0%5D=-16.7755%C2%B0%2C+49.26427%C2%B0 -16.7755°, 49.26427°]), rainforest, 430 m alt.; (-16.81745°, 49.2925°), rainforest, 400 m alt.; ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-16.81753%C2%B0%2C+49.29498%C2%B0&coordinates%5B0%5D=-16.81753%C2%B0%2C+49.29498%C2%B0 -16.81753°, 49.29498°]), rainforest, 360 m alt.; Ile Sainte Marie, Forêt Kalalao, 9.9 km 34° Ambodifotatra ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-16.9225%C2%B0%2C+49.88733%C2%B0&coordinates%5B0%5D=-16.9225%C2%B0%2C+49.88733%C2%B0 -16.9225°, 49.88733°]), rainforest, 100 m alt.; Réserve Forestière Tampolo, 95.2 km N Toamasina ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.27808%C2%B0%2C+49.42853%C2%B0&coordinates%5B0%5D=-17.27808%C2%B0%2C+49.42853%C2%B0 -17.27808°, 49.42853°]), littoral rainforest, 20 m alt.; S.F. Tampolo, 10 km NNE Fenoarivo Atn. ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.2825%C2%B0%2C+49.43%C2%B0&coordinates%5B0%5D=-17.2825%C2%B0%2C+49.43%C2%B0 -17.2825°, 49.43°]), littoral rainforest, 10 m alt.; Bridge at Onibi, NW of Mahavelona ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.65%C2%B0%2C+49.46667%C2%B0&coordinates%5B0%5D=-17.65%C2%B0%2C+49.46667%C2%B0 -17.65°, 49.46667°]), in clay forest; Mahavelona (Foulpointe) ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.66667%C2%B0%2C+49.5%C2%B0&coordinates%5B0%5D=-17.66667%C2%B0%2C+49.5%C2%B0 -17.66667°, 49.5°]), Forest; Parc National de Zahamena, Besaky River ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.75244%C2%B0%2C+48.85321%C2%B0&coordinates%5B0%5D=-17.75244%C2%B0%2C+48.85321%C2%B0 -17.75244°, 48.85321°]), rainforest, 760 m alt.; Sahavorondrano River ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.75257%C2%B0%2C+48.85725%C2%B0&coordinates%5B0%5D=-17.75257%C2%B0%2C+48.85725%C2%B0 -17.75257°, 48.85725°]), rainforest, 765 m alt.; Onibe River ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.75908%C2%B0%2C+48.85468%C2%B0&coordinates%5B0%5D=-17.75908%C2%B0%2C+48.85468%C2%B0 -17.75908°, 48.85468°]), rainforest, 780 m alt.; Reserve Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.88667%C2%B0%2C+49.2025%C2%B0&coordinates%5B0%5D=-17.88667%C2%B0%2C+49.2025%C2%B0 -17.88667°, 49.2025°]), rainforest, 520 m alt.; Réserve Naturelle Betampona, 34.1 km 332° Toamasina ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.916135%C2%B0%2C+49.20185%C2%B0&coordinates%5B0%5D=-17.916135%C2%B0%2C+49.20185%C2%B0 -17.916135°, 49.20185°]), rainforest, 550 m alt.; Réserve Nationale Intégrale Betampona, Betampona 35.1 km NW Toamasina ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.91801%C2%B0%2C+49.20074%C2%B0&coordinates%5B0%5D=-17.91801%C2%B0%2C+49.20074%C2%B0 -17.91801°, 49.20074°]), rainforest, 500 m alt.; Reserve Betampona, Camp Rendrirendry 34.1 km 332° Toamasina ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-17.924%C2%B0%2C+49.19967%C2%B0&coordinates%5B0%5D=-17.924%C2%B0%2C+49.19967%C2%B0 -17.924°, 49.19967°]), rainforest, 390 m alt.; F.C. Sandranantitra ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-18.04833%C2%B0%2C+49.09167%C2%B0&coordinates%5B0%5D=-18.04833%C2%B0%2C+49.09167%C2%B0 -18.04833°, 49.09167°]), rainforest, 450 m alt.; F.C. Andriantantely ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-18.695%C2%B0%2C+48.81333%C2%B0&coordinates%5B0%5D=-18.695%C2%B0%2C+48.81333%C2%B0 -18.695°, 48.81333°]), rainforest, 530 m alt.; Sahafina forest 11.4 km W Brickaville ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-18.81445%C2%B0%2C+48.96205%C2%B0&coordinates%5B0%5D=-18.81445%C2%B0%2C+48.96205%C2%B0 -18.81445°, 48.96205°]), rainforest, 140 m alt. Mahajanga. Réserve Spéciale Marotandrano, Marotandrano 48.3 km S Mandritsara ([http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=-16.28322%C2%B0%2C+48.81443%C2%B0&coordinates%5B0%5D=-16.28322%C2%B0%2C+48.81443%C2%B0 -16.28322°, 48.81443°]), transition humid forest, 865 m alt.
  
