Difference between revisions of "Temnothorax brackoi"

Latest revision as of 15:38, 1 October 2019

Taxonavigation

Ordo: Hymenoptera
Familia: Formicidae
Genus: Temnothorax

Name

Temnothorax brackoi Salata & Borowiec, 2019 sp. nov. – Wikispecies link – ZooBank link – Pensoft Profile

Type material

Holotype: worker (pin), (CASENT0846669): GREECE, Pel. Messinia | 1.8 km E of Saidona, 985 m | 36.88491N, 22.30419E | 20 VI 2016, L. Borowiec || Collection L. Borowiec | Formicidae | LBC-GR02195 (MNHW).
Paratypes: • 11w. (pin), 9w. (EtOH) (CASENT0846607-CASENT0846617): the same nest sample as holotype (DBET); • 11w. (pin), 5w. (EtOH)(CASENT0846618-CASENT0846628): GREECE, Pel. Messinia | 0.8 km SE of Exochori, 535 m, | [http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=36.89582N%2C+22.27464E&coordinates%5B0%5D=36.89582N%2C+22.27464E 36.89582N, 22.27464E] | 20 VI 2016, L. Borowiec || Collection L. Borowiec | Formicidae | LBC-GR02184 (DBET, MHNG, CASC, BMNH); • 8w. (pin), 1w. (EtOH) (CASENT0846629-CASENT0846636): GREECE, Pel. Messinia | n. Arachova, 860 m | [http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=37.03922N%2C+22.21876E&coordinates%5B0%5D=37.03922N%2C+22.21876E 37.03922N, 22.21876E] | 13 VI 2016, L. Borowiec || Collection L. Borowiec | Formicidae | LBC-GR02029 (DBET); • 2w. (pin) (CASENT0846637-CASENT0846638): GREECE, Pel. Messinia | 1.3 km S of Artemisia, 870 m | [http://ptp.pensoft.eu/redirect_to_googlemap.php?labels%5B0%5D=37.08738N%2C+22.23378E&coordinates%5B0%5D=37.08738N%2C+22.23378E 37.08738N, 22.23378E] | 17 VI 2016, L. Borowiec || Collection L. Borowiec | Formicidae | LBC-GR02121 (DBET).

