Difference between revisions of "Libanopsyllipsocus alexanderasnitsyni"

Latest revision as of 11:53, 24 September 2011

Taxonavigation

Ordo: Psocoptera
Familia: Psyllipsocidae
Genus: Libanopsyllipsocus

Name

Libanopsyllipsocus alexanderasnitsyni Azar & Nel, 2011 sp. n. – Wikispecies link – ZooBank link – Pensoft Profile

Holotype

Specimen n° 30 (male, coll. Azar); provisionally deposited in the Muséum National d’Histoire Naturelle, Paris, France.

Locality and horizon

Lower Cretaceous, Upper Barremian – Lower Aptian, amber of Hammana-Mdeyrij, Caza, Baabda, Mohafazat Jabal Loubnan, central Lebanon.

Etymology

In honor of our friend Prof. Alexandr P. Rasnitsyn, a world authority in entomology and palaeoentomology.

Diagnosis

As for the genus (vide supra).

Description

Dorsal parts of head and body not preserved. Body and wings without scales (Figs 1–2). Head elongate (but cannot be measured accurately due to state of preservation), sublingual sclerite and hypopharynx with two distinct filaments (Fig. 3). Forewing transparent (Fig. 4), with infrequent small setae, 0.96 mm long, 0.35 mm broad; apex rounded; Sc badly preserved, not reaching C nor R; R1 reaching costal margin 0.73 mm from wing base; pterostigma absent; Rs separating from R 0.46 mm from wing base; Rs bifurcating into R2+3 and R4+5 at 0.76 mm; R2+3 curved, reaching wing margin 0.83 mm distally; R4+5 curved, reaching wing margin at 0.95 mm distally; a transverse crossvein between Rs and M; fork of M into M1 and M2 0.8 mm from wing base, but one wing is aberrant in having M simple [a frequent teratology in Psyllipsocidae as noted by Lienhard (1998)^[1]]; Cu1 bifurcating into Cu1a and Cu1b 0.36 mm from wing base forming a free elongate and narrow areola postica; Cu1a much longer than Cu1b; Cu2 (badly preserved) joining anal vein at posterior wing margin in a nodulus 0.28 mm from wing base (Fig. 5); A strongly convex. Wing margin glabrous (Fig. 6), similar in structure to that illustrated by Lienhard (1998: 111, fig. 31b). Hind wing transparent (Fig. 4), bare and smaller than forewing, about 0.75 mm long, 0.24 mm broad; free or individualized and distinct arm of R1 missing; Rs forked into R2+3 and R4+5 0.16 mm from wing apex; basal cell present; simple M reaching wing margin apically; basal part of wing badly preserved. Hind leg with tibia + tarsi slightly longer than abdomen. Coxal rasp of Pearman’s organ present on hind leg (Fig. 7), but without tympanum (the coxal rasp is preserved on one leg as the coxal area is missing on the other leg). Tibiae with two apical spurs on inner sides; all tarsi three-segmented with first tarsomere longer than others; pretarsal claws with a preapical tooth, without microtrichia nor pulvillus (Fig. 8). Abdomen 0.52 mm long, 0.32 mm broad; telson with epiproct and paraproct as represented in Figs 9–12, paraproct with a rounded hemispherical sensory area of trichobotria, and a spine in its ventral inner side; male genitalia with simple hypandrium, aedeagus with phallosome presenting parameres with curved arms not fused anteriorly as represented in Fig. 9 (it can be seen owing to transparency of sclerites and is comparable in structure to what occurs in other Psyllipsocidae (cf. Lienhard 1998^[1]: 112, fig. 32b).

Original Description

• Azar, D; Nel, A; 2011: The oldest psyllipsocid booklice, in Lower Cretaceous amber from Lebanon (Psocodea, Trogiomorpha, Psocathropetae, Psyllipsocidae) ZooKeys, 130: 153-165. doi

Other References

1. ↑ ^{1.0 1.1} Lienhard C (1998) Psocoptères Euro-Méditerranéens. Faune de France 83, 1-533.

Images

Figure 1. Photograph of habitus of Libanopsyllipsocus alexanderasnitsyni gen. et sp. n., holotype, male, specimen number 30.  Figure 2. Drawing of habitus of Libanopsyllipsocus alexanderasnitsyni gen. et sp. n., holotype, male, scale bar = 0.3 mm.  Figure 3. Photograph of hypopharynx filaments (arrows) of Libanopsyllipsocus alexanderasnitsyni gen. et sp. n., holotype, male.  Figure 4. Drawing of wings of Libanopsyllipsocus alexanderasnitsyni gen. et sp. n., holotype, male, scale bar = 0.3 mm.  Figure 5. Microphotograph of nodulus, arrow showing the meeting area of Cu2 and A.  Figure 6. Microphotograph of structure of forewing margin.  Figure 7. Microphotograph of hind leg coxal rasp (Pearman’s organ).  Figure 8. Drawing of pretarsal claw of Libanopsyllipsocus alexanderasnitsyni gen. et sp. n., holotype, male, scale bar = 0.03 mm.  Figure 9. Drawing of aedeagus of Libanopsyllipsocus alexanderasnitsyni gen. et sp. n., holotype, male; Ep = epiproct, Hyp = hypandrium, Par = paraproct, par = paramers, Pha = phallosome, sp = anal spine, trich = trichobothria; scale bar = 0.3 mm.  Figure 10. Drawing of hypandrium of Libanopsyllipsocus alexanderasnitsyni gen. et sp. n., holotype, male; par = paraproct, ep = epiproct, trich = trichobothria; scale bar = 0.3 mm.  Figure 11. Photograph of hypandrium and aedeagus of Libanopsyllipsocus alexanderasnitsyni gen. et sp. n., holotype, male.  Figure 12. Photograph of paraproct of Libanopsyllipsocus alexanderasnitsyni gen. et sp. n., holotype, male; arrow shows the anal spine.