−
Durch die laugen Kopfdornen des &#x263F; major, durch die Skulptur und durch die Behaarung von allen andern Arten verschieden, dem ''[[Mystrium_voeltzkowi|voeltzkowi]]'' am naechsten stehend, aber groesser und weniger schlank.}}
+
==Remarks==
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The worker of ''{{Taxon name|Mystrium oberthueri}}'' is easily distinguished from that of other ''{{Taxon name|Mystrium}}'' species by the combination of the following characters: strong, deep, and thick longitudinal sculpture on the dorsal and lateral surface of pronotum (Fig. 17B); straight anteromedial margin of the clypeus (Fig. 18A); a sharper angle between the dorsal and posterior faces of the vertex on the median line of the head (Fig. 16A). The queen of ''{{Taxon name|Mystrium oberthueri}}'' is differentiated from other queens of ''{{Taxon name|Mystrium}}'' species by the vestigial wings reduced to small appendages with undeveloped wing sclerites (ergatoid: as in Fig. 25D), the posterior face of the vertex forming approximately a right angle with its dorsal face on the median line of the head (as in Fig. 21A), straight anterior clypeal margin (Fig. 22B), and moderately developed clypeal conical setae (Fig. 23B). The male of ''{{Taxon name|Mystrium oberthueri}}'' can be separated from the other known ''{{Taxon name|Mystrium}}'' males by its smooth abdominal tergum VIII (as in Fig. 28), small lateral ocelli situated distant from posterior margin of head in full-face view, and straight dorsal margin of head in full-face view (Fig. 30A). We should note that the current male character diagnoses may not be able to distinguish between ''{{Taxon name|Mystrium oberthueri}}'' and ''{{Taxon name|Mystrium eques}}'', which is not yet known from the male caste and may be quite similar to ''{{Taxon name|Mystrium oberthueri}}''.<br />
−
{{Section|id=materials_examined|heading = Materials Examined|content=Insel Ste. Marie de Madagascar. Aus der Sammlung des Herrn R. Oberthuer, von Herrn Perrot gesammelt, durch Herrn Prof. Wasmann erhalten.}}
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The lectotype for ''{{Taxon name|Mystrium oberthueri}}'' is designated. According to Forel (1897)<ref name="B17">{{aut|Forel A}} (1897) Ameisen aus Nossi-Bé, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar, gesammelt von Herrn Dr. A. Voeltzkow aus Berlin. Mit einem Anhang über die von Herrn Privatdocenten Dr. A. Brauer in Marburg auf den Seychellen und von Herrn Perrot auf Ste. Marie (Madagaskar) gesammelten Ameisen. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 21: 185-208.</ref >, a single major worker and multiple ergatoid queens (as minor workers) were used for the original description of ''{{Taxon name|Mystrium oberthueri}}''. We confirmed one worker and one ergatoid queen in the Forel collection in MHNG and one ergatoid queen in MCZC, all of which are labeled with the location “Ste. Marie Madagascar.” The single worker was chosen as the new representative specimen.<br />
 +
Although worker descriptions for ''{{Taxon name|Mystrium oberthueri}}'' are reliable, male descriptions of this species in previous papers are problematic. The original description by Forel (1897)<ref name="B17">{{aut|Forel A}} (1897) Ameisen aus Nossi-Bé, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar, gesammelt von Herrn Dr. A. Voeltzkow aus Berlin. Mit einem Anhang über die von Herrn Privatdocenten Dr. A. Brauer in Marburg auf den Seychellen und von Herrn Perrot auf Ste. Marie (Madagaskar) gesammelten Ameisen. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 21: 185-208.</ref > did not include males. Later male descriptions were given by Emery (1899)<ref name="B9">{{aut|Emery C}} (1899) Formiche di Madagascar raccolte dal Sig. A. Mocquerys nei pressi della Baia di Antongil (1897–1898). Bullettino della Societa Entomologica Italiana 31: 263-290.</ref > and Menozzi (1929)<ref name="B27">{{aut|Menozzi C}} (1929) Revisione delle formiche del genere ''Mystrium'' Roger. Zoologischer Anzeiger 82: 518-536.</ref >. As in the cases of ''{{Taxon name|Mystrium mysticum}}'' and ''{{Taxon name|Mystrium rogeri}}'', Emery’s and Menozzi’s descriptions of the male of ''{{Taxon name|Mystrium oberthueri}}'' seem to be based on specimens not associated with conspecific workers. We did not examine the actual male specimens they used, and cannot be sure that these descriptions are for the actual male of ''{{Taxon name|Mystrium oberthueri}}''. Emery mentioned Forel’s description (Forel 1899<ref name="B18">{{aut|Forel A}} (1899) Trois notices myrmécologiques. Annales de la Société Entomologique de Belgique 43: 303-310.</ref >) of ''{{Taxon name|Mystrium rogeri}}'' in the same paper. We have confirmed the male described as ''{{Taxon name|Mystrium rogeri}}'' was the actual ''{{Taxon name|Mystrium oberthueri}}''; however, Emery (1899)<ref name="B9">{{aut|Emery C}} (1899) Formiche di Madagascar raccolte dal Sig. A. Mocquerys nei pressi della Baia di Antongil (1897–1898). Bullettino della Societa Entomologica Italiana 31: 263-290.</ref > did not mention this problem. Moreover, Emery listed smaller body size as a character able to distinguish the male of ''{{Taxon name|Mystrium oberthueri}}'' from that of ''{{Taxon name|Mystrium mysticum}}''; however, according to our measurements, ''{{Taxon name|Mystrium oberthueri}}'' is the largest of the ''{{Taxon name|Mystrium}}'' males (''{{Taxon name|Mystrium oberthueri}}'': HW>2.00 mm vs. other known males: HW<1.88 mm). We believe that Emery’s male description for ''{{Taxon name|Mystrium oberthueri}}'' is doubtful, even though we have not been able to determine the males described in either ''{{Taxon name|Mystrium oberthueri}}'' or ''{{Taxon name|Mystrium mysticum}}'' in Emery (1899)<ref name="B9">{{aut|Emery C}} (1899) Formiche di Madagascar raccolte dal Sig. A. Mocquerys nei pressi della Baia di Antongil (1897–1898). Bullettino della Societa Entomologica Italiana 31: 263-290.</ref >. Second, Menozzi (1929)<ref name="B27">{{aut|Menozzi C}} (1929) Revisione delle formiche del genere ''Mystrium'' Roger. Zoologischer Anzeiger 82: 518-536.</ref > examined all types in the Forel collection, including those of ''{{Taxon name|Mystrium rogeri}}''. However, he did not mention anything in the male description of ''{{Taxon name|Mystrium oberthueri}}'' about the male in the syntypes of ''{{Taxon name|Mystrium rogeri}}''. Hence, the male description of the ''{{Taxon name|Mystrium oberthueri}}'' in Menozzi (1929)<ref name="B27">{{aut|Menozzi C}} (1929) Revisione delle formiche del genere ''Mystrium'' Roger. Zoologischer Anzeiger 82: 518-536.</ref > is also not likely to be the true ''{{Taxon name|Mystrium oberthueri}}'', although it probably is a male in the ''voeltzkowi'' species group.<br />
 +
We describe a new species, ''{{Taxon name|Mystrium eques}}'', with a worker similar to that of ''{{Taxon name|Mystrium oberthueri}}''. As we have already mentioned in the section of ''{{Taxon name|Mystrium eques}}'', the unknown male of ''{{Taxon name|Mystrium eques}}'' is likely to be quite similar to that of ''{{Taxon name|Mystrium oberthueri}}'', and the current male diagnosis for ''{{Taxon name|Mystrium oberthueri}}'' may be insufficient to distinguish these two males (see ''{{Taxon name|Mystrium eques}}'').<br />
 +
Material identified as ''{{Taxon name|Mystrium oberthueri}}'' in Molet et al. (2009)<ref name="B28">{{aut|Molet M}}, {{aut|Fisher B}}, {{aut|Ito F}}, {{aut|Peeters C}} (2009) Shift from independent to dependent colony foundation and evolution of ‘multi-purpose’ ergatoid queens in ''Mystrium'' ants (subfamily Amblyoponinae). Biological journal of the Linnean Society 98: 198-207. doi: [http://dx.doi.org/10.1111/j.1095-8312.2009.01257.x 10.1111/j.1095-8312.2009.01257.x]</ref > and Bouchet et al. (2013)<ref name="B4">{{aut|Bouchet D}}, {{aut|Peeters C}}, {{aut|Fisher B}}, {{aut|Molet M}} (2013) Both female castes contribute to colony emigration in the polygynous ant ''Mystrium oberthueri''. Ecological Entomology 38: 408-417. doi: [http://dx.doi.org/10.1111/een.12033 10.1111/een.12033]</ref > is confirmed here.
  
 
==Taxon Treatment==
 
==Taxon Treatment==
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*{{aut|Forel, A.}}; 1897: Ameisen aus Nossi-Be, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar. Gesammelt von Herrn Dr. A. Voeltzkow aus Berlin., Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft '''21''': 192-193. {{doi|TODO}}
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*{{aut|Yoshimura, M}}; {{aut|Fisher, B}}; 2014: A revision of the ant genus ''Mystrium'' in the Malagasy region with description of six new species and remarks on ''Amblyopone'' and ''Stigmatomma'' (Hymenoptera, Formicidae, Amblyoponinae) [http://www.pensoft.net/journals/zookeys/ ''ZooKeys'',] '''394''': 1-99. {{doi|10.3897/zookeys.394.6446}}
  
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[[Category:TODO]]
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==Other References==
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 +
<references />
 +
 