Other material

GREECE. Epirus, Preveza: • 4w. (pin) (CASENT0846670-CASENT0846673), 21w. (EtOH): 700 m E of Kanali, oak forest, 39.0638N/20.70621E, 50 m, 2016-08-27, leg. L. Borowiec; • 8w. (pin) (CASENT0846674-CASENT0846681), 14w. (EtOH): n. Kanali, oak forest, 39.05458N/20.70207E, 15 m, 2016-08-28, leg. L. Borowiec; • 27w (EtOH): n. Kanali, pine forest, 39.05388N/20.70118E, 20 m, 2016-08-28, leg. L. Borowiec. Ionian Islands, Cephalonia: • 6w. (pin) (CASENT0846682-CASENT0846687): Avithos Lake, area near a small lake in a moist, shaded valley of a small creek, 38.17293N/20.71233E, 278 m, 2014-06-25, leg. L. Borowiec; • 68w. (EtOH): Avithos Lake, shrubs around small lake, 38.17203N/ 20.71107E, 288 m, 2019-06-10, leg. L. Borowiec; • 29w. (EtOH): Katapodata; roadsides with shrubs, 38.23337N/20.64594E, 100 m, 2019-06-10, leg. L. Borowiec; • 4w. (EtOH): 1.5 km NE of Koulourata, mixed forest on shrubs, 38.20667N/ 20.67715E, 273 m, 2019-06-10, leg. L. Borowiec; • 11w. (EtOH): ancient Same; roadsides with shrubs, 38.2522N/20.66423E, 220 m, 2019-06-10, leg. L. Borowiec; • 13w. (EtOH): rd. Skala-Poros; Mediterranean shrubs, 38.12872N/20.79576E, 5 m, 2019-06-12, leg. L. Borowiec. Ionian Islands, Corfu: • 6w. (EtOH): Ag. Ilias, deciduous forest, 39.79367N/19.87508E, 191 m, 2013-06-10, leg. L. Borowiec; • 5w. (pin) (CASENT0846688-CASENT0846692), 5w. (EtOH): Akr. Kefali n. Agios Stefanos, frygana on sea coast, 39.75154N/19.63272E, 13 m, 2013-06-05, leg. L. Borowiec; • 4w. (EtOH): E of Nymfes, old olive tree plantation, 39.75047N/19.8057E, 179 m, 2013-06-09, leg. L. Borowiec; • 8w. (pin) (CASENT0846693-CASENT0846700), 1w. (EtOH): Klimatia, deciduous forest, 39.74123N/19.78953E, 311 m, 2013-06-06, leg. L. Borowiec; • 1w. (pin) (CASENT0846701): N of Ag. Stefanos, old olive forest, 39.76338N/19.65213E, 88 m, 2013-06-05, leg. L. Borowiec; • 4w. (pin) (CASENT0846702-CASENT0846705): n. Doukades, old olive tree plantation, 39.70075N/19.75055E, 174 m, 2013-06-08, leg. L. Borowiec; • 3w. (pin) (CASENT0846706-CASENT0846708), 1w. (EtOH): n. Vistonas, old olive tree plantation, 39.68549N/19.71453E, 422 m, 2013-06-08, leg. L. Borowiec; • 20w. (pin) (CASENT0846709-CASENT0846728), 15w. (EtOH): Nymfes, deciduous forest, 39.75478N/19.79535E, 162 m, 2013-06-06, leg. L. Borowiec; • 20w. (EtOH): Old Perithia, old mountain village, 39.76159N/19.87412E, 467 m, 2013-06-10, leg. L. Borowiec; • 3w. (pin) (CASENT0846729-CASENT0846731): Strinilas, deciduous forest, 39.73963N/19.84265E, 632 m, 2013-06-07, leg. L. Borowiec. Ionian Islands, Lefkada: • 3w. (pin) (CASENT0846732-CASENT0846734), 2w. (EtOH): Asprogerakata, stream valley with deciduous forest, 38.78278N /20.65262E, 430 m, 2016-09-02, leg. L. Borowiec; • 2w. (pin) (CASENT0846735-CASENT0846736), 8w. (EtOH): Egklouvi, stream valley with deciduous forest, 38.73212N/20.64821E, 690 m, 2016-09-02, leg. L. Borowiec. Ionian Islands, Zakynthos: • 4w. (EtOH): 1 km E of Elies, shrubs along roadsides, 37.90173N/20.68787E, 300 m, 2018-05-06, leg. L. Borowiec; • 18w. (EtOH): 1 km N of Exo Chora, mixed forest, 37.81063N/20.68459E, 430 m, 2018-05-08, leg. L. Borowiec; • 1w. (pin) (CASENT0846737): 1.2 km N of Ano Vasilikos, roadsides along olive plantation and pasture with oak shrubs, 37.72456N/20.97786E, 30 m, 2018-05-05, leg. L. Borowiec; • 128w. (EtOH): 1.2 km N of Vasilikos, roadsides along olive plantation and pasture with oak shrubs, 37.72456N/20.97786E, 30 m, 2018-05-05, leg. L. Borowiec; • 2w. (EtOH): 1.2 km NE of Anafonitria shrubs around burned forests, 37.85489N/20.64124E, 475 m, 2018-05-10, leg. L. Borowiec; • 1w. (pin) (CASENT0846738), 43w. (EtOH): 1.2 km SW of Ano Vasilikos loc. 1, roadsides along olive plantation, 37.72374N/20.95964E, 72 m, 2018-05-05, leg. L. Borowiec; • 1w. (pin) (CASENT0846739), 25w. (EtOH): 1.2 km SW of Ano Vasilikos loc. 2, roadsides along olive plantation, 37.72325N/20.96123E, 75 m, 2018-05-05, leg. L. Borowiec; • 1w. (pin): 1.2 km SW of Skinaria, limestone hills after burned forests, 37.87694N/20.69272E, 375 m, 2018-05-06, leg. L. Borowiec; • 1w. (pin) (CASENT0846740), 13w. (EtOH): 1.4 km E of Ano Volimes, mixed forest, 37.87594N/20.68525E, 430 m, 2018-05-06, leg. L. Borowiec; • 1w. (EtOH): 1.5 km N of Exo Chora: shrubs around meadow, 37.81446N/ 20.68826E, 460 m, 2018-05-08, leg. L. Borowiec; • 38w. (EtOH): 1.7 km NE of Ano Volimes, shrubs around pastures, 37.88627N/20.68456E, 445 m, 2018-05-06, leg. L. Borowiec; • 3w. (EtOH): 1.8 km SW of Volimes, shrubs along roadsides, 37.86472N/20.64234E, 350 m, 2018-05-10, leg. L. Borowiec; • 1w. (EtOH): 1.9 km W of Maries, roadsides in burned forests, 37.818N/20.65556E, 290 m, 2018-05-09, leg. L. Borowiec; • 1w. (pin), 30w. (EtOH): 2.5 km NE of Maries, mixed forest, 37.8299N/20.70151E, 475 m, 2018-05-08, leg. L. Borowiec; • 51w. (EtOH): 3.9 km NE of Maries, shrubs around olive plantation, 37.84202N/20.70938E, 370 m, 2018-05-08, leg. L. Borowiec; • 1w. (pin) (CASENT0846741), 16w. (EtOH): 470 m NE of Orthonies, shrubs along roadsides, 37.85435N/20.69843E, 405 m, 2018-05-10, leg. L. Borowiec; • 1w. (EtOH): 500 m S of Apelati, pine forest, 37.66873N/20.81407E, 160 m, 2018-05-07, leg. L. Borowiec; • 1w. (EtOH): 700 m SW of Koroni, maquis, 37.86582N/20.71753E, 290 m, 2018-05-10, leg. L. Borowiec; • 3w. (EtOH): 750 m S of Volimes, shrubs in cypress forest, 37.86762N/20.66191E, 360 m, 2018-05-10, leg. L. Borowiec; • 1w. (pin) (CASENT0846742), 81w. (EtOH): 800 m SE of Xirokastello, roadsides along olive plantation, 37.73491N/20.95139E, 75 m, 2018-05-05, leg. L. Borowiec; • 10w. (EtOH): 880 m S of Orthonies, shrubs in cypress forest, 37.84462N/20.69843E, 390 m, 2018-05-10, leg. L. Borowiec; • 66w. (EtOH): Ag. Georgiou monastery, shrubs along roadsides, 37.85971N/20.63646E, 330 m, 2018-05-10, leg. L. Borowiec; • 8w. (EtOH): Ag. Joannis, roadsides with shrubs, 37.72924N/20.94553E, 165 m, 2018-05-05, leg. L. Borowiec; • 6w. (EtOH): Argassi, urban area, 37.76182N/20.92704E, 10 m, 2018-05-04, leg. L. Borowiec; • 1w. (EtOH): Livia Mts. loc. 1, shrubs along roadsides, 37.83018N/20.71619E, 600 m, 2018-05-08, leg. L. Borowiec; • 15w. (EtOH): Vrachionas Mts., mountain pastures with shrubs, 37.81798N/ 20.70621E, 670 m, 2018-05-08, leg. L. Borowiec; • 1w. (EtOH): W of Kampi, pine forest, 37.78161N/20.68078E, 165 m, 2018-05-09, leg. L. Borowiec. Macedonia, Pieria: • 4w. (EtOH): Platamonas Castle hill, on stones, soil and herbs, 40.00868N/ 22.59654E, 12 m, 2019-05-11, leg. L. Borowiec; • 1w. (pin) (CASENT0846743): 2 km W of Panteleimonas, roadsides with shrubs, 39.98563N/ 22.59513E, 305 m, 2019-05-15, leg. L. Borowiec; • 20w. (EtOH): road to P. Poroi loc. 1, roadsides with shrubs, 39.97963N/ 22.61563E, 110 m, 2019-05-17, leg. L. Borowiec; • 7w. (EtOH): road to P. Poroi loc. 2, roadsides with shrubs, 39.97627N/ 22.61146E, 185 m, 2019-05-17, leg. L. Borowiec; •81w. (EtOH): road to P. Poroi loc. 3, roadsides with shrubs, 39. 96863N/22.60494E, 260 m, 2019-05-17, leg. L. Borowiec. Peloponnese, Korinthia: • 16w. (pin) (CASENT0846744-CASENT0846759): n. Evrostina, deciduous forest, 38.07386N/22.39388E, 662 m, 2013-09-01, leg. L. Borowiec. Peloponnese, Laconia: • 13w. (pin) (CASENT0846760-CASENT0846772), 2w. (EtOH): Taygetos Mts., 1.5 km SW of Anavryti, coniferous forest, 37.0191N/22.36003E, 990 m, 2016-06-21, leg. L. Borowiec. Peloponnese, Messinia: • 1w. (pin) (CASENT0846773): Taygetos Mts., 0.7 km S of Dyrrachio, coniferous forest, 37.1811N/22.2074E, 800 m, 2016-06-22, leg. L. Borowiec; • 5w. (pin) (CASENT0846774-CASENT0846778): Taygetos Mts., 2 km W of Arachova, stream valley with Platanus forest, 37.0357N/22.1978E, 680 m, 2016-06-13, leg. L. Borowiec; • 1w. (pin) (CASENT0846779): Taygetos Mts., Chora Getson, stream valley with deciduous forest, 36.94779N/22.25466E, 615 m, 2016-06-14, leg. L. Borowiec; • 4w. (pin) (CASENT0846780-CASENT0846783): Taygetos Mts., Karveli, rest area with stream, 37.07591N/22.20633E, 600 m, 2016-06-17, leg. L. Borowiec; • 2w. (pin) (CASENT0846784-CASENT0846785): Tetrazi Mts., 0.5 km E of Vastas, stream valley with deciduous forest, 37.36441N/21.99304E, 895 m, 2016-06-19, leg. L. Borowiec; • 1w. (pin) (CASENT0846786), 2w. (EtOH): Tetrazi Mts., Isaris, pine forest, 37.36644N/22.01516E, 790 m, 2016-06-19, leg. L. Borowiec; • 1w. (pin) (CASENT0846787): Tetrazi Mts., Karnasi, stream valley with Platanus forest, 37.31904N/21.99158E, 460 m, 2016-06-19, leg. L. Borowiec. Sterea Ellas, Aetolia-Acarnania: • 1w. (EtOH): Psila Alonia, stream valley with Platanus forest, 38.96255N/21.19527E, 62 m, 2016-09-04, leg. L. Borowiec. Sterea Ellas, Euboea: • 2w. (EtOH): 1.2 km NW of Gerontas, mediterranean shrubs along roadsides, 38.45885N/23.808E, 405 m, 2018-06-10, leg. L. Borowiec; • 2w. (EtOH): 2.3 km S of Stropones, mixed forest, 38.9933N/23.87807E, 860 m, 2018-06-10, leg. L. Borowiec; • 1w. (EtOH): 300 m NW of Agios, pine forest with mediterranean shrubs, 38.65856N/23.55525E, 600 m, 2018-06-11, leg. L. Borowiec; • 7w. (EtOH): 570 m NW of Drosia, stream valley with mixed forest, 38.61705N/23.59089E, 140 m, 2018-06-11, leg. L. Borowiec. Thessaly, Larissa: • 1w. (pin) (CASENT0846788): Mt. Olympus, 5.3 km E of Olympiada, alpine pastures with shrubs, 40.00989N/22.31096E, 800 m, 2017-05-09, leg. L. Borowiec; • 7w. (EtOH): Mt. Ossa, 2.4 km SE of Karitsa, stream valley in deciduous forest, 39.82632N/22.77557E, 425 m, 2017-05-04, leg. L. Borowiec; • 12w. (EtOH): Mt. Ossa, 600 m SE of Karitsa, stream valley in deciduous forest, 39.84009N/22.76983E, 300 m, 2017-05-04, leg. L. Borowiec; • 3w. (pin) (CASENT0846789-CASENT0846791), 44w. (EtOH): Mt. Ossa, Kokkino Nero, stream valley in urban area, 39.83389N/22.79379E, 3 m, 2017-05-10, leg. L. Borowiec. CROATIA, Podgorje: • 4w. (pin) (CASENT0846792-CASENT0846795), vicinity Sibenj, 9 km S Senj, 0–50 m, 3.6.1997, A. Schulz & K. Vock leg.