 +
==Images==
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{{Gallery | lines=5 | width=250
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|1= File:zookeys-394-001-g001.jpg|2= '''Figure 1.''' Colonies of ''{{Taxon name|Mystrium}}''. '''A''' ''{{Taxon name|Mystrium oberthueri}}'' '''B''' ''{{Taxon name|Mystrium voeltzkowi}}''. '''A''' a cocoon and a worker '''B''' larvae, workers, and ergatoid queens. Photos by Alex Wild.
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|3= File:zookeys-394-001-g016.jpg|4= '''Figure 16.''' Vertex of workers in ''voeltzkowi'' species group. '''A''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0499569 CASENT0499569]) '''B''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0002075 CASENT0002075]). '''A, B''' head in posterior view.
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|5= File:zookeys-394-001-g017.jpg|6= '''Figure 17.''' Differences in worker morphology between ''{{Taxon name|Mystrium janovitzi}}'' and ''{{Taxon name|Mystrium oberthueri}}''. '''A''' ''{{Taxon name|Mystrium janovitzi}}'' ([http://data.antweb.org/specimen/CASENT0006082 CASENT0006082]) '''B''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0499569 CASENT0499569]) '''C''' ''{{Taxon name|Mystrium janovitzi}}'' ([http://data.antweb.org/specimen/CASENT0482698 CASENT0482698]: holotype) '''D''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496066 CASENT0496066]). '''A, B''' mesosoma in dorsal view '''C, D''' clypeus and genal tooth in full-face view.
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|7= File:zookeys-394-001-g018.jpg|8= '''Figure 18.''' Differences in worker morphology between ''{{Taxon name|Mystrium oberthueri}}'' and ''{{Taxon name|Mystrium eques}}''. '''A''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496066 CASENT0496066]) '''B''' ''{{Taxon name|Mystrium eques}}'' ([http://data.antweb.org/specimen/CASENT0418314 CASENT0418314]: holotype) '''C, E''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0499569 CASENT0499569]) '''D, F''' ''{{Taxon name|Mystrium eques}}'' ([http://data.antweb.org/specimen/CASENT0317399 CASENT0317399]). '''A, B''' head in full-face view '''C, D''' pedicel and flagellum in lateral view '''E, F''' mandible in dorsal view.
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|9= File:zookeys-394-001-g021.jpg|10= '''Figure 21.''' Vertex of ergatoid queens in ''voeltzkowi'' species group. '''A''' ''{{Taxon name|Mystrium eques}}'' ([http://data.antweb.org/specimen/CASENT0418311 CASENT0418311]) '''B''' ''{{Taxon name|Mystrium shadow}}'' ([http://data.antweb.org/specimen/CASENT0077647 CASENT0077647]: paratype). '''A, B''' head in posterior view.
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|11= File:zookeys-394-001-g022.jpg|12= '''Figure 22.''' Clypeus of ergatoid queens in ''voeltzkowi'' species group. '''A''' ''{{Taxon name|Mystrium eques}}'' ([http://data.antweb.org/specimen/CASENT0418317 CASENT0418317]: paratype) '''B''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496064 CASENT0496064]). '''A, B''' head in full-face view.
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|13= File:zookeys-394-001-g023.jpg|14= '''Figure 23.''' Differences in ergatoid queen characters between ''{{Taxon name|Mystrium janovitzi}}'' and ''{{Taxon name|Mystrium oberthueri}}''. '''A''' ''{{Taxon name|Mystrium janovitzi}}'' ([http://data.antweb.org/specimen/CASENT0482696 CASENT0482696]) '''B''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496064 CASENT0496064]) '''C''' ''{{Taxon name|Mystrium janovitzi}}'' ([http://data.antweb.org/specimen/CASENT0003302 CASENT0003302]) '''D''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0095743 CASENT0095743]). '''A, B''' head in full-face view '''C, D''' pedicel and flagellum in lateral view.
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|15= File:zookeys-394-001-g025.jpg|16= '''Figure 25.''' Differences in ergatoid queen characters between ''{{Taxon name|Mystrium mirror}}'' and ''{{Taxon name|Mystrium shadow}}''. '''A, C''' ''{{Taxon name|Mystrium mirror}}'' ([http://data.antweb.org/specimen/CASENT0429899 CASENT0429899]) '''B, D''' ''{{Taxon name|Mystrium shadow}}'' ([http://data.antweb.org/specimen/CASENT0077647 CASENT0077647]: paratype). '''A, B''' head and pronotum in oblique dorsal view '''C, D''' mesosoma in oblique dorsal view.
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|17= File:zookeys-394-001-g028.jpg|18= '''Figure 28.''' Abdomen of ''{{Taxon name|Mystrium}}'' males. '''A''' ''{{Taxon name|Mystrium mysticum}}'' ([http://data.antweb.org/specimen/CASENT0318027 CASENT0318027]) '''B''' ''{{Taxon name|Mystrium janovitzi}}'' ([http://data.antweb.org/specimen/CASENT0007820 CASENT0007820]) '''C''' ''{{Taxon name|Mystrium mysticum}}'' ([http://data.antweb.org/specimen/CASENT0429983 CASENT0429983]) '''D''' ''{{Taxon name|Mystrium shadow}}'' ([http://data.antweb.org/specimen/CASENT0303561 CASENT0303561]). '''A, B''' abdominal segment VIII in oblique lateral view '''C, D''' petiole in dorsal view.
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|19= File:zookeys-394-001-g030.jpg|20= '''Figure 30.''' Head of ''{{Taxon name|Mystrium}}'' males in full-face view. '''A''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0179499 CASENT0179499]) '''B''' ''{{Taxon name|Mystrium shadow}}'' ([http://data.antweb.org/specimen/CASENT0107647 CASENT0107647]).
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|21= File:zookeys-394-001-g037.jpg|22= '''Figure 37.''' Males of ''{{Taxon name|Mystrium}}'' in lateral view. '''A''' ''{{Taxon name|Mystrium barrybressleri}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0135866 CASENT0135866]) '''B''' ''{{Taxon name|Mystrium mysticum}}'' ([http://data.antweb.org/specimen/CASENT0318027 CASENT0318027]) '''C''' ''{{Taxon name|Mystrium rogeri}}'' ([http://data.antweb.org/specimen/CASENT0001083 CASENT0001083]) '''D''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0007820 CASENT0007820]) '''E''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0318941 CASENT0318941]) '''F''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0179499 CASENT0179499]) '''G''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0107647 CASENT0107647]) '''H''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0101673 CASENT0101673]).
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|23= File:zookeys-394-001-g038.jpg|24= '''Figure 38.''' Head of ''{{Taxon name|Mystrium}}'' males in full-face view. '''A''' ''{{Taxon name|Mystrium barrybressleri}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0135866 CASENT0135866]) '''B''' ''{{Taxon name|Mystrium mysticum}}'' ([http://data.antweb.org/specimen/CASENT0318027 CASENT0318027]) '''C''' ''{{Taxon name|Mystrium rogeri}}'' ([http://data.antweb.org/specimen/CASENT0001083 CASENT0001083]) '''D''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0007820 CASENT0007820]) '''E''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0318941 CASENT0318941]) '''F''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0179499 CASENT0179499]) '''G''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0107647 CASENT0107647]) '''H''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0101673 CASENT0101673]).
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|25= File:zookeys-394-001-g039.jpg|26= '''Figure 39.''' Males of ''{{Taxon name|Mystrium}}'' in dorsal view. '''A''' ''{{Taxon name|Mystrium barrybressleri}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0135866 CASENT0135866]) '''B''' ''{{Taxon name|Mystrium mysticum}}'' ([http://data.antweb.org/specimen/CASENT0318027 CASENT0318027]) '''C''' ''{{Taxon name|Mystrium rogeri}}'' ([http://data.antweb.org/specimen/CASENT0001083 CASENT0001083]) '''D''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0007820 CASENT0007820]) '''E''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0318941 CASENT0318941]) '''F''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0179499 CASENT0179499]) '''G''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0107647 CASENT0107647]) '''H''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0101673 CASENT0101673]).
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|27= File:zookeys-394-001-g040.jpg|28= '''Figure 40.''' Genital capsule of ''{{Taxon name|Mystrium}}'' males in lateral view. '''A''' ''{{Taxon name|Mystrium barrybressleri}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0078803 CASENT0078803]) '''B''' ''{{Taxon name|Mystrium mysticum}}'' ([http://data.antweb.org/specimen/CASENT0218102 CASENT0218102]) '''C''' ''{{Taxon name|Mystrium rogeri}}'' ([http://data.antweb.org/specimen/CASENT0001152 CASENT0001152]) '''D''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0080644 CASENT0080644]) '''E''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0317444 CASENT0317444]) '''F''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0317422 CASENT0317422]) '''G''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0107647 CASENT0107647]) '''H''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0317583 CASENT0317583]).
 +
|29= File:zookeys-394-001-g041.jpg|30= '''Figure 41.''' Aedeagus of ''{{Taxon name|Mystrium}}'' males in lateral view. '''A''' ''{{Taxon name|Mystrium barrybressleri}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0078803 CASENT0078803]) '''B''' ''{{Taxon name|Mystrium mysticum}}'' ([http://data.antweb.org/specimen/CASENT0218102 CASENT0218102]) '''C''' ''{{Taxon name|Mystrium rogeri}}'' ([http://data.antweb.org/specimen/CASENT0001152 CASENT0001152]) '''D''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0080644 CASENT0080644]) '''E''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0317444 CASENT0317444]) '''F''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0317422 CASENT0317422]) '''G''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0107647 CASENT0107647]) '''H''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0317583 CASENT0317583]).
 +
|31= File:zookeys-394-001-g050.jpg|32= '''Figure 50.''' Workers of ''voeltzkowi'' species group in lateral view. '''A''' ''{{Taxon name|Mystrium eques}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0418314 CASENT0418314]: holotype) '''B''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0482698 CASENT0482698]: holotype) '''C''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0429897 CASENT0429897]: holotype) '''D''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496066 CASENT0496066]) '''E''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0318933 CASENT0318933]: holotype) '''F''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0002075 CASENT0002075]).
 +
|33= File:zookeys-394-001-g051.jpg|34= '''Figure 51.''' Head of the workers of ''voeltzkowi'' species group in full-face view. '''A''' ''{{Taxon name|Mystrium eques}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0418314 CASENT0418314]: holotype) '''B''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0482698 CASENT0482698]: holotype) '''C''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0429897 CASENT0429897]: holotype) '''D''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496066 CASENT0496066]) '''E''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0318933 CASENT0318933]: holotype) '''F''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0002075 CASENT0002075]).
 +
|35= File:zookeys-394-001-g052.jpg|36= '''Figure 52.''' Workers of ''voeltzkowi'' species group in dorsal view. '''A''' ''{{Taxon name|Mystrium eques}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0418314 CASENT0418314]: holotype) '''B''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0482698 CASENT0482698]: holotype) '''C''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0429897 CASENT0429897]: holotype) '''D''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496066 CASENT0496066]) '''E''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0318933 CASENT0318933]: holotype) '''F''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0002075 CASENT0002075]).
 +
|37= File:zookeys-394-001-g053.jpg|38= '''Figure 53.''' Ergatoid queens of ''voeltzkowi'' species group in lateral view. '''A''' ''{{Taxon name|Mystrium eques}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0418317 CASENT0418317]: paratype) '''B''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0482696 CASENT0482696]) '''C''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0318940 CASENT0318940]: paratype) '''D''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496064 CASENT0496064]) '''E''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0077647 CASENT0077647]: paratype) '''F''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0002092 CASENT0002092]).
 +
|39= File:zookeys-394-001-g054.jpg|40= '''Figure 54.''' Head of the ergatoid queens of ''voeltzkowi'' species group in full-face view. '''A''' ''{{Taxon name|Mystrium eques}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0418317 CASENT0418317]: paratype) '''B''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0482696 CASENT0482696]) '''C''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0318940 CASENT0318940]: paratype) '''D''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496064 CASENT0496064]) '''E''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0077647 CASENT0077647]: paratype) '''F''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0002092 CASENT0002092]).
 +
|41= File:zookeys-394-001-g055.jpg|42= '''Figure 55.''' Ergatoid queens of ''voeltzkowi'' species group in dorsal view. '''A''' ''{{Taxon name|Mystrium eques}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0418317 CASENT0418317]: paratype) '''B''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0482696 CASENT0482696]) '''C''' ''{{Taxon name|Mystrium mirror}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0318940 CASENT0318940]: paratype) '''D''' ''{{Taxon name|Mystrium oberthueri}}'' ([http://data.antweb.org/specimen/CASENT0496064 CASENT0496064]) '''E''' ''{{Taxon name|Mystrium shadow}}'' sp. n. ([http://data.antweb.org/specimen/CASENT0077647 CASENT0077647]: paratype) '''F''' ''{{Taxon name|Mystrium voeltzkowi}}'' ([http://data.antweb.org/specimen/CASENT0002092 CASENT0002092]).
 +
|43= File:zookeys-394-001-g056.jpg|44= '''Figure 56.''' Distribution maps for ''voeltzkowi'' species group. '''A''' ''{{Taxon name|Mystrium eques}}'' sp. n. '''B''' ''{{Taxon name|Mystrium janovitzi}}'' sp. n. '''C''' ''{{Taxon name|Mystrium mirror}}'' sp. n. '''D''' ''{{Taxon name|Mystrium oberthueri}}'' '''E''' ''{{Taxon name|Mystrium shadow}}'' sp. n. '''F''' ''{{Taxon name|Mystrium voeltzkowi}}''. Star symbols represent the type locality: the localities for ''{{Taxon name|Mystrium oberthueri}}'' and ''{{Taxon name|Mystrium voeltzkowi}}'' are approximate, which are assumed from the original descriptions.
 +
}}

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Yoshimura M, Fisher B (2014) A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae). ZooKeys 394 : 1–99, doi. Versioned wiki page: 2014-03-31, version 43936, https://species-id.net/w/index.php?title=Mystrium_oberthueri&oldid=43936 , contributors (alphabetical order): PlaziBot, Pensoft Publishers.

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@article{Yoshimura2014ZooKeys394,
author = {Yoshimura, Masashi AND Fisher, Brian L.},
journal = {ZooKeys},
publisher = {Pensoft Publishers},
title = {A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae)},
year = {2014},
volume = {394},
issue = {},
pages = {1--99},
doi = {10.3897/zookeys.394.6446},
url = {http://www.pensoft.net/journals/zookeys/article/6446/abstract},
note = {Versioned wiki page: 2014-03-31, version 43936, https://species-id.net/w/index.php?title=Mystrium_oberthueri&oldid=43936 , contributors (alphabetical order): PlaziBot, Pensoft Publishers.}

}

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TY - JOUR
T1 - A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae)
A1 - Yoshimura M
A1 - Fisher B
Y1 - 2014
JF - ZooKeys
JA -
VL - 394
IS -
UR - http://dx.doi.org/10.3897/zookeys.394.6446
SP - 1
EP - 99
PB - Pensoft Publishers
M1 - Versioned wiki page: 2014-03-31, version 43936, https://species-id.net/w/index.php?title=Mystrium_oberthueri&oldid=43936 , contributors (alphabetical order): PlaziBot, Pensoft Publishers.