Locus typicus

Peloponnese, Taygetos Mts.

Differential diagnosis

Temnothorax brackoi is most similar to T. aveli (Bondroit, 1918), a species widely distributed in the western part of the Mediterranean basin, east to Slovenia. It is similar to T. aveli especially in body shape, petiolar structure and unicolour yellow antennae. Temnothorax aveli differs in microreticulation of head not as regular as in T. brackoi, with slightly shiny surface between sculpture and often with presence of median stripe of diffused microsculpture on frons, while in T. brackoi head sculpture is always homogenous, without areas of diffused microreticulation and dull surface between sculpture. From Greek members of Temnothorax, T. brackoi is most similar to sympatric T. graecus, but differs from it in well-developed sculpture on the whole head surface (in T. graecus central part of frons is more or less shiny, without microreticulation) and larger, distinctly triangular propodeal spines (in T. graecus spines are short, needle-shaped); some specimens of T. brackoi can also be confused with species belonging to the T. tuberum group, but T. brackoi differs from them in low and arched petiolar node and lack of longitudinal striation on frons.

Description

Worker (n = 10): HL: 0.637 ± 0.04 (0.584–0.683); HW: 0.547 ± 0.03 (0.509–0.696); SL: 0.437 ± 0.02 (0.410–0.560); EL: 0.148 ± 0.01 (0.124–0.174); EW: 0.105 ± 0.01 (0.087–0.124); WL: 0.735 ± 0.05 (0.671–0.820); PSL: 0.155 ± 0.015 (0.130–0.174); SDL: 0.111 ± 0.01 (0.096–0.124); PEL: 0.271 ± 0.03 (0.224–0.311); PPL: 0.168 ± 0.01 (0.149–0.186); PEH: 0.208 ± 0.015 (0.186–0.236); PPH: 0.207 ± 0.015 (0.180–0.224); PNW: 0.368 ± 0.02 (0.329–0.401); PLW: 0.170 ± 0.01 (0.149–0.199); PPW: 0.223 ± 0.02 (0.196–0.242); CI: 85.9 ± 1.2 (83.1–87.5); SI1: 68.6 ± 1.6 (66.7–71.3); SI2: 79.9 ± 1.6 (77.1–81.9); MI: 50.0 ± 1.0 (48.4–51.3); EI1: 71.0 ± 5.0 (64.6–78.3); EI2: 16.5 ± 1.4 (14.8–18.75); PI: 130.0 ± 7.9 (120.0–145.5); PPI: 81.4 ± 6.1 (128.2–150.0); PSI: 135.8 ± 12.4 (120.0–155.6).