M3 - doi:10.3897/zookeys.394.6446

Wikipedia/ Citizendium:

<ref name="Yoshimura2014ZooKeys394">{{Citation
| author = Yoshimura M, Fisher B
| title = A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae)
| journal = ZooKeys
| year = 2014
| volume = 394
| issue =
| pages = 1--99
| pmid =
| publisher = Pensoft Publishers
| doi = 10.3897/zookeys.394.6446
| url = http://www.pensoft.net/journals/zookeys/article/6446/abstract
| pmc =
| accessdate = 2024-12-23

}} Versioned wiki page: 2014-03-31, version 43936, https://species-id.net/w/index.php?title=Mystrium_oberthueri&oldid=43936 , contributors (alphabetical order): PlaziBot, Pensoft Publishers.</ref>

See also the citation download page at the journal.


Taxonavigation

Ordo: Hymenoptera
Familia: Formicidae
Genus: Mystrium

Name

Mystrium oberthueri Forel, 1897 – Wikispecies link – Pensoft Profile

  • Mystrium oberthueri Forel, 1897. MADAGASCAR. Syntypes: one worker (as major) and two ergatoid queens (as minor) [lectotype is designated below].
  • Mystrium oberthueri: Menozzi 1929[1] (in part?: uncertain male identification)
  • Mystrium oberthueri (Not) (male): Emery 1899[2].
  • Mystrium rogeri (male): Forel 1899[3].

Lectotype

[here designated]. Worker: CASENT0101993, MADAGASCAR, Insel Ste. Marie (Island of Sainte Marie), Perrot [MHNG: examined].

Worker

Description. Measurements: lectotype. HL 2.40, HW 2.61, SL 1.80, ML 2.92, HD 1.67, WL 2.79, PnW 1.39, PpW 1.21, PtW 1.18, PtL 0.81, CI 108.7, SI 69.0, MI 111.8, PpI 87.4, PtI 144.9.
HL 2.00–2.58, HW 2.12–2.76, SL 1.55–1.96, ML 2.39–3.17, HD 1.33–1.69, WL 2.43–2.96, PnW 1.17–1.38, PpW 1.02–1.29, PtW 1.00–1.32, PtL 0.67–0.81, CI 104.4–108.5, SI 68.8–74.9, MI 106.9–122.9, PpI 87.2–93.8, PtI 139.5–163.2 (10 specimens measured).
Posterolateral corner of head moderately expanding posteriorly. Posterior face of vertex forming almost right angle with dorsal face on median line of head, so that declivity of vertex on lateral part as steep as on median part. Whole region of vertex finely striated. Eye moderately small. Anterior margin of clypeus straight or weakly convex with long conical setae. Genal tooth of head relatively long, as long as lateral lobe of clypeus. Masticatory surface of mandible almost invisible in full-face view, and width of dorsal surface of mandible almost identical from mandibular shaft to distal portion. Second maxillary palpomere longer than third. First flagellomere (third antennal segment) about 1.2–1.5× length of pedicel (second antennal segment). Strong, deep longitudinal striae regularly impressed on whole central part of pronotal dorsum. Strong, deep longitudinal striae impressed on lateral surface of pronotum. Mesonotum differentiated from propodeum in dorsal view, length slightly shorter than that of propodeum. Metanotal groove shallowly and gently impressed in lateral view. Metapleural gland bulla moderately developed, and propodeal declivity in lateral view almost straight. Petiole widened on posterior 1/3 and gently narrowing anteriorly in dorsal view, anterior margin straight to gently rounded and often edged by thick striae.
Body color black.

Ergatoid queen

Description. Measurements: HL 1.40–1.93, HW 1.46–1.96, SL 1.07–1.47, ML 1.49–2.00, HD 1.00–1.35, WL 1.92–2.56, PnW 0.83–1.11, PpW 0.85–1.13, PtW 0.81–1.07, PtL 0.53–0.70, CI 101.6–108.6, SI 71.6–76.7, MI 96.3–108.8, PpI 96.0–104.4, PtI 149.1–177.1 (10 specimens measured).
Wings vestigial and reduced to small but distinct appendages. Wing sclerites undeveloped. Posterolateral corner of head moderately expanding posteriorly, expansion not differentiated from that in workers. Posterior face of vertex forming almost a right angle with dorsal face on median line of head, so that declivity of vertex on lateral part as steep as on median part. Ventral half of vertex sculptured. Eye small but distinct. Ocelli absent. Anterior margin of clypeus almost straight with long to moderate conical setae. Anterolateral portion of head with short spine. Masticatory margin of mandible almost invisible in full-face view, and dorsal surface on distal portion as wide as that on mandibular shaft. Spatulate seta present on basal side of each basal denticle on masticatory margin of mandible. First flagellomere (third antennal segment) long, about 1.2-1.5× length of pedicel (second antennal segment). Setae on pronotum distinctly spatulate, widened distally with sharp or blunt apex. Metapleural gland bulla moderately developed, not expanding dorsally to propodeal spiracle, so that propodeal declivity in lateral view weakly convex and rounded posteriorly on its ventral 1/3. Petiole relatively long in dorsal view, about 0.8× length of abdominal segment III.
Body color blackish to reddish brown.

Male

Description. Measurements: HL 1.44, HW 2.09, SL 0.62, EL 0.95, WL 3.55, MnW 1.94, CI 145.1, SI 29.6, EI 66.2, MnI 92.9 (1 specimen measured).
Eye moderately large, occupying 0.6× of head length. Ocelli relatively distant from dorsal margin of head in full-face view. Dorsal margin of head in full-face view straight. Both anterior and lateral ocelli small. Lateral ocellus small and distant from eye: distance between these more than 1.5× maximum diameter of lateral ocellus. Posterior face of vertex not clearly differentiated from dorsal face, so that vertex almost continuously rounded. Palpal formula 4,3. First maxillary palpomere flattened and distinctly wider than second segment. Second maxillary palpomere longer than third. Notauli clearly impressed on mesoscutum. Petiole in dorsal view thin, 0.8× as long as abdominal tergite III. Petiolar dorsum covered with shallow, irregular punctures. Abdominal tergum VIII without deep punctures, almost smooth.
Distal portion of abdominal sternum IX smooth and not punctured. Basal ring moderately long, expanding basoventrally. Telomere distinctly extending distally farther than digitus. Basoventral expansion of aedeagus moderately developed basoventrally, longer than dorsal extension. Ventral margin of aedeagus gently convex in lateral view. Aedeagus moderately narrowing distally and distal portion rounded.
On forewing, cu-a located far basal from junction of Media (M) and Cubitus (Cu).
Body color reddish brown to black.

Distribution

MADAGASCAR: as in Figure 56D.

Additional material examined

In addition to the type material, specimens from the following localities were examined in this study: MADAGASCAR. Antsiranana. Forêt d’ Antsahabe, 11.4 km 275° W Daraina (-13.21167°, 49.55667°), tropical dry forest, 550 m alt.; Makirovana forest (-14.17066°, 49.95409°), rainforest, 415 m alt.; Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina (-14.435°, 49.76°), rainforest, 775 m alt.; 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina (-14.43667°, 49.775°), rainforest, 450 m alt.; Marojejy R.N.I. #12 (-14.43583°, 49.76056°), rainforest, 610 m alt.; Forêt Ambanitaza, 26.1 km 347° Antalaha (-14.67933°, 50.18367°), rainforest, 240 m alt.; 1 km W Andampibe, Cap Masoala (-15.69361°, 50.18139°), lowland rainforest, 125 m alt.; Fotodriana, Cap Masoala (-15.69694°, 50.27028°), lowland rainforest, 25 m alt.; Toamasina. Montagne d’Anjanaharibe, 18.0 km 21° NNE Ambinanitelo (-15.18833°, 49.615°), rainforest, 470 m alt.; Montagne d’Akirindro 7.6 km 341° NNW Ambinanitelo (-15.28833°, 49.54833°), rainforest, 600 m alt.; 19 km ESE Maroantsetra (-15.48333°, 49.9°), rainforest, 350 m alt.; 5.3 km SSE Ambanizana, Andranobe (-15.66667°, 49.96667°), rainforest, 425 m alt.; 6.3 km S Ambanizana, Andranobe (-15.6813°, 49.958°), rainforest, 5 m alt.; 6.3 km S Ambanizana, Andranobe (-15.68131°, 49.958°), rainforest, 25 m alt.; Parc National Mananara-Nord, 7.1 km 261° Antanambe (-16.455°, 49.7875°), rainforest, 225 m alt.; Res. Ambodiriana, 4.8 km 306° Manompana, along Manompana river (-16.67233°, 49.70117°), rainforest, 125 m alt.; Réserve Spéciale Ambatovaky, Sandrangato river (-16.7633°, 49.26692°), rainforest, 520 m alt.; (-16.77274°, 49.26551°), rainforest, 450 m alt.; (-16.7755°, 49.26427°), rainforest, 430 m alt.; (-16.81745°, 49.2925°), rainforest, 400 m alt.; (-16.81753°, 49.29498°), rainforest, 360 m alt.; Ile Sainte Marie, Forêt Kalalao, 9.9 km 34° Ambodifotatra (-16.9225°, 49.88733°), rainforest, 100 m alt.; Réserve Forestière Tampolo, 95.2 km N Toamasina (-17.27808°, 49.42853°), littoral rainforest, 20 m alt.; S.F. Tampolo, 10 km NNE Fenoarivo Atn. (-17.2825°, 49.43°), littoral rainforest, 10 m alt.; Bridge at Onibi, NW of Mahavelona (-17.65°, 49.46667°), in clay forest; Mahavelona (Foulpointe) (-17.66667°, 49.5°), Forest; Parc National de Zahamena, Besaky River (-17.75244°, 48.85321°), rainforest, 760 m alt.; Sahavorondrano River (-17.75257°, 48.85725°), rainforest, 765 m alt.; Onibe River (-17.75908°, 48.85468°), rainforest, 780 m alt.; Reserve Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina (-17.88667°, 49.2025°), rainforest, 520 m alt.; Réserve Naturelle Betampona, 34.1 km 332° Toamasina (-17.916135°, 49.20185°), rainforest, 550 m alt.; Réserve Nationale Intégrale Betampona, Betampona 35.1 km NW Toamasina (-17.91801°, 49.20074°), rainforest, 500 m alt.; Reserve Betampona, Camp Rendrirendry 34.1 km 332° Toamasina (-17.924°, 49.19967°), rainforest, 390 m alt.; F.C. Sandranantitra (-18.04833°, 49.09167°), rainforest, 450 m alt.; F.C. Andriantantely (-18.695°, 48.81333°), rainforest, 530 m alt.; Sahafina forest 11.4 km W Brickaville (-18.81445°, 48.96205°), rainforest, 140 m alt. Mahajanga. Réserve Spéciale Marotandrano, Marotandrano 48.3 km S Mandritsara (-16.28322°, 48.81443°), transition humid forest, 865 m alt.

Remarks

The worker of Mystrium oberthueri is easily distinguished from that of other Mystrium species by the combination of the following characters: strong, deep, and thick longitudinal sculpture on the dorsal and lateral surface of pronotum (Fig. 17B); straight anteromedial margin of the clypeus (Fig. 18A); a sharper angle between the dorsal and posterior faces of the vertex on the median line of the head (Fig. 16A). The queen of Mystrium oberthueri is differentiated from other queens of Mystrium species by the vestigial wings reduced to small appendages with undeveloped wing sclerites (ergatoid: as in Fig. 25D), the posterior face of the vertex forming approximately a right angle with its dorsal face on the median line of the head (as in Fig. 21A), straight anterior clypeal margin (Fig. 22B), and moderately developed clypeal conical setae (Fig. 23B). The male of Mystrium oberthueri can be separated from the other known Mystrium males by its smooth abdominal tergum VIII (as in Fig. 28), small lateral ocelli situated distant from posterior margin of head in full-face view, and straight dorsal margin of head in full-face view (Fig. 30A). We should note that the current male character diagnoses may not be able to distinguish between Mystrium oberthueri and Mystrium eques, which is not yet known from the male caste and may be quite similar to Mystrium oberthueri.
The lectotype for Mystrium oberthueri is designated. According to Forel (1897)[4], a single major worker and multiple ergatoid queens (as minor workers) were used for the original description of Mystrium oberthueri. We confirmed one worker and one ergatoid queen in the Forel collection in MHNG and one ergatoid queen in MCZC, all of which are labeled with the location “Ste. Marie Madagascar.” The single worker was chosen as the new representative specimen.
Although worker descriptions for Mystrium oberthueri are reliable, male descriptions of this species in previous papers are problematic. The original description by Forel (1897)[4] did not include males. Later male descriptions were given by Emery (1899)[2] and Menozzi (1929)[1]. As in the cases of Mystrium mysticum and Mystrium rogeri, Emery’s and Menozzi’s descriptions of the male of Mystrium oberthueri seem to be based on specimens not associated with conspecific workers. We did not examine the actual male specimens they used, and cannot be sure that these descriptions are for the actual male of Mystrium oberthueri. Emery mentioned Forel’s description (Forel 1899[3]) of Mystrium rogeri in the same paper. We have confirmed the male described as Mystrium rogeri was the actual Mystrium oberthueri; however, Emery (1899)[2] did not mention this problem. Moreover, Emery listed smaller body size as a character able to distinguish the male of Mystrium oberthueri from that of Mystrium mysticum; however, according to our measurements, Mystrium oberthueri is the largest of the Mystrium males (Mystrium oberthueri: HW>2.00 mm vs. other known males: HW<1.88 mm). We believe that Emery’s male description for Mystrium oberthueri is doubtful, even though we have not been able to determine the males described in either Mystrium oberthueri or Mystrium mysticum in Emery (1899)[2]. Second, Menozzi (1929)[1] examined all types in the Forel collection, including those of Mystrium rogeri. However, he did not mention anything in the male description of Mystrium oberthueri about the male in the syntypes of Mystrium rogeri. Hence, the male description of the Mystrium oberthueri in Menozzi (1929)[1] is also not likely to be the true Mystrium oberthueri, although it probably is a male in the voeltzkowi species group.
We describe a new species, Mystrium eques, with a worker similar to that of Mystrium oberthueri. As we have already mentioned in the section of Mystrium eques, the unknown male of Mystrium eques is likely to be quite similar to that of Mystrium oberthueri, and the current male diagnosis for Mystrium oberthueri may be insufficient to distinguish these two males (see Mystrium eques).
Material identified as Mystrium oberthueri in Molet et al. (2009)[5] and Bouchet et al. (2013)[6] is confirmed here.