Colour

Head, antennae, mesosoma, petiole, postpetiole and legs uniformly yellow to dark yellow, in few specimens, frons and femora partly darkened. Gaster yellow, only the first gaster tergite with wide, dark band on its posterior part (Figs 1, 2). Head. Rectangular, 1.16 times as long as wide, lateral surfaces below and above eyes gently convex, posterior edges convex, occipital margin of head straight or slightly concave (Figs 9, 13). Anterior margin of clypeus slightly convex, medial notch absent. Eyes moderate, oval, 1.4 times as long as wide. Antennal scape short, in lateral view slightly curved, 0.69 times as long as length of the head, in apex gradually widened, its base with small, triangular tooth, funiculus long, club 3-segmented (Fig. 9). Surface of scape with very fine microreticulation, shiny, covered with thin, moderate dense, decumbent to suberect setae. Mandibles rounded with thick sparse, longitudinal striae, shiny. Clypeus shiny with thick, sparse, longitudinal striae, area between striae smooth and shiny. Frontal carinae short, not extending beyond frontal lobes. Antennal fossa deep, with sparse, thin, roundly curved striae and dense reticulation. Frontal lobes narrow, smooth with slight, dense longitudinal striation (Fig. 13). Frons, vertex and temples with dense, thick, reticulation, sometimes central surface of frons with additional a few thin, longitudinal, interrupted wrinkles, surface between striation smooth; malar area with irregular, thick, reticulation, space between reticulation smooth or with very sparse microreticulation, shiny; genae with sparser, thank of frons, and thick reticulation, shiny. Frons and vertex with erect, pale, short and thick setae (Fig. 13). Mesosoma. Elongate, approximately twice as long as wide, slightly arched in profile. Metanotal groove absent. Pronotum convex on sides. Propodeal spines short to moderate, triangular, directed upward, base wide, tips sharp (Fig. 2). Whole surface with dense, reticulation, its dorsal surface and lateral surfaces of pronotum with thick and sparse longitudinal striation or reticulation. Area between thick sculpture shiny, smooth or sometimes with sparse, fait microreticulation. Entire mesosoma bearing erect, pale, short and thick setae (Fig. 2). Petiole. In lateral view, with short peduncle, node low, with anterior face flat, and posterior face convex and dorsum convex. Peduncle and petiolar node shiny, with thick, dense reticulation, area between rugae smooth, dorsum with sparser reticulation. Dorsal surface bearing sparse, short, erect setae (Fig. 2). Postpetiole. In lateral view, regularly convex, apical half with gently convex sides (Figs 1, 2), on the whole surface shiny, with thick, dense reticulation, dorsum with sparser reticulation; area between rugae smooth. Dorsal surface bearing sparse, short, semierect to erect setae (Fig. 2). Gaster. Gaster smooth and shiny, bearing erect, thin, pale setae (Figs 1, 2).

Etymology

Named after Gregor Bračko, a Slovenian myrmecologist, for his significant contributions to the studies on ants of the Balkan Peninsula.

General distribution

Greece: Epirus, Ionian Islands, Peloponnese, western Sterea Ellas, Thessaly; Croatia: Dalmatia.