Taxon Treatment

  • Yoshimura, M; Fisher, B; 2014: A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae) ZooKeys, 394: 1-99. doi

Other References

  1. ↑ 1.0 1.1 1.2 1.3 Menozzi C (1929) Revisione delle formiche del genere Mystrium Roger. Zoologischer Anzeiger 82: 518-536.
  2. ↑ 2.0 2.1 2.2 2.3 Emery C (1899) Formiche di Madagascar raccolte dal Sig. A. Mocquerys nei pressi della Baia di Antongil (1897–1898). Bullettino della Societa Entomologica Italiana 31: 263-290.
  3. ↑ 3.0 3.1 Forel A (1899) Trois notices myrmécologiques. Annales de la Société Entomologique de Belgique 43: 303-310.
  4. ↑ 4.0 4.1 Forel A (1897) Ameisen aus Nossi-Bé, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar, gesammelt von Herrn Dr. A. Voeltzkow aus Berlin. Mit einem Anhang über die von Herrn Privatdocenten Dr. A. Brauer in Marburg auf den Seychellen und von Herrn Perrot auf Ste. Marie (Madagaskar) gesammelten Ameisen. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 21: 185-208.
  5. ↑ Molet M, Fisher B, Ito F, Peeters C (2009) Shift from independent to dependent colony foundation and evolution of ‘multi-purpose’ ergatoid queens in Mystrium ants (subfamily Amblyoponinae). Biological journal of the Linnean Society 98: 198-207. doi: 10.1111/j.1095-8312.2009.01257.x
  6. ↑ Bouchet D, Peeters C, Fisher B, Molet M (2013) Both female castes contribute to colony emigration in the polygynous ant Mystrium oberthueri. Ecological Entomology 38: 408-417. doi: 10.1111/een.12033