Biology

Specimens of T. brackoi were collected from various localities: sunny and shadowy located on lowlands and highlands (9 - 990 m a.s.l). The species was noted in various habitats, most often on shrubs and trees growing along roadsides and olive plantations, maquis, phrygana, often inside forests (mostly deciduous, occasionally coniferous). We noted also its presence on the leaves of climbing plants growing on walls or trees. Nests were not found, probably like other species of this group, are located inside dry stems of herbs.
The following ant species were recorded in the same areas as T. brackoi:
Epirus, Preveza, 700 m E of Kanali: Aphaenogaster balcanica (Emery), A. subterranea (Latreille), Camponotus dalmaticus (Nylander), C. fallax (Nylander), C. lateralis (Olivier), C. piceus (Leach), C. vagus (Scopoli), Cataglyphis nodus (Brullé), Colobopsis truncata (Spinola), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Lasius lasioides (Emery), Messor wasmanni Krausse, Myrmecina graminicola (Latreille), Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Temnothorax bulgaricus (Forel), T. rogeri Emery; near Kanali, oak forest: Aphaenogaster balcanica (Emery), A. subterranea (Latreille), Camponotus dalmaticus (Nylander), C. fallax (Nylander), C. lateralis (Olivier), Cataglyphis nodus (Brullé), Colobopsis truncata (Spinola), Crematogaster schmidti (Mayr), Lasius lasioides (Emery), Pheidole pallidula (Nylander); near Kanali, pine forest: Temnothorax bulgaricus (Forel).
Ionian Islands, Cephalonia, Avithos Lake: Aphaenogaster muelleriana Wolf, Camponotus dalmaticus (Nylander), C. gestroi Emery, C. ionius Emery, C. lateralis (Olivier), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Lepisiota frauenfeldi (Mayr), Liometopum microcephalum (Panzer), Messor wasmanni Krausse, Monomorium monomorium Bolton, Myrmecina graminicola (Latreille), Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Tapinoma festae Emery, Temnothorax bulgaricus (Forel), T. exilis (Emery), T. laconicus Csősz et al., Tetramorium kephalosi Salata & Borowiec; Katapodata: Camponotus dalmaticus (Nylander), Crematogaster schmidti (Mayr), Lepisiota frauenfeldi (Mayr), Plagiolepis pygmaea (Latreille); 1.5 km NE of Koulourata: Aphaenogaster balcanica (Emery), Camponotus dalmaticus (Nylander), C. kiesenwetteri (Roger), C. lateralis (Olivier), Crematogaster schmidti (Mayr), Lepisiota frauenfeldi (Mayr), Pheidole balcanica Seifert, Plagiolepis pygmaea (Latreille), Temnothorax bulgaricus (Forel), T. leviceps (Emery). T. messiniaensis sp. nov.; ancient Same: Camponotus kiesenwetteri (Roger), C. lateralis (Olivier), Colobopsis truncata (Spinola), Crematogaster ionia Forel, C. sordidula (Nylander), Pheidole balcanica Seifert, Temnothorax bulgaricus (Forel), T. messiniaensis sp. nov.; road Skala-Poros: Camponotus aethiops (Latreille), C. gestroi Emery; C. dalmaticus (Nylander), C. kiesenwetteri (Roger), C. lateralis (Olivier), Colobopsis truncata (Spinola), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Temnothorax messiniaensis sp. nov.
Ionian Islands, Corfu, Ag. Ilias: Aphaenogaster subterranea (Latreille), Camponotus dalmaticus (Nylander), C. lateralis (Olivier), Temnothorax laconicus Csősz et al.; Akr. Kefali near Agios Stefanos: Aphaenogaster balcanica (Emery), Bothriomyrmex communista Santschi, Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. lateralis (Olivier), C. piceus (Leach), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Lasius illyricus Zimmermann, Lepisiota melas (Emery), L. nigra (Dalla Torre), Messor wasmanni Krausse, Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Prenolepis nitens (Mayr), Temnothorax laconicus Csősz et al.; E of Nymfes: Aphaenogaster muelleriana Wolf, A. subterranea (Latreille), Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. lateralis (Olivier), Crematogaster sordidula (Nylander), Dolichoderus quadripunctatus (Linnaeus), Formica gagates Latreille, Lasius bombycina Seifert & Galkowski, L. illyricus Zimmermann, Lepisiota frauenfeldi (Mayr), Temnothorax laconicus Csősz et al.; Klimatia: Aphaenogaster balcanica (Emery), A. muelleriana Wolf, A. subterranea (Latreille), Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. gestroi Emery, C. lateralis (Olivier), C. heidrunvogtae Seifert, C. piceus (Leach), Cataglyphis nodus (Brullé), Colobopsis truncata (Spinola), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Dolichoderus quadripunctatus (Linnaeus), Formica gagates Latreille, Lasius bombycina Seifert & Galkowski, L. illyricus Zimmermann, L. lasioides (Emery), Lepisiota frauenfeldi (Mayr), L. nigra (Dalla Torre), Messor ibericus Santschi, M. wasmanni Krausse, Myrmecina graminicola (Latreille), Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), P. xene Stärcke, Prenolepis nitens (Mayr), Temnothorax cf. exilis, T. laconicus Csősz et al., T. rogeri Emery, Tetramorium kephalosi Salata & Borowiec.; N of Ag. Stefanos: Aphaenogaster balcanica (Emery), Camponotus aethiops (Latreille), C. fallax (Nylander), C. lateralis (Olivier), C. piceus (Leach), Crematogaster schmidti (Mayr), Lasius bombycina Seifert & Galkowski, L. illyricus Zimmermann, Lepisiota frauenfeldi (Mayr), Messor ibericus Santschi, Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Solenopsis cf. fugax, Temnothorax clypeatus (Mayr); near Doukades: Camponotus lateralis (Olivier), C. piceus (Leach), Lasius lasioides (Emery); near Vistonas: Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. piceus (Leach), Crematogaster sordidula (Nylander), Lepisiota melas (Emery), Messor ibericus Santschi, M. wasmanni Krausse, Plagiolepis pygmaea (Latreille), Temnothorax cf. exilis, T. rogeri Emery, Tetramorium kephalosi Salata & Borowiec; Nymfes: Aphaenogaster cf. subterranea, Temnothorax laconicus Csősz et al., T. cf. tuberum, Tetramorium cf. caespitum; Corfu, Old Perithia: Aphaenogaster balcanica (Emery), A. epirotes (Emery), A. muelleriana Wolf, Bothriomyrmex communista Santschi, Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. gestroi Emery, C. lateralis (Olivier), C. piceus (Leach), Crematogaster schmidti (Mayr), Lasius illyricus Zimmermann, L. lasioides (Emery), Lepisiota frauenfeldi (Mayr), L. melas (Emery), Messor ibericus Santschi, M. wasmanni Krausse, Plagiolepis pygmaea (Latreille), Prenolepis nitens (Mayr), Tapinoma erraticum (Latreille), Temnothorax affinis (Mayr), T. cf. exilis, T. laconicus Csősz et al., T. cf. nigriceps, T. rogeri Emery, Tetramorium cf. caespitum, T. kephalosi Salata & Borowiec; Strinilas: Aphaenogaster balcanica (Emery), A. epirotes (Emery), Bothriomyrmex communista Santschi, Camponotus dalmaticus (Nylander), C. lateralis (Olivier), Formica gagates Latreille, Lasius lasioides (Emery), Plagiolepis perperamus Salata et al., P. pygmaea (Latreille), Ponera coarctata (Latreille), Solenopsis cf. fugax, Tapinoma erraticum (Latreille), T. laconicus Csősz et al., Tetramorium kephalosi Salata & Borowiec, T. cf. punctatum;