Images

Figure 1. Colonies of Mystrium. A Mystrium oberthueri B Mystrium voeltzkowi. A a cocoon and a worker B larvae, workers, and ergatoid queens. Photos by Alex Wild.
Figure 1. Colonies of Mystrium. A Mystrium oberthueri B Mystrium voeltzkowi. A a cocoon and a worker B larvae, workers, and ergatoid queens. Photos by Alex Wild. 
Figure 16. Vertex of workers in voeltzkowi species group. A Mystrium oberthueri (CASENT0499569) B Mystrium voeltzkowi (CASENT0002075). A, B head in posterior view.
Figure 16. Vertex of workers in voeltzkowi species group. A Mystrium oberthueri (CASENT0499569) B Mystrium voeltzkowi (CASENT0002075). A, B head in posterior view. 
Figure 17. Differences in worker morphology between Mystrium janovitzi and Mystrium oberthueri. A Mystrium janovitzi (CASENT0006082) B Mystrium oberthueri (CASENT0499569) C Mystrium janovitzi (CASENT0482698: holotype) D Mystrium oberthueri (CASENT0496066). A, B mesosoma in dorsal view C, D clypeus and genal tooth in full-face view.
Figure 17. Differences in worker morphology between Mystrium janovitzi and Mystrium oberthueri. A Mystrium janovitzi (CASENT0006082) B Mystrium oberthueri (CASENT0499569) C Mystrium janovitzi (CASENT0482698: holotype) D Mystrium oberthueri (CASENT0496066). A, B mesosoma in dorsal view C, D clypeus and genal tooth in full-face view. 
Figure 18. Differences in worker morphology between Mystrium oberthueri and Mystrium eques. A Mystrium oberthueri (CASENT0496066) B Mystrium eques (CASENT0418314: holotype) C, E Mystrium oberthueri (CASENT0499569) D, F Mystrium eques (CASENT0317399). A, B head in full-face view C, D pedicel and flagellum in lateral view E, F mandible in dorsal view.
Figure 18. Differences in worker morphology between Mystrium oberthueri and Mystrium eques. A Mystrium oberthueri (CASENT0496066) B Mystrium eques (CASENT0418314: holotype) C, E Mystrium oberthueri (CASENT0499569) D, F Mystrium eques (CASENT0317399). A, B head in full-face view C, D pedicel and flagellum in lateral view E, F mandible in dorsal view. 
Figure 21. Vertex of ergatoid queens in voeltzkowi species group. A Mystrium eques (CASENT0418311) B Mystrium shadow (CASENT0077647: paratype). A, B head in posterior view.
Figure 21. Vertex of ergatoid queens in voeltzkowi species group. A Mystrium eques (CASENT0418311) B Mystrium shadow (CASENT0077647: paratype). A, B head in posterior view. 
Figure 22. Clypeus of ergatoid queens in voeltzkowi species group. A Mystrium eques (CASENT0418317: paratype) B Mystrium oberthueri (CASENT0496064). A, B head in full-face view.
Figure 22. Clypeus of ergatoid queens in voeltzkowi species group. A Mystrium eques (CASENT0418317: paratype) B Mystrium oberthueri (CASENT0496064). A, B head in full-face view. 
Figure 23. Differences in ergatoid queen characters between Mystrium janovitzi and Mystrium oberthueri. A Mystrium janovitzi (CASENT0482696) B Mystrium oberthueri (CASENT0496064) C Mystrium janovitzi (CASENT0003302) D Mystrium oberthueri (CASENT0095743). A, B head in full-face view C, D pedicel and flagellum in lateral view.
Figure 23. Differences in ergatoid queen characters between Mystrium janovitzi and Mystrium oberthueri. A Mystrium janovitzi (CASENT0482696) B Mystrium oberthueri (CASENT0496064) C Mystrium janovitzi (CASENT0003302) D Mystrium oberthueri (CASENT0095743). A, B head in full-face view C, D pedicel and flagellum in lateral view. 
Figure 25. Differences in ergatoid queen characters between Mystrium mirror and Mystrium shadow. A, C Mystrium mirror (CASENT0429899) B, D Mystrium shadow (CASENT0077647: paratype). A, B head and pronotum in oblique dorsal view C, D mesosoma in oblique dorsal view.
Figure 25. Differences in ergatoid queen characters between Mystrium mirror and Mystrium shadow. A, C Mystrium mirror (CASENT0429899) B, D Mystrium shadow (CASENT0077647: paratype). A, B head and pronotum in oblique dorsal view C, D mesosoma in oblique dorsal view. 
Figure 28. Abdomen of Mystrium males. A Mystrium mysticum (CASENT0318027) B Mystrium janovitzi (CASENT0007820) C Mystrium mysticum (CASENT0429983) D Mystrium shadow (CASENT0303561). A, B abdominal segment VIII in oblique lateral view C, D petiole in dorsal view.
Figure 28. Abdomen of Mystrium males. A Mystrium mysticum (CASENT0318027) B Mystrium janovitzi (CASENT0007820) C Mystrium mysticum (CASENT0429983) D Mystrium shadow (CASENT0303561). A, B abdominal segment VIII in oblique lateral view C, D petiole in dorsal view. 
Figure 30. Head of Mystrium males in full-face view. A Mystrium oberthueri (CASENT0179499) B Mystrium shadow (CASENT0107647).
Figure 30. Head of Mystrium males in full-face view. A Mystrium oberthueri (CASENT0179499) B Mystrium shadow (CASENT0107647). 
Figure 37. Males of Mystrium in lateral view. A Mystrium barrybressleri sp. n. (CASENT0135866) B Mystrium mysticum (CASENT0318027) C Mystrium rogeri (CASENT0001083) D Mystrium janovitzi sp. n. (CASENT0007820) E Mystrium mirror sp. n. (CASENT0318941) F Mystrium oberthueri (CASENT0179499) G Mystrium shadow sp. n. (CASENT0107647) H Mystrium voeltzkowi (CASENT0101673).
Figure 38. Head of Mystrium males in full-face view. A Mystrium barrybressleri sp. n. (CASENT0135866) B Mystrium mysticum (CASENT0318027) C Mystrium rogeri (CASENT0001083) D Mystrium janovitzi sp. n. (CASENT0007820) E Mystrium mirror sp. n. (CASENT0318941) F Mystrium oberthueri (CASENT0179499) G Mystrium shadow sp. n. (CASENT0107647) H Mystrium voeltzkowi (CASENT0101673).
Figure 39. Males of Mystrium in dorsal view. A Mystrium barrybressleri sp. n. (CASENT0135866) B Mystrium mysticum (CASENT0318027) C Mystrium rogeri (CASENT0001083) D Mystrium janovitzi sp. n. (CASENT0007820) E Mystrium mirror sp. n. (CASENT0318941) F Mystrium oberthueri (CASENT0179499) G Mystrium shadow sp. n. (CASENT0107647) H Mystrium voeltzkowi (CASENT0101673).
Figure 40. Genital capsule of Mystrium males in lateral view. A Mystrium barrybressleri sp. n. (CASENT0078803) B Mystrium mysticum (CASENT0218102) C Mystrium rogeri (CASENT0001152) D Mystrium janovitzi sp. n. (CASENT0080644) E Mystrium mirror sp. n. (CASENT0317444) F Mystrium oberthueri (CASENT0317422) G Mystrium shadow sp. n. (CASENT0107647) H Mystrium voeltzkowi (CASENT0317583).
Figure 41. Aedeagus of Mystrium males in lateral view. A Mystrium barrybressleri sp. n. (CASENT0078803) B Mystrium mysticum (CASENT0218102) C Mystrium rogeri (CASENT0001152) D Mystrium janovitzi sp. n. (CASENT0080644) E Mystrium mirror sp. n. (CASENT0317444) F Mystrium oberthueri (CASENT0317422) G Mystrium shadow sp. n. (CASENT0107647) H Mystrium voeltzkowi (CASENT0317583).
Figure 50. Workers of voeltzkowi species group in lateral view. A Mystrium eques sp. n. (CASENT0418314: holotype) B Mystrium janovitzi sp. n. (CASENT0482698: holotype) C Mystrium mirror sp. n. (CASENT0429897: holotype) D Mystrium oberthueri (CASENT0496066) E Mystrium shadow sp. n. (CASENT0318933: holotype) F Mystrium voeltzkowi (CASENT0002075).
Figure 50. Workers of voeltzkowi species group in lateral view. A Mystrium eques sp. n. (CASENT0418314: holotype) B Mystrium janovitzi sp. n. (CASENT0482698: holotype) C Mystrium mirror sp. n. (CASENT0429897: holotype) D Mystrium oberthueri (CASENT0496066) E Mystrium shadow sp. n. (CASENT0318933: holotype) F Mystrium voeltzkowi (CASENT0002075). 
Figure 51. Head of the workers of voeltzkowi species group in full-face view. A Mystrium eques sp. n. (CASENT0418314: holotype) B Mystrium janovitzi sp. n. (CASENT0482698: holotype) C Mystrium mirror sp. n. (CASENT0429897: holotype) D Mystrium oberthueri (CASENT0496066) E Mystrium shadow sp. n. (CASENT0318933: holotype) F Mystrium voeltzkowi (CASENT0002075).
Figure 51. Head of the workers of voeltzkowi species group in full-face view. A Mystrium eques sp. n. (CASENT0418314: holotype) B Mystrium janovitzi sp. n. (CASENT0482698: holotype) C Mystrium mirror sp. n. (CASENT0429897: holotype) D Mystrium oberthueri (CASENT0496066) E Mystrium shadow sp. n. (CASENT0318933: holotype) F Mystrium voeltzkowi (CASENT0002075). 
Figure 52. Workers of voeltzkowi species group in dorsal view. A Mystrium eques sp. n. (CASENT0418314: holotype) B Mystrium janovitzi sp. n. (CASENT0482698: holotype) C Mystrium mirror sp. n. (CASENT0429897: holotype) D Mystrium oberthueri (CASENT0496066) E Mystrium shadow sp. n. (CASENT0318933: holotype) F Mystrium voeltzkowi (CASENT0002075).
Figure 52. Workers of voeltzkowi species group in dorsal view. A Mystrium eques sp. n. (CASENT0418314: holotype) B Mystrium janovitzi sp. n. (CASENT0482698: holotype) C Mystrium mirror sp. n. (CASENT0429897: holotype) D Mystrium oberthueri (CASENT0496066) E Mystrium shadow sp. n. (CASENT0318933: holotype) F Mystrium voeltzkowi (CASENT0002075). 
Figure 53. Ergatoid queens of voeltzkowi species group in lateral view. A Mystrium eques sp. n. (CASENT0418317: paratype) B Mystrium janovitzi sp. n. (CASENT0482696) C Mystrium mirror sp. n. (CASENT0318940: paratype) D Mystrium oberthueri (CASENT0496064) E Mystrium shadow sp. n. (CASENT0077647: paratype) F Mystrium voeltzkowi (CASENT0002092).
Figure 53. Ergatoid queens of voeltzkowi species group in lateral view. A Mystrium eques sp. n. (CASENT0418317: paratype) B Mystrium janovitzi sp. n. (CASENT0482696) C Mystrium mirror sp. n. (CASENT0318940: paratype) D Mystrium oberthueri (CASENT0496064) E Mystrium shadow sp. n. (CASENT0077647: paratype) F Mystrium voeltzkowi (CASENT0002092). 
Figure 54. Head of the ergatoid queens of voeltzkowi species group in full-face view. A Mystrium eques sp. n. (CASENT0418317: paratype) B Mystrium janovitzi sp. n. (CASENT0482696) C Mystrium mirror sp. n. (CASENT0318940: paratype) D Mystrium oberthueri (CASENT0496064) E Mystrium shadow sp. n. (CASENT0077647: paratype) F Mystrium voeltzkowi (CASENT0002092).
Figure 54. Head of the ergatoid queens of voeltzkowi species group in full-face view. A Mystrium eques sp. n. (CASENT0418317: paratype) B Mystrium janovitzi sp. n. (CASENT0482696) C Mystrium mirror sp. n. (CASENT0318940: paratype) D Mystrium oberthueri (CASENT0496064) E Mystrium shadow sp. n. (CASENT0077647: paratype) F Mystrium voeltzkowi (CASENT0002092). 
Figure 55. Ergatoid queens of voeltzkowi species group in dorsal view. A Mystrium eques sp. n. (CASENT0418317: paratype) B Mystrium janovitzi sp. n. (CASENT0482696) C Mystrium mirror sp. n. (CASENT0318940: paratype) D Mystrium oberthueri (CASENT0496064) E Mystrium shadow sp. n. (CASENT0077647: paratype) F Mystrium voeltzkowi (CASENT0002092).
Figure 55. Ergatoid queens of voeltzkowi species group in dorsal view. A Mystrium eques sp. n. (CASENT0418317: paratype) B Mystrium janovitzi sp. n. (CASENT0482696) C Mystrium mirror sp. n. (CASENT0318940: paratype) D Mystrium oberthueri (CASENT0496064) E Mystrium shadow sp. n. (CASENT0077647: paratype) F Mystrium voeltzkowi (CASENT0002092). 
Figure 56. Distribution maps for voeltzkowi species group. A Mystrium eques sp. n. B Mystrium janovitzi sp. n. C Mystrium mirror sp. n. D Mystrium oberthueri E Mystrium shadow sp. n. F Mystrium voeltzkowi. Star symbols represent the type locality: the localities for Mystrium oberthueri and Mystrium voeltzkowi are approximate, which are assumed from the original descriptions.
Figure 56. Distribution maps for voeltzkowi species group. A Mystrium eques sp. n. B Mystrium janovitzi sp. n. C Mystrium mirror sp. n. D Mystrium oberthueri E Mystrium shadow sp. n. F Mystrium voeltzkowi. Star symbols represent the type locality: the localities for Mystrium oberthueri and Mystrium voeltzkowi are approximate, which are assumed from the original descriptions. 
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