Ionian Islands, Lefkada, Asprogerakata: Aphaenogaster balcanica (Emery), A. muelleriana Wolf, Camponotus aethiops (Latreille), C. dalmaticus (Nylander), Crematogaster schmidti (Mayr), Lepisiota frauenfeldi (Mayr), Pheidole cf. pallidula, Prenolepis nitens (Mayr), Tetramorium kephalosi Salata & Borowiec; Egklouvi: Aphaenogaster balcanica (Emery), Camponotus dalmaticus (Nylander), Cataglyphis nodus (Brullé), Crematogaster schmidti (Mayr), Messor wasmanni Krausse, Pheidole cf. pallidula, Temnothorax lichtensteini (Bondroit), T. rogeri Emery;
Ionian Islands, Zakynthos, 1 km E of Elies: Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. kiesenwetteri (Roger), C. lateralis (Olivier), C. oertzeni Forel, Crematogaster schmidti (Mayr), Messor wasmanni Krausse, Plagiolepis pygmaea (Latreille), Temnothorax cf. tergestinus; 1 km N of Exo Chora: Aphaenogaster balcanica (Emery), Camponotus dalmaticus (Nylander), C. kiesenwetteri (Roger), C. oertzeni Forel, Crematogaster schmidti (Mayr), C. sordidula (Nylander), Lepisiota frauenfeldi (Mayr), Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Temnothorax bulgaricus (Forel), T. rogeri Emery, T. cf. tergestinus; 1.2 km N of Ano Vasilikos: Temnothorax exilis (Emery); 1.2 km N of Vasilikos: Aphaenogaster balcanica (Emery), Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. kiesenwetteri (Roger), C. lateralis (Olivier), Colobopsis truncata (Spinola), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Lepisiota frauenfeldi (Mayr), Plagiolepis pygmaea (Latreille), Temnothorax exilis (Emery), T. graecus (Forel), T. rogeri Emery, T. cf. tergestinus; 1.2 km NE of Anafonitria: Aphaenogaster balcanica (Emery), A. cf. epirotes, Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. kiesenwetteri (Roger), Crematogaster schmidti (Mayr), Lepisiota frauenfeldi (Mayr), Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Temnothorax exilis (Emery), T. muellerianus (Finzi), T. cf. tergestinus; 1.2 km SW of Ano Vasilikos loc. 1: Aphaenogaster balcanica (Emery), Camponotus dalmaticus (Nylander), C. kiesenwetteri (Roger), C. lateralis (Olivier), C. oertzeni Forel, Crematogaster schmidti (Mayr), C. sordidula (Nylander), Lepisiota frauenfeldi (Mayr), Plagiolepis pygmaea (Latreille), Temnothorax exilis (Emery); 1.2 km SW of Ano Vasilikos loc. 2: Aphaenogaster balcanica (Emery), Aphaenogaster cf. epirotes, Camponotus dalmaticus (Nylander), C. lateralis (Olivier), C. oertzeni Forel, Cataglyphis nodus (Brullé), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Lepisiota melas (Emery), Messor wasmanni Krausse, Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Temnothorax exilis (Emery); 1.2 km SW of Skinaria: Aphaenogaster balcanica (Emery), Camponotus aethiops (Latreille), C. gestroi Emery, C. ionius Emery, C. kiesenwetteri (Roger), C. oertzeni Forel, Crematogaster sordidula (Nylander), Lepisiota melas (Emery), Messor wasmanni Krausse, Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Temnothorax exilis (Emery), T. rogeri Emery, T. cf. tergestinus, Tetramorium diomedeum Emery, T. kephalosi Salata & Borowiec; 1.4 km E of Ano Volimes: Aphaenogaster balcanica (Emery), A. cf. epirotes, A. muelleriana Wolf, Camponotus dalmaticus (Nylander), Crematogaster schmidti (Mayr), Lepisiota frauenfeldi (Mayr), Messor wasmanni Krausse, Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Prenolepis nitens (Mayr), Temnothorax bulgaricus (Forel), T. cf. tergestinus, Tetramorium kephalosi Salata & Borowiec; 1.5 km N of Exo Chora: Aphaenogaster cf. epirotes, Camponotus dalmaticus (Nylander), C. kiesenwetteri (Roger), C. lateralis (Olivier), C. oertzeni Forel, Crematogaster schmidti (Mayr), Lepisiota frauenfeldi (Mayr), Messor wasmanni Krausse, Plagiolepis pygmaea (Latreille), Temnothorax bulgaricus (Forel), T. cf. tergestinus, Tetramorium kephalosi Salata & Borowiec; 1.7 km NE of Ano Volimes: Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. gestroi Emery, C. kiesenwetteri (Roger), C. lateralis (Olivier), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Pheidole cf. pallidula, Temnothorax cf. tergestinus; 1.8 km SW of Volimes: Aphaenogaster balcanica (Emery), Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. kiesenwetteri (Roger), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Plagiolepis pygmaea (Latreille), Temnothorax cf. tergestinus; 1.9 km W of Maries: Aphaenogaster balcanica (Emery), A. cf. epirotes, Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. kiesenwetteri (Roger), Crematogaster sordidula (Nylander), Lepisiota melas (Emery), Messor wasmanni Krausse, Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Temnothorax cf. tergestinus; 2.5 km NE of Maries: Aphaenogaster balcanica (Emery), A. cf. epirotes, A. subterraneoides Emery, Bothriomyrmex communista Santschi, Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. kiesenwetteri (Roger), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Plagiolepis pygmaea (Latreille), Temnothorax rogeri Emery, Tetramorium kephalosi Salata & Borowiec; 3.9 km NE of Maries: Camponotus aethiops (Latreille), C. dalmaticus (Nylander), Crematogaster schmidti (Mayr), Prenolepis nitens (Mayr), Temnothorax cf. tergestinus, Tetramorium diomedeum Mayr; 470 m NE of Orthonies: Aphaenogaster balcanica (Emery), Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. kiesenwetteri (Roger), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Lepisiota melas (Emery), Plagiolepis pygmaea (Latreille), Temnothorax bulgaricus (Forel), T. exilis (Emery), Temnothorax cf. tergestinus; 500 m S of Apelati: Aphaenogaster balcanica (Emery), Camponotus dalmaticus (Nylander), C. lateralis (Olivier), Crematogaster ionia Forel, C. schmidti (Mayr), Messor wasmanni Krausse, Myrmecina graminicola (Latreille), Plagiolepis pygmaea (Latreille), Temnothorax cf. tergestinus; 700 m SW of Koroni: Camponotus aethiops (Latreille), C. kiesenwetteri (Roger), C. lateralis (Olivier), Lepisiota melas (Emery), Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Temnothorax exilis (Emery), Temnothorax cf. tergestinus; 750 m S of Volimes: Camponotus dalmaticus (Nylander), C. gestroi Emery, C. lateralis (Olivier), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Messor wasmanni Krausse, Plagiolepis pygmaea (Latreille), Temnothorax bulgaricus (Forel), T. rogeri Emery, T. cf. tergestinus; 800 m SE of Xirokastello: Camponotus dalmaticus (Nylander), C. kiesenwetteri (Roger), C. lateralis (Olivier), C. oertzeni Forel, Colobopsis truncata (Spinola), Crematogaster schmidti (Mayr), Lepisiota melas (Emery), Messor wasmanni Krausse, Plagiolepis pygmaea (Latreille), Temnothorax exilis (Emery), T. cf. tergestinus, Tetramorium cf. punctatum; 880 m S of Orthonies: Aphaenogaster balcanica (Emery), A. cf. epirotes, Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. kiesenwetteri (Roger), C. lateralis (Olivier), Crematogaster schmidti (Mayr), Lepisiota frauenfeldi (Mayr), L. melas (Emery), Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Temnothorax bulgaricus (Forel), T. rogeri Emery, T. cf. tergestinus; Ag. Georgiou monastery: Camponotus aethiops (Latreille), C. dalmaticus (Nylander), Crematogaster schmidti (Mayr), Plagiolepis pygmaea (Latreille); Ag. Joannis: Camponotus aethiops (Latreille), C. kiesenwetteri (Roger), C. lateralis (Olivier), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Pheidole cf. pallidula, Solenopsis cf. lusitanica, Temnothorax graecus (Forel), T. rogeri Emery, T. cf. tergestinus, Tetramorium kephalosi Salata & Borowiec; Argassi: Aphaenogaster balcanica (Emery), Camponotus dalmaticus (Nylander), C. kiesenwetteri (Roger), C. oertzeni Forel, Colobopsis truncata (Spinola), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Lasius alienus (Foerster), Lepisiota melas (Emery), Messor ibericus Santschi, Monomorium monomorium Bolton, Nylanderia jaegerskioeldi (Mayr), Pheidole cf. pallidula, Plagiolepis perperamus Salata et al., P. pygmaea (Latreille), Tapinoma erraticum (Latreille), Temnothorax rogeri Emery, Tetramorium immigrans Santschi, T. cf. punctatum; Livia Mts. loc. 1: Aphaenogaster balcanica (Emery), A. cf. epirotes, Bothriomyrmex communista Santschi, Camponotus aethiops (Latreille), C. kiesenwetteri (Roger), Crematogaster schmidti (Mayr), C. sordidula (Nylander), Messor wasmanni Krausse, Prenolepis nitens (Mayr), Tetramorium cf. punctatum; Vrachionas Mts.: Aphaenogaster balcanica (Emery), Bothriomyrmex communista Santschi, Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. kiesenwetteri (Roger), C. oertzeni Forel, Crematogaster schmidti (Mayr), Messor wasmanni Krausse, Plagiolepis pygmaea (Latreille), Prenolepis nitens (Mayr), Temnothorax cf. tergestinus, Tetramorium cf. punctatum; s, W of Kampi: Aphaenogaster balcanica (Emery), A. cf. epirotes, A. muelleriana Wolf, Camponotus aethiops (Latreille), Crematogaster schmidti (Mayr), Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Solenopsis cf. lusitanica, Temnothorax rogeri Emery.
Macedonia, Pieria, Platamonas Castle hill: Aphaenogaster epirotes (Emery), A. muelleriana Wolf, A. cf. subterranea, Camponotus dalmaticus (Nylander), C. lateralis (Olivier), C. piceus (Leach), Crematogaster ionia Forel, C. sordidula (Nylander), Lasius emarginatus (Olivier), L. turcicus (Santschi), Messor hellenius Agosti & Collingwood, Pheidole balcanica Seifert, Plagiolepis pygmaea (Latreille), Prenolepis nitens (Mayr), Temnothorax lichtensteini (Bondroit); road to P. Poroi loc. 1: Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. ionius Emery, C. piceus (Leach), Chalepoxenus muellerianus (Finzi), Crematogaster ionia Forel, Dolichoderus quadripunctatus (Linnaeus), Formica gagates Latreille, Lasius emarginatus (Olivier), Messor hellenius Agosti & Collingwood, Plagiolepis pygmaea (Latreille), Tapinoma erraticum (Latreille), Temnothorax cf. exilis, T. graecus (Forel), T. turcicus (Santschi); road to P. Poroi loc. 2: Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. piceus (Leach), Cataglyphis nodus (Brullé), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus); Lasius emarginatus (Olivier), Lepisiota frauenfeldi (Mayr), Pheidole balcanica Seifert, Plagiolepis pygmaea (Latreille), Prenolepis nitens (Mayr), Tapinoma erraticum (Latreille), Temnothorax graecus (Forel), T. turcicus (Santschi), Tetramorium kephalosi Salata & Borowiec; road to P. Poroi loc. 3: Camponotus dalmaticus (Nylander), C. lateralis (Olivier), C. vagus (Scopoli), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Lasius emarginatus (Olivier), Messor hellenius Agosti & Collingwood, Pheidole balcanica Seifert, Plagiolepis pygmaea (Latreille), Prenolepis nitens (Mayr), Temnothorax bulgaricus (Forel), T. morea Csősz, Salata & Borowiec, T. turcicus (Santschi).
Peloponnese, Korinthia, near Evrostina: Aphaenogaster subterranea (Latreille), Camponotus lateralis (Olivier), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Lasius emarginatus (Olivier), L. turcicus (Santschi), Ponera coarctata (Latreille), Temnothorax laconicus Csősz et al.
Peloponnese, Laconia, Taygetos Mts., 1.5 km SW of Anavryti: Aphaenogaster balcanica (Emery), A. subterranea (Latreille), Camponotus dalmaticus (Nylander), C. oertzeni Forel, C. piceus (Leach), Cataglyphis nodus (Brullé), Crematogaster ionia Forel, Temnothorax laconicus Csősz et al.
Peloponnese, Messinia, Taygetos Mts., 0.7 km S of Dyrrachio: Camponotus lateralis (Olivier), Crematogaster schmidti (Mayr), Lasius flavus (Fabricius), L. illyricus Zimmermann, Temnothorax helenae Csősz et al.; Taygetos Mts., 0.8 km SE of Exochori: Aphaenogaster balcanica (Emery), Camponotus dalmaticus (Nylander), C. lateralis (Olivier), Crematogaster ionia Forel, Lepisiota frauenfeldi (Mayr), Messor structor (Latreille), M. wasmanni Krausse, Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Temnothorax cf. bulgaricus, T. exilis (Emery), T. helenae Csősz et al., T. recedens (Nylander), Tetramorium kephalosi Salata & Borowiec; Taygetos Mts., 1.3 km S of Artemisia: Aphaenogaster subterranea (Latreille), Camponotus boghossiani Forel, Crematogaster schmidti (Mayr), Lasius cf. alienus, L. illyricus Zimmermann, Myrmecina graminicola (Latreille), Plagiolepis pygmaea (Latreille), Temnothorax crasecundus Seifert & Csősz, T. helenae Csősz et al.; Taygetos Mts., 1.8 km E of Saidona: Lasius illyricus Zimmermann, Plagiolepis pygmaea (Latreille), Temnothorax crasecundus Seifert & Csősz, Tetramorium cf. caespitum; Taygetos Mts., 2 km W of Arachova: Aphaenogaster cf. muelleriana, Bothriomyrmex communista Santschi, Camponotus boghossiani Forel, C. dalmaticus (Nylander), C. laconicus Emery, Cataglyphis nodus (Brullé), Crematogaster schmidti (Mayr), Lepisiota frauenfeldi (Mayr), Messor wasmanni Krausse, Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Stigmatomma denticulatum Roger, Tapinoma erraticum (Latreille), Temnothorax cf. bulgaricus, T. laconicus Csősz et al., T. cf. luteus, T. rogeri Emery, Tetramorium kephalosi Salata & Borowiec, Tetramorium cf. punctatum; Taygetos Mts., Arachova: Camponotus aethiops (Latreille), C. laconicus Emery, C. piceus (Leach), Cataglyphis nodus (Brullé), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Lasius cf. alienus, L. illyricus Zimmermann, Lepisiota frauenfeldi (Mayr), Messor wasmanni Krausse, Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Tapinoma erraticum (Latreille), Temnothorax cf. bulgaricus, T. parvulus (Schenck); Taygetos Mts., Chora Getson: Aphaenogaster balcanica (Emery), A. cf. muelleriana, Camponotus boghossiani Forel, Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Lasius cf. alienus, L. illyricus Zimmermann, Lepisiota frauenfeldi (Mayr), Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Temnothorax cf. bulgaricus, T. laconicus Csősz et al., T. recedens (Nylander), T. strymonensis Csősz, Salata & Borowiec; Taygetos Mts., Karveli: Aphaenogaster cf. muelleriana, Crematogaster schmidti (Mayr), Lasius cf. alienus, L. illyricus Zimmermann, Myrmecina graminicola (Latreille), Temnothorax helenae Csősz et al., T. strymonensis Csősz, Salata & Borowiec; Tetrazi Mts., 0.5 km E of Vastas: Aphaenogaster balcanica (Emery), Camponotus dalmaticus (Nylander), C. fallax (Nylander), C. laconicus Csősz et al., C. piceus (Leach), Colobopsis truncata (Spinola), Crematogaster schmidti (Mayr), Formica gagates Latreille, Lasius illyricus Zimmermann, Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Temnothorax helenae Csősz et al., T. laconicus Csősz et al., T. morea Csősz, Salata & Borowiec, T. rogeri Emery; Tetrazi Mts., Isaris: Aphaenogaster balcanica (Emery), A. subterranea (Latreille), Camponotus dalmaticus (Nylander), C. laconicus Csősz et al., C. lateralis (Olivier), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Lasius cf. alienus, L. illyricus Zimmermann, Lepisiota frauenfeldi (Mayr), Temnothorax cf. bulgaricus, T. helenae Csősz et al.,; Tetrazi Mts., Karnasi: Aphaenogaster balcanica (Emery), A. cf. muelleriana, Camponotus dalmaticus (Nylander), C. lateralis (Olivier), Crematogaster schmidti (Mayr), Lasius illyricus Zimmermann, Messor hellenius Agosti & Collingwood, Pheidole pallidula (Nylander), Temnothorax cf. bulgaricus, Temnothorax laconicus Csősz et al., T. morea Csősz, Salata & Borowiec, T. rogeri Emery, Tetramorium kephalosi Salata & Borowiec.
Sterea Ellas, Aetolia-Acarnania, Psila Alonia: Aphaenogaster balcanica (Emery), A. cf. muelleriana, A. subterranea (Latreille), Camponotus dalmaticus (Nylander), C. lateralis (Olivier), Cataglyphis nodus (Brullé), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Lasius illyricus Zimmermann, Messor wasmanni Krausse, Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Solenopsis cf. lusitanica, Temnothorax crassispinus (Karavaiev).
Sterea Ellas, Euboea, 1.2 km NW of Gerontas: Aphaenogaster balcanica (Emery), Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. gestroi Emery, C. kiesenwetteri (Roger), C. laconicus Emery, C. lateralis (Olivier), C. piceus (Leach), Cataglyphis aenescens (Nylander), Crematogaster ionia Forel, C. schmidti (Mayr), Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Tapinoma erraticum (Latreille), Temnothorax graecus (Forel), T. recedens (Nylander); 2.3 km S of Stropones: Camponotus aethiops (Latreille), C. piceus (Leach), C. vagus (Scopoli), Cataglyphis nodus (Brullé), Formica fusca Karavaiev, Lasius alienus Förster, L. flavus (Fabricius), Pheidole pallidula (Nylander), Temnothorax crasecundus Seifert & Csősz, T. helenae Csősz et al., T. unifasciatus (Latreille); 300 m NW of Agios: Camponotus aethiops (Latreille), C. dalmaticus (Nylander), Cataglyphis nodus (Brullé), Crematogaster ionia Forel, Dolichoderus quadripunctatus (Linnaeus), Formica fusca Karavaiev, Lasius lasioides (Emery), Pheidole cf. pallidula, Plagiolepis pygmaea (Latreille), Temnothorax lichtensteini (Bondroit), T. parvulus (Schenck), T. recedens (Nylander), T. unifasciatus (Latreille); 570 m NW of Drosia: Aphaenogaster balcanica (Emery), A. subterranea (Latreille), Camponotus dalmaticus (Nylander), C. ionius Emery, C. lateralis (Olivier), C. piceus (Leach), C. vagus (Scopoli), Cataglyphis nodus (Brullé), Crematogaster ionia Forel, C. schmidti (Mayr), Lepisiota frauenfeldi (Mayr), Messor hellenius Agosti & Collingwood, Temnothorax bulgaricus (Forel).
Thessaly, Larissa, Mt. Olympus, 5.3 km E of Olympiada: Aphaenogaster subterranea (Latreille), Camponotus aethiops (Latreille), C. lateralis (Olivier), C. piceus (Leach), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Formica cunicularia Latreille, F. gagates Latreille, Lasius alienus Förster, L. emarginatus (Olivier), Messor hellenius Agosti & Collingwood, Pheidole pallidula (Nylander), Plagiolepis pygmaea (Latreille), Prenolepis nitens (Mayr), Solenopsis cf. lusitanica, Tapinoma erraticum (Latreille), Temnothorax recedens (Nylander), Tetramorium kephalosi Salata & Borowiec, Tetramorium moravicum Kratochvil; Mt. Ossa, 2.4 km SE of Karitsa: Aphaenogaster subterranea (Latreille), Camponotus lateralis (Olivier), Cataglyphis nodus (Brullé), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Formica gagates Latreille, Lasius emarginatus (Olivier), Myrmoxenus gordiagini Ruzsky, Plagiolepis pygmaea (Latreille), Temnothorax crasecundus Seifert & Csősz, T. cf. subtilis-helenae, T. tauricus (Ruzsky), T. cf. unifasciatus, Tetramorium moravicum Kratochvil; Mt. Ossa, 600 m SE of Karitsa: Aphaenogaster subterranea (Latreille), Camponotus fallax (Nylander), Colobopsis truncata (Spinola), Lasius emarginatus (Olivier), Plagiolepis pygmaea (Latreille), Prenolepis nitens (Mayr), Temnothorax crasecundus Seifert & Csősz, T. lichtensteini (Bondroit), T. cf. subtilis-helenae, T. cf. unifasciatus; Mt. Ossa, Kokkino Nero: Aphaenogaster subterranea (Latreille), Camponotus aethiops (Latreille), C. dalmaticus (Nylander), C. lateralis (Olivier), Colobopsis truncata (Spinola), Crematogaster schmidti (Mayr), Dolichoderus quadripunctatus (Linnaeus), Formica cunicularia Latreille, Lasius alienus Förster, L. emarginatus (Olivier), Messor hellenius Agosti & Collingwood, Pheidole pallidula (Nylander), Prenolepis nitens (Mayr), Temnothorax tauricus (Ruzsky), T. cf. unifasciatus, Tetramorium cf. caespitum.

Original Description

• Salata, S; Borowiec, L; 2019: Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species ZooKeys, 877: 81-131. doi

Images

Figures 1, 2. Worker of Temnothorax brackoi sp. nov. 1 Dorsal 2 Lateral.  Figures 9–12. Head and antennae 9 Worker of Temnothorax brackoi sp. nov. 10 Worker of Temnothorax messiniaensis sp. nov. 11 Worker of Temnothorax turcicus (Santschi) 12 Gyne of Temnothorax messiniaensis sp. nov.  Figures 13–15. Head sculpture 13 Worker of Temnothorax brackoi sp. nov. 14 Worker of Temnothorax messiniaensis sp. nov. 15 Worker of Temnothorax turcicus (Santschi).